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    • To provide a comprehensive and accurate resource of cell markers for brain, 328 brain cell markers were collected from CellMarker Database.

    Cell Marker Cell Type Source PMID
    A2M Endothelial cell Single-cell sequencing 26060301
    ABCA1 Astrocyte Single-cell sequencing 27806376
    ACSL6 Astrocyte Single-cell sequencing 27806376
    ACTG1 Oligodendrocyte Single-cell sequencing 27806376
    ADAMTS4 Oligodendrocyte Single-cell sequencing 26060301
    ADCYAP1R1 Astrocyte Single-cell sequencing 27806376
    ADORA3 Microglial cell Single-cell sequencing 29262845
    ADRB2 Microglial cell Single-cell sequencing 29262845
    AFAP1L2 Oligodendrocyte Single-cell sequencing 27806376
    AGT Astrocyte Single-cell sequencing 27806376
    AGXT Astrocyte Single-cell sequencing 27806376
    ALDH1L1 Astrocyte Experiment 26295203
    ALDH1L1 Astrocyte Single-cell sequencing 26060301
    ANGPTL2 Oligodendrocyte Single-cell sequencing 27806376
    ANLN Astrocyte Single-cell sequencing 27806376
    APOLD1 Endothelial cell Single-cell sequencing 26060301
    ARHGEF26 Astrocyte Single-cell sequencing 27806376
    ASCL1 Neural stem cell Company
    ASIC1 Oligodendrocyte Single-cell sequencing 27806376
    ASPA Oligodendrocyte Single-cell sequencing 26060301
    ATP13A4 Astrocyte Single-cell sequencing 27806376
    ATP13A4 Astrocyte Single-cell sequencing 26060301
    BBOX1 Astrocyte Single-cell sequencing 27806376
    BCL2A1 Microglial cell Single-cell sequencing 26060301
    BHLHE40 Macrophage Single-cell sequencing 29262845
    BHLHE41 Microglial cell Single-cell sequencing 29262845
    BIN1 Oligodendrocyte Single-cell sequencing 27806376
    BIN1 Microglial cell Single-cell sequencing 29262845
    BMI1 Neural stem cell Company
    BMPR1B Astrocyte Single-cell sequencing 26060301
    BTG2 Astrocyte Single-cell sequencing 27806376
    ARMH4 Neuron Single-cell sequencing 26060301
    SNHG29 Oligodendrocyte Single-cell sequencing 27806376
    MIR9-1HG Astrocyte Single-cell sequencing 27806376
    CASQ1 Astrocyte Single-cell sequencing 27806376
    CCL2 Microglial cell Single-cell sequencing 26060301
    CCL4 Microglial cell Single-cell sequencing 26060301
    CCL4 Microglial cell Single-cell sequencing 29262845
    PROM1 Neural stem cell Experiment 25082219
    MCAM Neural stem cell Company
    CD163 M2 macrophage Experiment 21737148
    MS4A1 B cell Experiment 18825001
    NGFR Neural progenitor cell Experiment 24604282
    ITGB1 Neural stem cell Company
    TNFRSF8 T helper2 cell Experiment 12396479
    ABCG2 Neural stem cell Company
    CD40 Astrocyte Company
    CD68 Macrophage Experiment 18825001
    CD68 Microglial cell Experiment 18825001
    CD68 Macrophage Experiment 24405578
    CD68 Macrophage Experiment 28987033
    CD80 Astrocyte Company
    CD81 Neural stem cell Company
    CD83 Microglial cell Single-cell sequencing 26060301
    CD86 Astrocyte Company
    C5AR1 Astrocyte Company
    CD93 Macrophage Single-cell sequencing 29262845
    CDC42EP4 Astrocyte Single-cell sequencing 27806376
    CDHR1 Oligodendrocyte Single-cell sequencing 27806376
    CDK18 Oligodendrocyte Single-cell sequencing 26060301
    CHRDL1 Astrocyte Single-cell sequencing 26060301
    CHST9 Astrocyte Single-cell sequencing 27806376
    CIB1 Macrophage Single-cell sequencing 29262845
    CIITA Macrophage Single-cell sequencing 29262845
    CLDN11 Oligodendrocyte Single-cell sequencing 26060301
    CLU Astrocyte Single-cell sequencing 27806376
    CPNE5 Astrocyte Single-cell sequencing 27806376
    CRB1 Oligodendrocyte Single-cell sequencing 27806376
    CRYAB Astrocyte Single-cell sequencing 27806376
    CSF1R Microglial cell Single-cell sequencing 26060301
    CSPG4 Oligodendrocyte Single-cell sequencing 27806376
    VSIG2 Astrocyte Single-cell sequencing 26060301
    VSIG2 Astrocyte Single-cell sequencing 26060301
    CXCR4 Neural stem cell Company
    CYBB Macrophage Single-cell sequencing 29262845
    CYBRD1 Astrocyte Single-cell sequencing 26060301
    CYTH1 Macrophage Single-cell sequencing 29262845
    DAND5 Astrocyte Single-cell sequencing 27806376
    DCN Neuron Single-cell sequencing 26060301
    DCX Neural stem cell Experiment 21275797
    DCX Neural progenitor cell Experiment 21275797
    DHCR24 Oligodendrocyte Single-cell sequencing 27806376
    DKK3 Astrocyte Single-cell sequencing 27806376
    DNAJB1 Astrocyte Single-cell sequencing 27806376
    DNASE2 Astrocyte Single-cell sequencing 27806376
    DOK5 Astrocyte Single-cell sequencing 27806376
    DPYD Oligodendrocyte Single-cell sequencing 26060301
    DPYSL5 Neuron Single-cell sequencing 26060301
    DSE Macrophage Single-cell sequencing 29262845
    DTNA Astrocyte Single-cell sequencing 27806376
    EBP Oligodendrocyte Single-cell sequencing 27806376
    SH3D19 Oligodendrocyte Single-cell sequencing 27806376
    EDNRB Astrocyte Single-cell sequencing 27806376
    EEF2 Oligodendrocyte Single-cell sequencing 27806376
    EFEMP1 Astrocyte Single-cell sequencing 27806376
    EGFR Neuron Single-cell sequencing 26060301
    EGLN3 Astrocyte Single-cell sequencing 27806376
    EIF3E Oligodendrocyte Single-cell sequencing 27806376
    EIF3L Oligodendrocyte Single-cell sequencing 27806376
    EPAS1 Astrocyte Single-cell sequencing 27806376
    EPN2 Oligodendrocyte Single-cell sequencing 27806376
    ERMN Oligodendrocyte Single-cell sequencing 26060301
    F3 Astrocyte Single-cell sequencing 27806376
    CNTN1 Astrocyte Single-cell sequencing 27806376
    FA2H Oligodendrocyte Single-cell sequencing 27806376
    FA2H Oligodendrocyte Single-cell sequencing 26060301
    FABP7 Oligodendrocyte Single-cell sequencing 27806376
    CYRIA Macrophage Single-cell sequencing 29262845
    FAU Oligodendrocyte Single-cell sequencing 27806376
    FERMT1 Oligodendrocyte Single-cell sequencing 27806376
    FGFBP3 Oligodendrocyte Single-cell sequencing 27806376
    FGFR3 Astrocyte Single-cell sequencing 26060301
    FGR Macrophage Single-cell sequencing 29262845
    FLT1 Endothelial cell Single-cell sequencing 26060301
    FOS Astrocyte Single-cell sequencing 27806376
    FOSB Astrocyte Single-cell sequencing 27806376
    FOSL2 Macrophage Single-cell sequencing 29262845
    FPR3 Macrophage Single-cell sequencing 29262845
    FTH1 Astrocyte Single-cell sequencing 27806376
    GADD45B Astrocyte Single-cell sequencing 27806376
    GAP43 Oligodendrocyte Single-cell sequencing 27806376
    GJB1 Oligodendrocyte Single-cell sequencing 26060301
    GLI3 Astrocyte Single-cell sequencing 26060301
    GLIS3 Astrocyte Single-cell sequencing 26060301
    RACK1 Oligodendrocyte Single-cell sequencing 27806376
    GPR37 Oligodendrocyte Single-cell sequencing 26060301
    ADGRG1 Neural stem cell Experiment 19525879
    ADGRG1 Neural progenitor cell Experiment 19525879
    GPR65 Macrophage Single-cell sequencing 29262845
    GRAMD2B Astrocyte Single-cell sequencing 27806376
    GRIA1 Astrocyte Single-cell sequencing 27806376
    GRIA2 Oligodendrocyte Single-cell sequencing 27806376
    GRIA3 Neuron Single-cell sequencing 26060301
    GRIA4 Oligodendrocyte Single-cell sequencing 27806376
    GRIK2 Oligodendrocyte Single-cell sequencing 27806376
    GRIK5 Oligodendrocyte Single-cell sequencing 27806376
    GRM3 Oligodendrocyte Single-cell sequencing 26060301
    HEPACAM Astrocyte Single-cell sequencing 27806376
    HEY1 Astrocyte Single-cell sequencing 27806376
    HGF Astrocyte Single-cell sequencing 26060301
    IL6 Astrocyte Single-cell sequencing 26060301
    HIF1A Astrocyte Single-cell sequencing 27806376
    HIF3A Astrocyte Single-cell sequencing 27806376
    HLA-DMB Macrophage Single-cell sequencing 29262845
    HLA-DQA1 Macrophage Single-cell sequencing 29262845
    HSPB8 Astrocyte Single-cell sequencing 27806376
    AIF1 Macrophage Experiment 28987033
    ID1 Astrocyte Single-cell sequencing 27806376
    IFITM2 Macrophage Single-cell sequencing 29262845
    IGFBP7 Astrocyte Single-cell sequencing 27806376
    IL10 Macrophage Single-cell sequencing 29262845
    IL1RN Macrophage Single-cell sequencing 29262845
    IL33 Astrocyte Single-cell sequencing 27806376
    IQGAP1 Macrophage Single-cell sequencing 29262845
    ITGA4 Macrophage Single-cell sequencing 29262845
    JUN Astrocyte Single-cell sequencing 27806376
    KCNH7 Astrocyte Single-cell sequencing 27806376
    KCNH8 Oligodendrocyte Single-cell sequencing 26060301
    KCNIP3 Oligodendrocyte Single-cell sequencing 27806376
    KCNK1 Neuron Single-cell sequencing 26060301
    ELAPOR1 Neuron Single-cell sequencing 26060301
    KIF21A Astrocyte Single-cell sequencing 27806376
    KLK6 Oligodendrocyte Single-cell sequencing 26060301
    KLRC2 Oligodendrocyte Single-cell sequencing 27806376
    L1CAM Astrocyte Single-cell sequencing 27806376
    LGR5 Stem cell Experiment 25451725
    LHFPL6 Astrocyte Single-cell sequencing 27806376
    LIMS2 Oligodendrocyte Single-cell sequencing 27806376
    LIX1 Astrocyte Single-cell sequencing 27806376
    LMF1 Oligodendrocyte Single-cell sequencing 27806376
    LNX1 Neuron Single-cell sequencing 26060301
    LPAR1 Oligodendrocyte Single-cell sequencing 26060301
    PLPPR1 Oligodendrocyte Single-cell sequencing 27806376
    LRIG1 Astrocyte Single-cell sequencing 27806376
    LRRC8A Astrocyte Single-cell sequencing 27806376
    LRRN1 Oligodendrocyte Single-cell sequencing 27806376
    LRCH4 Oligodendrocyte Single-cell sequencing 27806376
    LYPD1 Astrocyte Single-cell sequencing 27806376
    MAG Oligodendrocyte Single-cell sequencing 26060301
    ERCC2 Oligodendrocyte Single-cell sequencing 26060301
    MAL Oligodendrocyte Single-cell sequencing 26060301
    MRTFA Oligodendrocyte Single-cell sequencing 26060301
    MAML2 Oligodendrocyte Single-cell sequencing 27806376
    MAP1A Oligodendrocyte Single-cell sequencing 27806376
    METAP1 Oligodendrocyte Single-cell sequencing 27806376
    MAP2 Neuron Experiment 24142904
    METAP2 Neuron Experiment 24142904
    MAP2 Oligodendrocyte Single-cell sequencing 27806376
    METAP2 Oligodendrocyte Single-cell sequencing 27806376
    MAP6D1 Oligodendrocyte Single-cell sequencing 26060301
    MARCKSL1 Oligodendrocyte Single-cell sequencing 27806376
    MARK4 Neuron Experiment 16973293
    PRG2 Oligodendrocyte Single-cell sequencing 26060301
    PRG2 Oligodendrocyte Single-cell sequencing 26060301
    MEGF11 Oligodendrocyte precursor cell Single-cell sequencing 26060301
    MICAL1 Oligodendrocyte Single-cell sequencing 27806376
    MIF Oligodendrocyte Single-cell sequencing 27806376
    S100A9 Oligodendrocyte Single-cell sequencing 27806376
    MLC1 Astrocyte Single-cell sequencing 27806376
    MS4A6A Macrophage Single-cell sequencing 29262845
    MS4A7 Macrophage Single-cell sequencing 29262845
    MS4A4A Macrophage Single-cell sequencing 29262845
    MT1X Astrocyte Single-cell sequencing 27806376
    MTSS1 Oligodendrocyte Single-cell sequencing 27806376
    MXD1 Macrophage Single-cell sequencing 29262845
    MYT1 Oligodendrocyte Single-cell sequencing 27806376
    PKMYT1 Oligodendrocyte Single-cell sequencing 27806376
    NAP1L1 Oligodendrocyte Single-cell sequencing 27806376
    NAV3 Microglial cell Single-cell sequencing 29262845
    NDRG2 Astrocyte Single-cell sequencing 27806376
    NELL1 Neuron Single-cell sequencing 26060301
    RBFOX3 Neuron Experiment 28877758
    RBFOX3 Neuron Experiment 9809557
    NEUROD1 Neural stem cell Experiment 21275797
    NEUROD1 Neural progenitor cell Experiment 21275797
    NFASC Neuron Single-cell sequencing 26060301
    NFIA Astrocyte Experiment 26295203
    NFIL3 Macrophage Single-cell sequencing 29262845
    NFKBIA Astrocyte Single-cell sequencing 27806376
    CSPG4 Oligodendrocyte progenitor cell Experiment 27105664
    CSPG4 Oligodendrocyte progenitor cell Experiment 18281150
    CSPG4 Pericyte Experiment 25360079
    NGFR Neural stem cell Experiment 25514637
    NGFR Neural progenitor cell Experiment 25514637
    NHSL1 Astrocyte Single-cell sequencing 27806376
    NME1 Oligodendrocyte Single-cell sequencing 27806376
    RMRP Oligodendrocyte Single-cell sequencing 27806376
    NME2 Oligodendrocyte Single-cell sequencing 27806376
    NOG Astrocyte Single-cell sequencing 27806376
    RTN4 Oligodendrocyte Experiment 18515322
    NPM1 Oligodendrocyte Single-cell sequencing 27806376
    NPPA Oligodendrocyte Single-cell sequencing 27806376
    NRP1 Astrocyte Single-cell sequencing 27806376
    OLIG1 Oligodendrocyte Single-cell sequencing 27806376
    OLIG2 Oligodendrocyte Experiment 27596440
    OLIG2 Oligodendrocyte progenitor cell Experiment 27105664
    OLIG2 Oligodendrocyte Experiment 26110560
    OLIG2 Oligodendrocyte Single-cell sequencing 27806376
    OPALIN Oligodendrocyte Single-cell sequencing 26060301
    OPCML Oligodendrocyte Single-cell sequencing 27806376
    P2RX7 Oligodendrocyte Single-cell sequencing 27806376
    P2RY1 Astrocyte Single-cell sequencing 27806376
    P2RY12 Microglial cell Single-cell sequencing 29262845
    PAQR6 Astrocyte Single-cell sequencing 26060301
    PAX6 Stem cell Experiment 24113065
    PCDH15 Oligodendrocyte precursor cell Single-cell sequencing 26060301
    PDGFRA Oligodendrocyte precursor cell Single-cell sequencing 26060301
    PEA15 Astrocyte Single-cell sequencing 27806376
    PER1 Astrocyte Single-cell sequencing 27806376
    PGAP3 Astrocyte Single-cell sequencing 27806376
    PHACTR3 Oligodendrocyte Single-cell sequencing 27806376
    PHLDA1 Oligodendrocyte Single-cell sequencing 27806376
    PHLDB1 Oligodendrocyte Single-cell sequencing 27806376
    PID1 Oligodendrocyte Single-cell sequencing 27806376
    PIM1 Macrophage Single-cell sequencing 29262845
    LONP1 Macrophage Single-cell sequencing 29262845
    PLAC8 Macrophage Single-cell sequencing 29262845
    LGALS13 Macrophage Single-cell sequencing 29262845
    PLEKHB1 Oligodendrocyte Single-cell sequencing 26060301
    PLP1 Oligodendrocyte Single-cell sequencing 26060301
    PNOC Neuron Single-cell sequencing 26060301
    POLR2F Oligodendrocyte Single-cell sequencing 27806376
    PPP1R14A Oligodendrocyte Single-cell sequencing 26060301
    PPP1R3C Astrocyte Single-cell sequencing 26060301
    SLC66A3 Macrophage Single-cell sequencing 29262845
    PRLHR Astrocyte Single-cell sequencing 27806376
    PRRT2 Astrocyte Single-cell sequencing 27806376
    PSAP Astrocyte Single-cell sequencing 27806376
    MTCH1 Astrocyte Single-cell sequencing 27806376
    PTPN7 Macrophage Single-cell sequencing 29262845
    RASSF4 Astrocyte Single-cell sequencing 27806376
    RELN Neuron Single-cell sequencing 26060301
    RGMB Oligodendrocyte Single-cell sequencing 27806376
    RHOB Astrocyte Single-cell sequencing 27806376
    RHOB Microglial cell Single-cell sequencing 29262845
    RND3 Astrocyte Single-cell sequencing 27806376
    RNF43 Astrocyte Single-cell sequencing 26060301
    RPL13A Oligodendrocyte Single-cell sequencing 27806376
    RPS17 Oligodendrocyte Single-cell sequencing 27806376
    RPS2 Oligodendrocyte Single-cell sequencing 27806376
    RPS23 Oligodendrocyte Single-cell sequencing 27806376
    SALL1 Microglial cell Single-cell sequencing 29262845
    SAMHD1 Macrophage Single-cell sequencing 29262845
    SCD Oligodendrocyte Single-cell sequencing 27806376
    SLC22A5 Oligodendrocyte Single-cell sequencing 27806376
    SCG2 Astrocyte Single-cell sequencing 27806376
    SCG3 Astrocyte Single-cell sequencing 27806376
    SGCG Astrocyte Single-cell sequencing 27806376
    SEC14L5 Oligodendrocyte Single-cell sequencing 26060301
    SEMA6A Astrocyte Single-cell sequencing 27806376
    SERINC5 Oligodendrocyte Single-cell sequencing 27806376
    SERPINA3 Astrocyte Single-cell sequencing 27806376
    SF3A1 Astrocyte Single-cell sequencing 27806376
    SHC4 Oligodendrocyte Single-cell sequencing 26060301
    SHISA4 Oligodendrocyte Single-cell sequencing 27806376
    SIGLEC8 Microglial cell Single-cell sequencing 29262845
    SLC14A1 Astrocyte Single-cell sequencing 26060301
    SLC1A1 Oligodendrocyte Single-cell sequencing 27806376
    SLC1A2 Astrocyte Single-cell sequencing 27806376
    SLC1A3 Astrocyte Single-cell sequencing 27806376
    SLC1A3 Microglial cell Single-cell sequencing 29262845
    SLC30A10 Astrocyte Single-cell sequencing 26060301
    SLC4A4 Astrocyte Single-cell sequencing 26060301
    SLITRK1 Neuron Single-cell sequencing 26060301
    SNX1 Oligodendrocyte Single-cell sequencing 27806376
    SORL1 Astrocyte Single-cell sequencing 27806376
    SOX9 Neural stem cell Experiment 16797497
    SOX9 Astrocyte Experiment 28336567
    SOX9 Neural stem cell Company
    SOX9 Astrocyte Single-cell sequencing 27806376
    SPINT2 Macrophage Single-cell sequencing 29262845
    SPOCK1 Astrocyte Single-cell sequencing 27806376
    SPON1 Astrocyte Single-cell sequencing 27806376
    SST Neuron Single-cell sequencing 26060301
    ST6GAL2 Astrocyte Single-cell sequencing 27806376
    TACR1 Astrocyte Single-cell sequencing 27806376
    TAGLN3 Oligodendrocyte Single-cell sequencing 27806376
    TAL1 Microglial cell Single-cell sequencing 29262845
    TBC1D10A Astrocyte Single-cell sequencing 27806376
    TCF12 Oligodendrocyte Single-cell sequencing 27806376
    TGFBI Macrophage Single-cell sequencing 29262845
    THBD Macrophage Single-cell sequencing 29262845
    TM4SF1 Endothelial cell Single-cell sequencing 26060301
    TM7SF2 Oligodendrocyte Single-cell sequencing 27806376
    TMEFF2 Oligodendrocyte Single-cell sequencing 27806376
    TMEM123 Macrophage Single-cell sequencing 29262845
    TMEM130 Neuron Single-cell sequencing 26060301
    TMEM151B Astrocyte Single-cell sequencing 27806376
    TMEM97 Oligodendrocyte Single-cell sequencing 27806376
    TMSB4X Oligodendrocyte Single-cell sequencing 27806376
    TNFSF13 Macrophage Single-cell sequencing 29262845
    TNK2 Oligodendrocyte Single-cell sequencing 27806376
    TNS1 Astrocyte Single-cell sequencing 27806376
    TOB2 Astrocyte Single-cell sequencing 27806376
    TPCN1 Astrocyte Single-cell sequencing 27806376
    TRAF4 Oligodendrocyte Single-cell sequencing 27806376
    TREM1 Macrophage Single-cell sequencing 29262845
    TRIM66 Astrocyte Single-cell sequencing 26060301
    TSC22D4 Astrocyte Single-cell sequencing 27806376
    TSPAN12 Astrocyte Single-cell sequencing 27806376
    TSPO M1 macrophage Experiment 28968465
    TUBB Oligodendrocyte Single-cell sequencing 27806376
    TUBB2A Oligodendrocyte Single-cell sequencing 27806376
    TUBB3 Oligodendrocyte Single-cell sequencing 27806376
    TUBB4A Oligodendrocyte Single-cell sequencing 27806376
    UQCRB Oligodendrocyte Single-cell sequencing 27806376
    SLC17A7 Glutamatergic neuron Experiment 17651422
    SLC17A6 Glutamatergic neuron Experiment 17651422
    VIPR2 Oligodendrocyte Single-cell sequencing 27806376
    VOPP1 Macrophage Single-cell sequencing 29262845
    ZCCHC24 Oligodendrocyte Single-cell sequencing 27806376
    ZDHHC9 Oligodendrocyte Single-cell sequencing 27806376
    ZEB2 Oligodendrocyte Single-cell sequencing 27806376
    ZFAND5 Astrocyte Single-cell sequencing 27806376
    ZFP36 Astrocyte Single-cell sequencing 27806376
    ZFP36L1 Astrocyte Single-cell sequencing 27806376
    ZFP36L2 Astrocyte Single-cell sequencing 27806376