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Anatomical record (Hoboken, N.J. : 2007)
May 29, 2023;

The turtles from the middle Paleolithic site of Gruta Nova da Columbeira (Bombarral, Portugal): Update through an archaeozoological perspective.

Iratxe Boneta Jiménez, João Luis Cardoso, Adán Pérez-García
Author Information
  1. Iratxe Boneta Jiménez: Departamento de Prehistoria y Arqueología, Facultad de Filosofía y Letras, Universidad Autónoma de Madrid, Carretera de Colmenar Viejo, Madrid, Spain. ORCID
  2. João Luis Cardoso: Departamento de Ciências Sociais e de Gestão, Universidade Aberta. R. da Escola Politécnica 141, Lisboa, Portugal. ORCID
  3. Adán Pérez-García: Grupo de Biología Evolutiva, Facultad de Ciencias, Universidad Nacional de Educación a Distancia, Madrid, Spain. ORCID
PMID: 37246494 DOI: 10.1002/ar.25229

Abstract

Twenty-five years after the preliminary systematic study of the turtle remains (Agrionemys [=Testudo] hermanni and Emys or Mauremys) recovered from Gruta Nova da Columbeira site (Bombarral, Portugal), the results of its review from systematic and archaeozoological perspectives are presented here. Tortoise remains studies from pre-Upper Paleolithic sites worldwide have provided relevant data confirming its role as a dietary supply for hominid populations and informing about their ability to adapt to local environmental resources. The Iberian Peninsula record in general, and specifically, that from Portugal, have yielded substantial evidence to this highly debated topic. In this sense, turtle remains recovered in Gruta Nova da Columbeira site, discovered in the 1960s and the main ensemble chronologically ascribed to the MIS-5 (87.1 ± 6.3 ka BP), offer new information to this debate. Its detailed restudy, has allowed us the identification, justification, and figuration of remains attributed to two Iberian turtle taxa, Chersine hermanni and Emys orbicularis. Therefore, this update on the data concerning the turtle record from Gruta Nova da Columbeira provides new justified taxonomic evidence regarding the Iberian turtle taxa distribution during the Upper Pleistocene. The previously suggested hypothesis about the tortoise human consumption on the site is here evaluated through the development of an archaeozoological and taphonomical analysis, as well as considering the potential documentation of anthropic alterations (e.g., burning, cutmarks, percussion marks). In this sense, this hypothesis is confirmed. In addition, the presence of carnivore activity evidence indicates the engagement of other agents in the deposit formation.

Keywords

Chersine hermanni Emys orbicularis Neanderthal tortoise consumption upper Pleistocene

References

  1. Albizuri, S., Colomer, S., & Buisan, C. (1993). Experimentación sobre la exposición del tejido óseo a focos de calor. Estudios de la Antigüedad, 6(7), 91-97.
  2. Antunes, M. T., & Cardoso, J. L. (2000). Gruta Nova da Columbeira, Gruta das Salemas and Gruta da Figueira Brava. Stratigraphy, and chronology of the Pleistocene deposits. In M. T. Antunes (Ed.), Last Neanderthals in Portugal odontologic and other evidence (pp. 23-67). Academia das Ciências de Lisboa.
  3. Antunes, M. T., Santinho Cunha, A., Schwartz, J. H., & Tattersall, I. (2000). The latest Neanderthals: Evidence from Portugal. In M. T. Antunes (Ed.), Last Neanderthals in Portugal odontologic and other evidence (pp. 283-317). Lisboa.
  4. Archer, W., & Braun, D. (2013). Investigating the signature of aquatic resource use within Pleistocene Hominin Dietary Adaptations. PLoS One, 8(8), e69899.
  5. Archer, W., Braun, D., Harris, J., McCoy, J., & Richmond, B. (2014). Early Pleistocene aquatic resource use in the Turkana Basin. Journal of Human Evolution, 77, 74-87.
  6. Arenas, P. (2003). Etnografía y Alimentación Entre los Tobas Ñachilamoleek y Wichi-Lhukutas del Chaco Central (Argentina). Arenas editor.
  7. Auffenberg, W. (1981). The fossil turtles of Olduvai Gorge, Tanzania, Africa. Copeia, 3, 509-522.
  8. Ayres, C. (2015). Galápago europeo - Emys orbicularis (Linnaeus, 1758). In A. Salvador & A. Marco (Eds.), Enciclopedia Virtual de los Vertebrados Españoles (pp. 1-22). Museo Nacional de Ciencias Naturales. http://www.vertebradosibericos.org/
  9. Bailón, S. (2010). Quelonios fósiles del yacimiento de Barranco León (Pleistoceno Inferior, Orce, Granada, España). In I. Toro, B. Martínez-Navarro, & J. Agustí (Eds.), Ocupaciones humanas en el Pleistoceno Inferior y Medio de la Cuenca de Guadix-Baza (pp. 185-195). Sevilla.
  10. Bertolero, A. (2015). Tortuga mediterránea - Testudo hermanni Gmelin, 1789. In A. Salvador & A. Marco (Eds.), Enciclopedia Virtual de los Vertebrados Españoles (pp. 1-34). Museo Nacional de Ciencias Naturales. http://www.vertebradosibericos.org/
  11. Bicho, N. F. (2004a). The middle Paleolithic occupation of southern Portugal. In N. Conard (Ed.), Settlement dynamics of the middle Paleolithic and middle stone age II (pp. 513-531). Tübingen.
  12. Bicho, N. F. (2004b). As comunidades humanas de caçadores-recolectores do Algarve Ocidental. Perspectiva ecológica. In A. A. Tavares, M. J. F. Tavares, & J. L. Cardoso (Eds.), Evolução Geohistórica do Litoral Portugês e Fenómenos Correlativos. Geologia, Historia, Arqueologia e Climatologia. Actas (pp. 359-396). Universidade Aberta.
  13. Blain, H.-A., Agustí, J., Lordkipanidze, D., Rook, L., & Delfino, M. (2014). Paleoclimatic and paleoenvironmental context of the early Pleistocene hominins from Dmanisi (Georgia, lesser Caucasus) inferred from the herpetofaunal assemblage. Quaternary Science Reviews, 105, 136-150.
  14. Blasco, R. (2008). Human consumption of tortoises at level IV of Bolomor Cave (Valencia, Spain). Journal of Archaeological Science, 35, 2839-2848.
  15. Blasco, R., Blain, H.-A., Rosell, J., Díez, J. C., Huguet, R., Rodríguez, J., Arsuaga, J. L., Bermúdez de Castro, J. M., & Carbonell, E. (2011). Earliest evidence for human consumption of tortoises in the European early Pleistocene from Sima del Elefante, Sierra de Atapuerca, Spain. Journal of Human Evolution, 61, 503-509.
  16. Blasco, R., Rosell, J., Smith, K., Maul, L., Sañudo, P., Barkai, R., & Gopher, A. (2016). Tortoises as a dietary supplement: A view from the middle Pleistocene site of Qesem Cave, Israel. Quaternary Science Reviews, 133, 165-182.
  17. Boneta Jiménez, I. (2022). Los quelonios en el registro arqueológico de la península ibérica: Aproximación a su estudio a través del conjunto del yacimiento calcolítico de Camino de las Yeseras. PhD Dissertation. Universidad Autónoma de Madrid.
  18. Boneta Jiménez, I., Pérez-García, A., & Liesau von Lettow-Vorbeck, C. (2023). Chelonians from the Middle Palaeolithic Site of Mealhada (Coimbra, Portugal): An Update. Diversity, 15(2), 243. https://doi.org/10.3390/d15020243
  19. Braun, D. R., Harris, J. W. K., Levin, N. E., McCoy, J. T., Herries, A. I. R., Bamford, M. K., Bishop, L. C., Richmond, B. G., & Kibunjia, M. (2010). Early hominin diet included diverse terrestrial and aquatic animals 1.95 Ma in East Turkana, Kenya. Proceedings of the National Academy of Sciences of the United States of America, 107(22), 10002-10007.
  20. Broadley, D. (1997). A new species of Psammobates (Reptilia: Testudinidae) from the early Pleistocene of South Africa. Palaeontologica Africana, 42, 111-114.
  21. Brugal, J.-P., & Desse, J. (2004). Petits Animaux et Sociétés Humaines. Du Complément Alimentaire aux Ressources Utilitaires. In XXIV Rencontres Internationales d'Archéologie et d'Histoire d'Antibes (pp. 255-259). APDCA.
  22. Cardoso, J. L. (1993). Contribuçâo para o conhecimento dos grandes mamíferos do Pleistocénico Superior de Portugal. Cámara Municipal de Oeiras.
  23. Cardoso, J. L. (2006). The Mousterian complex in Portugal. Zephyrvs, 59, 21-50.
  24. Cardoso, J. L., Raposo, L., & da Veiga Ferreira, O. (2002). A Gruta Nova da Columbeira Bombarral. Cãmara Municipal do Bombarral.
  25. Carvalho, M., Pereira, T., & Manso, C. (2018). Rabbit exploitation in the middle Palaeolithic at Gruta Nova da Columbeira, Portugal. Journal of Archaeological Science: Reports, 21, 821-832.
  26. Clarke, R. (1998). First ever discovery of a well-preserved skull and associated skeleton of Autralopithecus. South African Journal of Science, 94, 460-463.
  27. Clason, A. T. (1972). Some remarks on the use and presentation of archaezoological data. Helinium, 12, 139-153.
  28. Crespo, E. (2002). Paleoherpetofauna portuguesa. Revista Española de Herpetología, Volumen Especial, 17-35.
  29. Daura, J., Sanz, M., Arsuaga, J.-L., Hoffman, D. L., Quam, R. M., Ortega, M. C., Santos, E., Gómez, S., Rubio, A., Villaescusa, L., Souto, P., Mauricio, J., Rodrigues, F., Ferreira, A., Godinho, P., Trinkaus, E., & Zilhão, J. (2017). New middle Pleistocene hominin cranium from Gruta da Aroeira (Portugal). Proceedings of the National Academy of Sciences of the United States of America, 114(13), 3397-3402.
  30. Davis, S. J. M. (2002). The mammals and birds from the Gruta do Caldeirão, Portugal. Revista Portuguesa de Arqueologia, 5(2), 29-98.
  31. Delgado, J. F. N. (1884). La grotte de Furninha a Peniche. In Congrès International d'Anthropologie et d'Archéologie Préhistoriques. Compte-Rendu de la neuvième Session à Lisbonne, 1880 (pp. 207-278). Typographie de l'Académie Royale des Sciences.
  32. Domínguez-Rodrigo, M., de Juana, S., Galán, A. B., & Rodríguez, M. (2009). A new protocol to differentiate trampling marks from butchery cut marks. Journal of Archaeological Science, 36, 2643-2654.
  33. Ferembach, D. (1964-1965). La molaire humaine inférieure moustérienne de Bombarral (Portugal). Comunicações dos Serviços Geológicos de Portugal, 48, 185-191.
  34. Fernández-Jalvo, Y., & Andrews, P. (2016). Atlas of Taphonomic identifications. 1001+ images of fossil and recent mammal bone modification. Springer.
  35. Fernández-Laso, M. C., Brugal, J.-P., & Raposo, L. (2015). Gruta Nova da Columbeira (Bombarral, Portugal): Un modelo de ocupación en cueva durante el Paleolítico Medio. Resultados del estudio del registro de macromamíferos. Trabajos de Prehistoria, 72(2), 304-326.
  36. Ferreira, O. d. V. (1984). O mais importante nível de ocupação do caçador Neandertal da Gruta Nova da Columbeira (Bombarral). In Volume d'Hommage au géologue Georges Zbyszewski (pp. 365-370). Éditions Recherche sur les Civilisations.
  37. Figueiredo, S., & Raposo, L. (2018). As Aves Como Recurso Alimentar do Homem do Paleolítico Médio: Interpretação tafonómica das acumulações faunísticas da Gruta Nova da Columbeira e da Foz do Enxarrique. Boletim do Centro Português de Geo-História e Pré-História, 1(1), 57-63.
  38. Figueiredo, S., Sousa, F., Silva, S., Raposo, L., Pereira, T., & Rosa, A. (2018). Pleistocene birds of Gruta Nova da Columbeira (Bombarral-Portugal): A paleontological and Paleoenvironmental approach. Journal of Environmental Health Science and Engineering, A7, 246-254.
  39. Grayson, D. K. (1984). Quantitative zooarchaeology: Topics in the analysis of archaeological faunas. Academic Press.
  40. Hans-Volker, K. (2012). Human consumption of turtles of the Homo rudolfensis site Uraha (Malawi, East Africa). Archeofauna, 21, 267-279.
  41. Harlé, E. (1910-1911). Les mammifères et oiseaux quaternaires connus jusqu'ici en Portugal. Communition du Service Géologique du Portugal, 8, 22-85.
  42. Hervet, S. (2000). Tortues du Quaternaire de France: Critères de détermination, répartition chronologique et géographique. Mésogée, 58, 3-47.
  43. Huguet, R. (2007). Primeras ocupaciones humanas en la Península Ibérica: Paleoeconomía en la Sierra de Atapuerca (Burgos) y la Cuenca de Guadix-Baza (Grana-da) durante el Pleistoceno Inferior. PhD Dissertation. Universitat Rovira i Virgili.
  44. Jiménez Fuentes, E., Cardoso, J. L., & Crespo, E. (1998). Presencia de Agrionemys (=Testudo) hermanni (Gmelin, 1789) en el Paleolítico Medio de la Gruta Nova da Columbeira (Bombarral, Provincia de Estremadura, Portugal). Studia Geologica Salmantisensia, 34, 123-139.
  45. Jiménez Fuentes, E., Gil, S., & Pollos, S. (1995). Quelonios del Pleistoceno Medio de las Grajas (Archidona: Málaga). Studia Geologica Salmantisensia, 31, 55-62.
  46. Jiménez-Fuentes, E. (1980). Los quelonios del sitio de ocupación achelense de Aridos-1 (Arganda, Madrid). In M. Santoja, L. López-Martínez, & A. Pérez-González (Eds.), Ocupaciones Achelenses en el Valle del Jarama. Geología, Paleontología, Paleoecología y Prehistoria (pp. 139-143). Publicaciones de la Excelentísima Diputación Provincial de Madrid.
  47. Keller, C. (1997). Ecología de las poblaciones de Mauremys Leprosa y Emys orbicularis en el Parque Nacional de Doñana. PhD Dissertation. Universidad de Sevilla.
  48. Keller, C., & Andreu, A. C. (2002). Emys orbicularis Galápago europeo. In J. M. Pleguezuelos, R. Márquez, & M. Lizana (Eds.), Atlas y Libro Rojo de los Anfibios y Reptiles de España, 2ª impresión (pp. 137-142). Dirección General de Conservación de la Naturaleza-Asociación Herpetológica Española.
  49. Klein, R., & Cruz-Uribe, K. (1987). Large mammals and tortoise bones from Eland's bay cave and nearby sites, western Cape Province, South Africa. In J. Parkington & M. Hall (Eds.), Papers in the prehistory of the Western Cape, South Africa (pp. 132-163). BAR International Series, 332.
  50. Lapparent de Broin, F. d., Bour, R., Parham, J. F., & Perälä, J. (2006). Eurotestudo, a new genus for the species Testudo hermanni Gmelin, 1789 (Chelonii, Testudinidae). Comptes Rendus Palevol, 5, 803-811.
  51. Lapparent de Broin, F. d., Bour, R., & Perälä, J. (2006b). Morphological definition of Eurotestudo (Testudinidae, Chelonii): First part. Annales de Paléontologie, 92, 255-304.
  52. Lapparent de Broin, F. d., Bour, R., & Perälä, J. (2006c). Morphological definition of Eurotestudo (Testudinidae, Chelonii): Second part. Annales de Paléontologie, 92, 325-357.
  53. Lapparent de Broin, F. D., & Antunes, M. T. (2000). Pleistocene Chelonians from Gruta da Figueira Brava (Arrábida, Portgal). In M. T. Antunes (Ed.), Last Neanderthals in Portugal odontologic and other evidence (pp. 105-116). Lisboa.
  54. Liesau von Lettow-Vorbeck, C. (1998). El Soto de Medinilla: Faunas de Mamíferos de la Edad del Hierro en el Valle del Duero (Valladolid, España). Archaeofauna, 7, 11-210.
  55. Lyman, R. L. (1994). Vertebrate Taphonomy. Cambridge University Press.
  56. Marks, A., Brugal, J., Chabai, V., Monigal, K., Goldberg, P., Hockett, B., Peman, E., Elorza, M., & Mallol, C. (2002). Le gisement pléistocène moyen de Galeria Pesada (Estrémadure, Portugal): Premiers résultats. Paleo, 14, 77-100.
  57. Morales, J., & Sanchis, A. (2009). The quaternary fossil record of the genus Testudo in the Iberian Peninsula. Archaeological implications and diachronic distribution in the western Mediterranean. Journal of Archaeological Science, 36, 1152-1162.
  58. Nabais, M. (2012). Middle Palaeolithic tortoise use at Gruta da Oliveira (Torres Novas, Portugal). In Actas das IV Jornadas de Jovens em Investigação Arqueológica - JIA 2011 (Vol. 1, pp. 251-258). Universidade do Algarve.
  59. Nabais, M., & Zilhão, J. (2019). The consumption of tortoise among last interglacial Iberian Neanderthals. Quaternary Science Reviews, 217, 225-246.
  60. Nicholson, R. (1993). A morphological investigation of burnt animal bone and an evaluation of its utility in archaeology. Journal of Archaeological Science, 20, 411-428.
  61. Partridge, T., Granger, D., Caffee, M., & Clarke, R. (2003). Lower Pliocene hominid remains from Sterkfontein. Science, 25, 607-612.
  62. Pérez Ripoll, M. (1992). Marcas de carnicería, fracturas intencionadas y mordeduras de carnívoros en huesos prehistóricos del Mediterráneo español. Instituto de Cultura “Juan-Gil Albert”.
  63. Pérez-García, A., Martín-Jiménez, M., Vlachos, E., & Codrea, V. (2022). The most complete extinct species of Testudo (Testudines, Testudinidae) defined by several well-preserved skeletons from the late Miocene of Romania. Journal of Systematic Palaeontology, 19-18, 1237-1270.
  64. Pérez-García, A., Murelaga, X., Mancheño, M. A., Aberasturi Rodríguez, A., & Romero, G. (2015). The tortoises from the lower Pleistocene paleontological site of Quibas (Región de Murcia, Spain). Comptes Rendus Palevol, 14, 598-603.
  65. Raposo, L., & Cardoso, J. L. (1997). Notas acerca das industrias musterienses da Gruta Nova da Columbeira. In Actas do II Congreso de Arqueología Peninsular (Zamora 1996), 1. Zamora (pp. 27-33). Edinburgh University Press.
  66. Raposo, L., & Cardoso, J. L. (1998). Las industrias líticas de la Gruta Nova de Columbeira (Bombarral, Portugal) en el contexto del Musteriense final de la Península Ibérica. Trabajos de Prehistoria, 55(1), 39-62.
  67. Real Margalef, C., Sanchis Serra, A., Morales Pérez, J. V., Bel Martínez, M. Á., Eixea VIlanova, A., Zilhão, J., & Villaverde Bonilla, V. (2019). Estudio de la fauna del nivel IV del Abrigo de la Quebrada y su aportación al conocimiento de la economía y el comportamiento humano en el Paleolítico medio de la vertiente Mediterránea Ibérica. SPAL, 28(2), 17-49.
  68. Rhodin, A. G. J., Thomson, S., Georgalis, G. L., Karl, H.-V., Danilov, I. G., Takahashi, A., De La Fuente, M. S., Bourque, J. R., Delfino, M., Bour, R., Iverson, J. B., Shaffer, H. B., & Van Dijk, P. P. (2015). Turtles and tortoises of the world during the rise and global spread of humanity: First checklist and review of extinct Pleistocene and Holocene Chelonians. In A. G. J. Rhodin, P. C. H. Pritchard, P. P. Van Dijk, R. Sumure, K. A. Buhlmann, J. B. Iverson, & R. A. Mittermeier (Eds.), Conservation biology of freshwater turtles and tortoises: A compilation project of the IUCN/SSC tortoise and freshwater turtle specialist group (Vols. Chelonian research monographs, 5) (pp. e.1-e.66). Chelonian Research Foundation.
  69. Royer, A., Laroulandie, V., Cochard, D., & Binder, D. (2011). Les brulures, des traces ambiguës aux origines multiples. Application aux vestiges des tortues de l'Abri du Mourre de Seve (Sorges, Vaucluse). In V. Laroulandie, J.-B. Mallye, & C. Denys (Eds.), Taphonomie des Petits Vertébrés: Référentiels et Transferts aux Fossiles (pp. 181-194). British Archaeological Reports, International Series, 2269.
  70. Sampson, C. (2000). Taphonomy of tortoises deposited by birds and bushmen. Journal of Archaeological Science, 27, 779-788.
  71. Sanchis, A., Morales, J., Pérez, L., Hernández, C., & Galván, B. (2015). La tortuga mediterránea en yacimientos valencianos del Paleolítico medio: Distribución, origen de las acumulaciones y nuevos datos procedentes del Abric del Pastor (Alcoi, Alacant). In A. Sanchis & J. Pascual (Eds.), Preses petites i grups humans en el passat (pp. 97-120). Valencia.
  72. Sanchis, S. A. (2010). Los lagomorfos del Paleolítico medio de la región central y sudoriental del Mediterráneo ibérico. Caracterización tafonómica y taxonómica. PhD Dissertation. Universitat de València.
  73. Sanchis Serra, A., & Fernández Peris, J. (2008). Procesado y consumo antrópico de conejo en la Cova del Bolomor (Tavernes de la Valldigna, Valencia). El nivel XVIIc (ca 350 ka)/Anthropic Processing and Consumption of Rabbit at the Bolomor Cave (Tavernes de la Valldigna, Valencia). The XVIIc level (ca 350 ka). Complutum, 19(1), 25-46.
  74. Speth, J., & Tchernov, E. (2002). Middle Paleolithic tortoise use at Kebara Cave (Israel). Journal of Archaeological Science, 29, 471-483.
  75. Steele, T. (2010). A unique hominin menu dated to 1.95 million years ago. Proceedings of the National Academy of Sciences of the United States of America, 107(24), 10771-10772.
  76. Stiner, M. C. (1993). Small animal exploitation and its relation to hunting, scavenging, and gathering in the Italian Mousterian. Archaeological Papers of the American Anthropological Association, 4(1), 107-125.
  77. Stiner, M. C. (2001). Thirty years on the “Broad Spectrum Revolution” and paleolithic demography. Proceedings of the National Academy of Sciences of the United States of America, 98(13), 6993-6996.
  78. Stiner, M. C. (2005). The faunas of the Hayonim Cave, Israel. A 20.000-year record of Paleolithic diet, demography and society. Peabody Museum of Archaeology and Ethnology, Harvard University.
  79. Stiner, M. C. (2013). An unshakable middle Paleolithic? Trends versus conservatism in the predatory niche and their social ramifications. Current Anthropology, 54(S8), S288-S304.
  80. Stiner, M. C., & Kuhn, S. L. (2006). Changes in the ‘connectedness’ and resilience of Paleolithic societies in the Mediterranean ecosystems. Human Ecology, 34, 693-712.
  81. Stiner, M. C., Kuhn, S. L., Weiner, S., & Bar-Yosef, O. (1995). Differential burning, recrystallization, and fragmentation of archaeological bone. Journal of Archaeological Science, 22, 223-237.
  82. Stiner, M. C., Munro, N., & Surovell, T. (2000). The tortoise and the hare: Small-game use, the broad-spectrum revolution, and Paleolithic demography. Current Anthropology, 41(1), 39-79.
  83. Stiner, M. C., Munro, N., Surovell, T., Tchernov, E., & Bar-Yosef, O. (1999). Paleolithic population growth pulses evidenced by small animal exploitation. Science, 283, 190-194.
  84. Stuart, A. J. (1979). Pleistocene occurrences of the European pond tortoise (Emys orbicularis L.) in Britain. Boreas, 8, 359-371.
  85. Thompson, J. (2010). Taphonomic analysis of the middle stone age faunal assemblage from pinnacle point cave 13B, Western Cape, South Africa. Journal of Human Evolution, 59, 321-339.
  86. Thompson, J., & Henshilwood, C. (2014). Tortoise taphonomy and tortoise butchery patterns at Blombos Cave, South Africa. Journal of Archaeological Science, 41, 214-229.
  87. Zangerl, R. (1969). The turtle shell. In C. Gans (Ed.), Biology of the Reptilia (Vol. 1, pp. 311-339). Academic Press Inc..
  88. Zilhão, J., Cardoso, J. L., Pike, A. W., & Weninger, B. (2011). Gruta Nova da Columbeira (Bombarral, Portugal): Site stratigraphy, age of the Mousterian sequence, and implications for the timing of Neanderthal extinction in Iberia. In W. Müller, B. V. Eriksen, M. Street, & G.-C. Weniger (Eds.), Quartär, Internationales Jahr-buch zur Eiszeitalter- und Steinzeitforschung; band 58/international yearbook for ice age and stone age research (Vol. 58, pp. 93-112). Leidorf.

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