http://192.168.164.12:81/ricewiki/api.php?action=feedcontributions&feedformat=atom&user=MikaRiceWiki - User contributions [en]2026-08-27T17:46:19ZUser contributionsMediaWiki 1.30.0https://ngdc.cncb.ac.cn/ricewiki/index.php?title=Os01g0182600&diff=183866Os01g01826002014-06-11T04:40:45Z<p>Mika: /* Evolution */</p>
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<div>The rice gene of ''Gigantea(GI)'' is solo ortholog of ''GI'' in the Arabidopsis circadian system and related to photoperiodic control of flowering.<br />
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==Annotated Information==<br />
===Function===<br />
*''OsGI'',a ''GI'' ortholog, regulates flowering time in response to photoperiodic conditions. Mutations in this gene delay flowering in inductive SDs (short-days),whereas flowering is marginally induced under LDs(long-days).In comparison, over expression of ''OsGI'' in transgenic rice causes late flowering in both SDs and LDs.In addition, ''OsGI'' has function to regulate ''Hd1'', a gene that also shows circadian rhythm<ref name="ref1" />. ''Os-GI'' affects gene expression of approximately 75% of genes among 27,201 genes and is required for strong amplitudes and proper phase-setting of global gene expression under natural field conditions<ref name="ref2" />. <br />
*The cDNA of ''OsGI'' firstly found in ''se5'' mutant that is completely photoperiod-insensitive, the expression patterns of rice and Arabidopsis ''GI'' gene are both photoperiodically regulated and are very similar under LD and SD conditions('''Figure 1''')<ref name="ref3" />. [[File:Limeijia2.png|frame|'''Figure 1''' Possible roles of isolated genes in the photoperiodic regulation of flowering in rice, including ''OsGI'', may function to inhibit flowering under LD conditions.(from reference <ref name="ref3" />).]]''GI'' is required for phase-dependent ''CO''transcription through the interaction with FKF1, which leads to cycling DNA binding with one finger (CDF) degradation and consequent derepression of CO. In addition, it has been known that GI protein interacts with ZEITLUPE (ZTL) F-box protein to control the diurnal rhythm of TOC1 protein in regulation of the Arabidopsis circadian clock<ref name="ref4" />. <br />
*''Os-GI'' also has function to maintain bioactive GA level through the regulation of the GA-deactivating enzyme genes in rice, ''Os-GI'' represses major ''OsGA2ox'' genes in order to maintain a proper bioactive GA level, ''osgi-1'' plants showed a semi-dwarf phenotype with elevated ''OsGA2ox'' gene expression('''Figure 2''') <ref name="ref4" /> .[[File:Limeijia1.png|frame|'''Figure 2''' WT rice and osgi-1 plants grown under natural field conditions and ''osgi-1'' plant exhibits Semi-dwarf phenotype(from reference <ref name="ref4" />).]]<br />
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===Expression===<br />
*''GI'' encodes a nuclear protein that defines the proper amplitude and period length of circadian rhythms. ''GI'' expression is regulated by the circadian clock, with a peak in transcript levels at 8 to 10 h after dawn. The ''GI'' protein acts between the circadian oscillator and ''CO'' to promote flowering by increasing ''CO'' and ''FT'' mRNA abundance<ref name="ref1" />. The expression of the ''OsGI'' messenger RNA was circadian-controlled and that its temporal expression pattern was very similar to that of ''GI'' under both SD and LD conditions. Decreased ''OsGI'' expression led to early flowering under LD conditions and late flowering under SD conditions relative to the wild type, and that this early flowering phenotype under LD conditions was the opposite of that of the Arabidopsis ''gi'' mutants, confirming the opposite role of the ''GI'' gene in flowering under LD conditions in rice and Arabidopsis<ref name="ref5" />.<br />
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===Mutation===<br />
*The single ''osgi-1'' mutation extended the time to flowering only under SD conditions. Under LD conditions the single ''hd1'' mutation induced early flowering, but ''osgi-1'' plants exhibited no phenotypic change<ref name="ref4" /><br />
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===Evolution===<br />
* There is some evolutionary conservation between rice circadian clock–related genes and those associated with Arabidopsis circadian clocks, where the circadian clock consists of interlocked subloops, it is also possible that ''Os-GI'' is a member of a subloop in rice circadian clocks<ref name="ref4" />.But there is a divergence in the photoperiodic pathways of these two species arises from the molecular mechanisms controlling the expression of ''FT/Hd3a'' by ''CO/Hd1(Se1)'' ('''Figure 3''')<ref name="ref5" />[[File:Limeijia3.png|frame|'''Figure 3''' The molecular mechanisms for the regulation of ''GI/OsGI'' expression, and the regulation of ''CO/Hd1(Se1)'' expression by ''GI/OsGI'', might be conserved between ''Arabidopsis'' and rice. Both the ''GI–CO'' and ''OsGI–Hd1(Se1)'' pathways are activated under LD conditions; the former promotes the flowering of ''Arabidopsis'', whereas the latter inhibits the flowering of rice. The divergence in the photoperiodic pathways of these two species arises from the molecular mechanisms controlling the expression of ''FT/Hd3a'' by ''CO/Hd1(Se1)''(from reference <ref name="ref5" />).]]<br />
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==Labs working on this gene==<br />
*Photosynthesis and Photobiology Research Unit, National Institute of Agrobiological Sciences, 2-1-2 Kannondai, Tsukuba 305-8602, Japan<br />
*Laboratory of Plant Molecular Genetics, Nara Institute of Science and Technology, 8916-5 Takayama, Ikoma 630-0101, Japan<br />
*Plant Genomics Research Unit, National Institute of Agrobiological Sciences, 2-1-2 Kannondai, Tsukuba 305-8602, Japan<br />
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==Knowledge Extension==<br />
*The rice orthologues of the two key regulators of flowering time in Arabidopsis, ''CO'' and ''FT'', were recently shown to be important in the photoperiodic control of flowering in rice. The rice ''Hd1(Se1)''gene, which is an orthologue of the Arabidopsis ''CO'' gene, was required for the suppression of flowering under LD conditions and for the promotion of flowering under SD conditions. Furthermore, the rice ''Hd3a'' gene, an orthologue of the Arabidopsis ''FT'' gene, was shown to be an activator of flowering in rice. It was previously demonstrated that ''CO'' controls the flowering time by integrating signals from the circadian clock and light to regulate ''FT'' expression in Arabidopsis<ref name="ref5" />.Unlike Arabidopsis ''FT'', expression of ''Hd3a'' mRNA is more or less opposite to that of ''Hd1'' under inductive short-day conditions<ref name="ref6" />.<br />
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==References==<br />
<references><br />
<ref name="ref1">Song Lim Kim, Shinyoung Lee, Hyo Jung Kim, Hong Gil Nam, Gynheung An（2007）''OsMADS51'' is a short-day flowering promoter that functions upstream of ''Ehd1'', ''OsMADS14'', and ''Hd3a''.''Plant Physiology'' 145(4): 1484-1494.</ref><br />
<ref name="ref2">Hironori Itoh, Takeshi Izawa (2011) A study of phytohormone biosynthetic gene expression using a circadian clock-related mutant in rice. ''Plant Signaling & Behavior'' 6(12): 1932-1936.</ref> <br />
<ref name="ref3">Ryousuke Hayama, Takeshi Izawa, Ko Shimamoto (2002) Isolation of rice genes possibly involved in the photoperiodic control of flowering by a<br />
fluorescent differential display method. ''Plant and Cell Physiology'' 43(5): 494-504.</ref> <br />
<ref name="ref4">Takeshi Izawa, Motohiro Mihara, Yuji Suzuki, Meenu Gupta, Hironori Itoh, et al. (2011) ''Os-GIGANTEA'' confers robust diurnal rhythms on the global transcriptome of rice in the field. ''The Plant Cell'' 23(5): 1741-1755.</ref> <br />
<ref name="ref5">Ryosuke Hayama, Shuji Yokoi, Shojiro Tamaki, Masahiro Yano, Ko Shimamoto (2003) Adaptation of photoperiodic control pathways produces short-day flowering in rice. ''Nature'' 422(6933): 719-722.</ref> <br />
<ref name="ref6">Bong-Soo Shin, Jeong-Hyun Lee, Jeong-Hwan Lee, Hyun-Joo Jeong, Choong-Hyo Yun, Jeong-Kook Kim (2004) Circadian regulation of rice (''Oryza sativa L.'') ''CONSTANS-like'' gene transcripts. ''Molecules and Cells'' 17(1): 10-16.</ref><br />
</references><br />
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==Structured Information==<br />
{{JaponicaGene|<br />
GeneName = Os01g0182600|<br />
Description = GIGANTEA protein|<br />
Version = NM_001048755.1 GI:115434923 GeneID:4325329|<br />
Length = 9083 bp|<br />
Definition = Oryza sativa Japonica Group Os01g0182600, complete gene.|<br />
Source = Oryza sativa Japonica Group<br />
<br />
ORGANISM Oryza sativa Japonica Group<br />
Eukaryota; Viridiplantae; Streptophyta; Embryophyta; Tracheophyta;<br />
Spermatophyta; Magnoliophyta; Liliopsida; Poales; Poaceae; BEP<br />
clade; Ehrhartoideae; Oryzeae; Oryza.<br />
|<br />
Chromosome = [[:category:Japonica Chromosome 1|Chromosome 1]]|<br />
AP = Chromosome 1:4328362..4337444|<br />
CDS = 4328725..4329030,4329123..4329215,4329333..4329551,4329752..4329820,4329925..4331421<br>,4332220..4332429,4332563..4332713,4332791..4333031,4333898..4333956<br>,4334379..4334653,4334813..4334955,4335034..4335088,4335181..4335258<br>,4335351..4335437|<br />
GCID = <gbrowseImage1><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage1>|<br />
GSID = <gbrowseImage2><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage2>|<br />
CDNA = <cdnaseq>atgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactgcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcatagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcggcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctga</cdnaseq>|<br />
AA = <aaseq>MSASNEKWIDGLQFSSLFWPPPQDSQQKQAQILAYVEYFGQFTA DSEQFPEDIAQLIQSCYPSKEKRLVDEVLATFVLHHPEHGHAVVHPILSRIIDGTLSY DRNGFPFMSFISLFSHTSEKEYSEQWALACGEILRVLTHYNRPIFKVDHQHSEAECSS TSDQASSCESMEKRANGSPRNEPDRKPLRPLSPWITDILLAAPLGIRSDYFRWCGGVM GKYAAGGELKPPTTAYSRGSGKHPQLMPSTPRWAVANGAGVILSVCDEEVARYETANL TAAAVPALLLPPPTTPLDEHLVAGLPPLEPYARLFHRYYAIATPSATQRLLFGLLEAP PSWAPDALDAAVQLVELLRAAEDYDSGMRLPKNWMHLHFLRAIGTAMSMRAGIAADTS AALLFRILSQPTLLFPPLRHAEGVELHHEPLGGYVSSYKRQLEVPASEATIDATAQGI ASMLCAHGPDVEWRICTIWEAAYGLLPLSSSAVDLPEIVVAAPLQPPTLSWSLYLPLL KVFEYLPRGSPSEACLMRIFVATVEAILRRTFPSETSEQSRKPRSQSKNLAVAELRTM IHSLFVESCASMDLASRLLFVVLTVCVSHQALPGGSKRPTGSDNHSSEEVTNDSRLTN GRNRCKKRQGPVATFDSYVLAAVCALSCELQLFPFISKNGNHSNLKDSIKIVIPGKTT GISNELHNSISSAILHTRRILGILEALFSLKPSSVGTSWSYSSNEIVAAAMVAAHVSE LFRRSRPCLNALSALKQCKWDAEISTRASSLYHLIDLHGKTVTSIVNKAEPLEAHLTL TPVKKDEPPIEEKNINSSDGGALEKKDASRSHRKNGFARPLLKCAEDVILNGDVASTS GKAIASLQVEASDLANFLTMDRNGGYRGSQTLLRSVLSEKQELCFSVVSLLWQKLIAS PEMQMSAESTSAHQGWRKVVDALCDIVSASPTKASAAIVLQAEKDLQPWIARDDEQGQ KMWRVNQRIVKLIAELMRNHDSPEALVILASASDLLLRATDGMLVDGEACTLPQLELL EVTARAVHLIVEWGDSGVSVADGLSNLLKCRLSTTIRCLSHPSAHVRALSMSVLRDIL NSGQINSSKLIQGEHRNGIQSPTYQCLAASIINWQADVERCIEWEAHSRRATGLTLAF LTAAAKELGCPLTC</aaseq>|<br />
DNA = <dnaseqindica>8415..8720#8230..8322#7894..8112#7625..7693#6024..7520#5016..5225#4732..4882#4414..4654#3489..3547#2792..3066#2490..2632#2357..2411#2187..2264#2008..2094#ccgagccgtttctctccctctctctctcctcctccttcccctgcgcctcctccaccgcccgtgtcgccgccgcgcgcgtctatcccctcgccgccgccgccgtctacgccgcgtgaggtccggattggttgaaaggctgatcgaggtaatgcttttttatttttcccgtgcggttttagtttttctgtgctgtgcttggtggttttttctttaaaactatttttaggctggttttgatgatggcctttctaaggttgtcgtgtatggggggcgaggtggttcgggtggatttgtggtgttatgttgaggggatttttgtttggggggagagagttttggtggtagatgcggttgggattttgaagacgtgaaagggaaattagttttgaaatttgacggcgaggttgagctgattagtattgtaataattcgtggcatcggatagatcaacgcgagatggatgcttattactgcgaatttcaggaggtttttgtggtttttttaggcgagtagcgttgattctagctggattttgctgcttcgtggaggcgtttagatgtggtgcatacggctgtggatgtaatttgtgcggggataattactgaaaattaattcgcagtaagctagctctgctttaatattttttcccctgaaacagtaacaagggatcccttcctcatttcgttaacagtggaagtgaatacagacaccaaccgatgagcagagacacctgtaattaaacaccaaaagaagaacctgttttgttcctatatattttccatttctatgctttaggtaatatactttctggttcggtgtcatagtacccgaaatagaatctttgcttgccaaaaaaagaaagaataacaagatagaacaacatcattaaattctgtcttagacaatacatttataatcgaccaacaaaaacagaccgatgagtgacgtggaataacaaatgaagtataaatgtcacgaaaaacacatctattgattttgtgtgataagccataggagcctacgaggaaccaagtgaatggttctgtacaccacatccattagcattaccataaatttgcctatttctatgatgccttccttaacatatttatcgacatgtgagcttgcaccataactaaatgttatggtaaactttaccttataaatgattgcaacatgtaccttttcatttcgataattaaaaacgagaaagcttccacctttttcctagatatattttttcgtttgattatgaatttagttgtacttgacaataaaataatgcgcttgtaagattatgcaattttcgttgcattacctccctctcttttcatttgtcagtttttttctactccctccgtttcatattttaagtcgtttgacttttttcctagtcagactttgttaagtttgacaaagtttatagtaaaatttagcaatatctaaaatattaaattggtttcatcaaatctaacattgaatatattttgataatatgttttttttgtgttgaaaatacttctatatttttctataaacttagtcaaacttggaagaaatttcagtaagaaaaaagtcaaacggcttataatatgaaacggagggagtaaatttttttttgcattaattaaaattcatgtattttcctccttttttggtttttgttgtgcgatatattacagtgtagagactctgaagctcatatggttggcattggttgatatgataacgtgtcacgttgtgagatgcagacccccgttatgggggttgattgaatgaagtggatagcactcttataactttcttgctcgaagtttcttgataccaaggcaagtagcttataagaagatttggactttggtggcataaatggatgaactaactggcggcttcaacgtggtgttgattacaggtgcttttattgctaactggagcagtttgtgtagtttctttctttctctttgagaattctctcagtttcttgggcaggctcttgagctaccaagtatgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggtaacattattcctatttgattcttgttagaaattctatatgttgaacatacttgcctcaatgggaaacttggttgcgtctcctctctgaaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcaggtactgtatctctaagcatgtttagtatcttctgtgataaaatctttctccagtgtgttaacttttgcattctttttcttgcatgcttgcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattaggtaatttgatccagtgatcaatacatgttccacaaacaattgaagaaagagtttgacatatttttctcatatttccagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgaggtaaggcccctactctaatggttagttccaatttattggttcatttctctgtgtttgataagattttgcctctgctgtgcatttcaatattatatctaaactgtgctgcttatagcttatcaaactactcttgctcataaagatgcaacttttatgcagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgagtcataataaatgataacatattattgttctgtggactgaaaatctgaaatcctcctttcttgagaaatacaacctcaattttatggtgatattgacaagaaggaagagcgcatataatttaaagttaaaacacggaaaataatagaagtatgtgagcagaacgcaggagcaaggggtggaactcataacaagttcaaacttcatgcatgttcataattccccaatgcaagtatggcaagaaaaatagacaacttttaaaagtatgctaatggaactgcatgcattgtgtaaacttgcttttgtcaagcatgttggaacatactatatatttgctgctgatgatgaaaaatttgcttgagtactcttctattgctaatggacgcttaccattccctgttttgtcttcaaatgcatcaggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactggttggtaaaatcagcaatggaattttctatcgtgaattttttataattattgtaacattaattactcccatgtctttttattctgttgagataccatgtgacaatgtctacaattgcatattttgcattctttggtagatcagttcccttcttaagatccccaaagggaaactggagtatcaataacttactccctcaaataaaaaaaaagtgatactttaggcataaacacattcttcaaaacaaaactttgaatagcagtttcttttgtaatatattaataaagcatggaaaaattataatgtgaagaatgtacattcagggacaaatatactcatcatttttgcatggttgatcttaatgtccatagagagattgatcagaccaaaagtttatttttgacaggagtgagtatggattataataggagtgaatggcaatatgctgtgaaagtgagcattttatatttgtttactactagcatttggtgacaaagcaaatgatgtatattgtaaatcaaagtaaacaagtagaacatgtcctgtggcgacaagacataacctacaaattaaagttagcctgattgtttattttactgggagcacattttttctagattctagagtacatattattaaaaaaggtgtgatatgtttgcaaattactttttaatggaagtttgtaaattactttgaaacttgcattcagtttttttgcttcttaaaaaacatagtgcatatgggctttatgagctcatactatgcatgtttttctgcattggagtgggacctttatctttttcttgcaactcggatgctaatgcttattataaacacaagttactaacatgtgcttgggtcattcagcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcataggtaagctcttatgatcctgtccactgttgtaccatgtgtactttccaaatgattgctaacatgctatttttcgtcagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtac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/>
Link = [http://www.ncbi.nlm.nih.gov/nuccore/NM_001048755.1 RefSeq:Os01g0182600]|<br />
}}<br />
[[Category:Genes]]<br />
[[Category:Japonica mRNA]]<br />
[[Category:Oryza Sativa Japonica Group]]<br />
[[Category:Japonica Genes]]<br />
[[Category:Japonica Chromosome 1]]<br />
[[Category:Chromosome 1]]</div>Mikahttps://ngdc.cncb.ac.cn/ricewiki/index.php?title=Os01g0182600&diff=183864Os01g01826002014-06-11T04:38:54Z<p>Mika: /* References */</p>
<hr />
<div>The rice gene of ''Gigantea(GI)'' is solo ortholog of ''GI'' in the Arabidopsis circadian system and related to photoperiodic control of flowering.<br />
<br />
==Annotated Information==<br />
===Function===<br />
*''OsGI'',a ''GI'' ortholog, regulates flowering time in response to photoperiodic conditions. Mutations in this gene delay flowering in inductive SDs (short-days),whereas flowering is marginally induced under LDs(long-days).In comparison, over expression of ''OsGI'' in transgenic rice causes late flowering in both SDs and LDs.In addition, ''OsGI'' has function to regulate ''Hd1'', a gene that also shows circadian rhythm<ref name="ref1" />. ''Os-GI'' affects gene expression of approximately 75% of genes among 27,201 genes and is required for strong amplitudes and proper phase-setting of global gene expression under natural field conditions<ref name="ref2" />. <br />
*The cDNA of ''OsGI'' firstly found in ''se5'' mutant that is completely photoperiod-insensitive, the expression patterns of rice and Arabidopsis ''GI'' gene are both photoperiodically regulated and are very similar under LD and SD conditions('''Figure 1''')<ref name="ref3" />. [[File:Limeijia2.png|frame|'''Figure 1''' Possible roles of isolated genes in the photoperiodic regulation of flowering in rice, including ''OsGI'', may function to inhibit flowering under LD conditions.(from reference <ref name="ref3" />).]]''GI'' is required for phase-dependent ''CO''transcription through the interaction with FKF1, which leads to cycling DNA binding with one finger (CDF) degradation and consequent derepression of CO. In addition, it has been known that GI protein interacts with ZEITLUPE (ZTL) F-box protein to control the diurnal rhythm of TOC1 protein in regulation of the Arabidopsis circadian clock<ref name="ref4" />. <br />
*''Os-GI'' also has function to maintain bioactive GA level through the regulation of the GA-deactivating enzyme genes in rice, ''Os-GI'' represses major ''OsGA2ox'' genes in order to maintain a proper bioactive GA level, ''osgi-1'' plants showed a semi-dwarf phenotype with elevated ''OsGA2ox'' gene expression('''Figure 2''') <ref name="ref4" /> .[[File:Limeijia1.png|frame|'''Figure 2''' WT rice and osgi-1 plants grown under natural field conditions and ''osgi-1'' plant exhibits Semi-dwarf phenotype(from reference <ref name="ref4" />).]]<br />
<br />
===Expression===<br />
*''GI'' encodes a nuclear protein that defines the proper amplitude and period length of circadian rhythms. ''GI'' expression is regulated by the circadian clock, with a peak in transcript levels at 8 to 10 h after dawn. The ''GI'' protein acts between the circadian oscillator and ''CO'' to promote flowering by increasing ''CO'' and ''FT'' mRNA abundance<ref name="ref1" />. The expression of the ''OsGI'' messenger RNA was circadian-controlled and that its temporal expression pattern was very similar to that of ''GI'' under both SD and LD conditions. Decreased ''OsGI'' expression led to early flowering under LD conditions and late flowering under SD conditions relative to the wild type, and that this early flowering phenotype under LD conditions was the opposite of that of the Arabidopsis ''gi'' mutants, confirming the opposite role of the ''GI'' gene in flowering under LD conditions in rice and Arabidopsis<ref name="ref5" />.<br />
<br />
===Mutation===<br />
*The single ''osgi-1'' mutation extended the time to flowering only under SD conditions. Under LD conditions the single ''hd1'' mutation induced early flowering, but ''osgi-1'' plants exhibited no phenotypic change<ref name="ref4" /><br />
<br />
===Evolution===<br />
* There is some evolutionary conservation between rice circadian clock–related genes and those associated with Arabidopsis circadian clocks, where the circadian clock consists of interlocked subloops, it is also possible that ''Os-GI'' is a member of a subloop in rice circadian clocks<ref name="ref4" />.But there is a divergence in the photoperiodic pathways of these two species arises from the molecular mechanisms controlling the expression of ''FT/Hd3a'' by ''CO/Hd1(Se1)'' ('''Figure 3''')<ref name="ref5" />[[File:Limeijia3.png|frame|'''Figure 3'''The molecular mechanisms for the regulation of ''GI/OsGI'' expression, and the regulation of ''CO/Hd1(Se1)'' expression by ''GI/OsGI'', might be conserved between ''Arabidopsis'' and rice. Both the ''GI–CO'' and ''OsGI–Hd1(Se1)'' pathways are activated under LD conditions; the former promotes the flowering of ''Arabidopsis'', whereas the latter inhibits the flowering of rice. The divergence in the photoperiodic pathways of these two species arises from the molecular mechanisms controlling the expression of ''FT/Hd3a'' by ''CO/Hd1(Se1)''(from reference <ref name="ref5" />).]]<br />
<br />
==Labs working on this gene==<br />
*Photosynthesis and Photobiology Research Unit, National Institute of Agrobiological Sciences, 2-1-2 Kannondai, Tsukuba 305-8602, Japan<br />
*Laboratory of Plant Molecular Genetics, Nara Institute of Science and Technology, 8916-5 Takayama, Ikoma 630-0101, Japan<br />
*Plant Genomics Research Unit, National Institute of Agrobiological Sciences, 2-1-2 Kannondai, Tsukuba 305-8602, Japan<br />
<br />
==Knowledge Extension==<br />
*The rice orthologues of the two key regulators of flowering time in Arabidopsis, ''CO'' and ''FT'', were recently shown to be important in the photoperiodic control of flowering in rice. The rice ''Hd1(Se1)''gene, which is an orthologue of the Arabidopsis ''CO'' gene, was required for the suppression of flowering under LD conditions and for the promotion of flowering under SD conditions. Furthermore, the rice ''Hd3a'' gene, an orthologue of the Arabidopsis ''FT'' gene, was shown to be an activator of flowering in rice. It was previously demonstrated that ''CO'' controls the flowering time by integrating signals from the circadian clock and light to regulate ''FT'' expression in Arabidopsis<ref name="ref5" />.Unlike Arabidopsis ''FT'', expression of ''Hd3a'' mRNA is more or less opposite to that of ''Hd1'' under inductive short-day conditions<ref name="ref6" />.<br />
<br />
==References==<br />
<references><br />
<ref name="ref1">Song Lim Kim, Shinyoung Lee, Hyo Jung Kim, Hong Gil Nam, Gynheung An（2007）''OsMADS51'' is a short-day flowering promoter that functions upstream of ''Ehd1'', ''OsMADS14'', and ''Hd3a''.''Plant Physiology'' 145(4): 1484-1494.</ref><br />
<ref name="ref2">Hironori Itoh, Takeshi Izawa (2011) A study of phytohormone biosynthetic gene expression using a circadian clock-related mutant in rice. ''Plant Signaling & Behavior'' 6(12): 1932-1936.</ref> <br />
<ref name="ref3">Ryousuke Hayama, Takeshi Izawa, Ko Shimamoto (2002) Isolation of rice genes possibly involved in the photoperiodic control of flowering by a<br />
fluorescent differential display method. ''Plant and Cell Physiology'' 43(5): 494-504.</ref> <br />
<ref name="ref4">Takeshi Izawa, Motohiro Mihara, Yuji Suzuki, Meenu Gupta, Hironori Itoh, et al. (2011) ''Os-GIGANTEA'' confers robust diurnal rhythms on the global transcriptome of rice in the field. ''The Plant Cell'' 23(5): 1741-1755.</ref> <br />
<ref name="ref5">Ryosuke Hayama, Shuji Yokoi, Shojiro Tamaki, Masahiro Yano, Ko Shimamoto (2003) Adaptation of photoperiodic control pathways produces short-day flowering in rice. ''Nature'' 422(6933): 719-722.</ref> <br />
<ref name="ref6">Bong-Soo Shin, Jeong-Hyun Lee, Jeong-Hwan Lee, Hyun-Joo Jeong, Choong-Hyo Yun, Jeong-Kook Kim (2004) Circadian regulation of rice (''Oryza sativa L.'') ''CONSTANS-like'' gene transcripts. ''Molecules and Cells'' 17(1): 10-16.</ref><br />
</references><br />
<br />
==Structured Information==<br />
{{JaponicaGene|<br />
GeneName = Os01g0182600|<br />
Description = GIGANTEA protein|<br />
Version = NM_001048755.1 GI:115434923 GeneID:4325329|<br />
Length = 9083 bp|<br />
Definition = Oryza sativa Japonica Group Os01g0182600, complete gene.|<br />
Source = Oryza sativa Japonica Group<br />
<br />
ORGANISM Oryza sativa Japonica Group<br />
Eukaryota; Viridiplantae; Streptophyta; Embryophyta; Tracheophyta;<br />
Spermatophyta; Magnoliophyta; Liliopsida; Poales; Poaceae; BEP<br />
clade; Ehrhartoideae; Oryzeae; Oryza.<br />
|<br />
Chromosome = [[:category:Japonica Chromosome 1|Chromosome 1]]|<br />
AP = Chromosome 1:4328362..4337444|<br />
CDS = 4328725..4329030,4329123..4329215,4329333..4329551,4329752..4329820,4329925..4331421<br>,4332220..4332429,4332563..4332713,4332791..4333031,4333898..4333956<br>,4334379..4334653,4334813..4334955,4335034..4335088,4335181..4335258<br>,4335351..4335437|<br />
GCID = <gbrowseImage1><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage1>|<br />
GSID = <gbrowseImage2><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage2>|<br />
CDNA = <cdnaseq>atgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactgcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcatagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcggcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctga</cdnaseq>|<br />
AA = <aaseq>MSASNEKWIDGLQFSSLFWPPPQDSQQKQAQILAYVEYFGQFTA DSEQFPEDIAQLIQSCYPSKEKRLVDEVLATFVLHHPEHGHAVVHPILSRIIDGTLSY DRNGFPFMSFISLFSHTSEKEYSEQWALACGEILRVLTHYNRPIFKVDHQHSEAECSS TSDQASSCESMEKRANGSPRNEPDRKPLRPLSPWITDILLAAPLGIRSDYFRWCGGVM GKYAAGGELKPPTTAYSRGSGKHPQLMPSTPRWAVANGAGVILSVCDEEVARYETANL TAAAVPALLLPPPTTPLDEHLVAGLPPLEPYARLFHRYYAIATPSATQRLLFGLLEAP PSWAPDALDAAVQLVELLRAAEDYDSGMRLPKNWMHLHFLRAIGTAMSMRAGIAADTS AALLFRILSQPTLLFPPLRHAEGVELHHEPLGGYVSSYKRQLEVPASEATIDATAQGI ASMLCAHGPDVEWRICTIWEAAYGLLPLSSSAVDLPEIVVAAPLQPPTLSWSLYLPLL KVFEYLPRGSPSEACLMRIFVATVEAILRRTFPSETSEQSRKPRSQSKNLAVAELRTM IHSLFVESCASMDLASRLLFVVLTVCVSHQALPGGSKRPTGSDNHSSEEVTNDSRLTN GRNRCKKRQGPVATFDSYVLAAVCALSCELQLFPFISKNGNHSNLKDSIKIVIPGKTT GISNELHNSISSAILHTRRILGILEALFSLKPSSVGTSWSYSSNEIVAAAMVAAHVSE LFRRSRPCLNALSALKQCKWDAEISTRASSLYHLIDLHGKTVTSIVNKAEPLEAHLTL TPVKKDEPPIEEKNINSSDGGALEKKDASRSHRKNGFARPLLKCAEDVILNGDVASTS GKAIASLQVEASDLANFLTMDRNGGYRGSQTLLRSVLSEKQELCFSVVSLLWQKLIAS PEMQMSAESTSAHQGWRKVVDALCDIVSASPTKASAAIVLQAEKDLQPWIARDDEQGQ KMWRVNQRIVKLIAELMRNHDSPEALVILASASDLLLRATDGMLVDGEACTLPQLELL EVTARAVHLIVEWGDSGVSVADGLSNLLKCRLSTTIRCLSHPSAHVRALSMSVLRDIL NSGQINSSKLIQGEHRNGIQSPTYQCLAASIINWQADVERCIEWEAHSRRATGLTLAF LTAAAKELGCPLTC</aaseq>|<br />
DNA = <dnaseqindica>8415..8720#8230..8322#7894..8112#7625..7693#6024..7520#5016..5225#4732..4882#4414..4654#3489..3547#2792..3066#2490..2632#2357..2411#2187..2264#2008..2094#ccgagccgtttctctccctctctctctcctcctccttcccctgcgcctcctccaccgcccgtgtcgccgccgcgcgcgtctatcccctcgccgccgccgccgtctacgccgcgtgaggtccggattggttgaaaggctgatcgaggtaatgcttttttatttttcccgtgcggttttagtttttctgtgctgtgcttggtggttttttctttaaaactatttttaggctggttttgatgatggcctttctaaggttgtcgtgtatggggggcgaggtggttcgggtggatttgtggtgttatgttgaggggatttttgtttggggggagagagttttggtggtagatgcggttgggattttgaagacgtgaaagggaaattagttttgaaatttgacggcgaggttgagctgattagtattgtaataattcgtggcatcggatagatcaacgcgagatggatgcttattactgcgaatttcaggaggtttttgtggtttttttaggcgagtagcgttgattctagctggattttgctgcttcgtggaggcgtttagatgtggtgcatacggctgtggatgtaatttgtgcggggataattactgaaaattaattcgcagtaagctagctctgctttaatattttttcccctgaaacagtaacaagggatcccttcctcatttcgttaacagtggaagtgaatacagacaccaaccgatgagcagagacacctgtaattaaacaccaaaagaagaacctgttttgttcctatatattttccatttctatgctttaggtaatatactttctggttcggtgtcatagtacccgaaatagaatctttgcttgccaaaaaaagaaagaataacaagatagaacaacatcattaaattctgtcttagacaatacatttataatcgaccaacaaaaacagaccgatgagtgacgtggaataacaaatgaagtataaatgtcacgaaaaacacatctattgattttgtgtgataagccataggagcctacgaggaaccaagtgaatggttctgtacaccacatccattagcattaccataaatttgcctatttctatgatgccttccttaacatatttatcgacatgtgagcttgcaccataactaaatgttatggtaaactttaccttataaatgattgcaacatgtaccttttcatttcgataattaaaaacgagaaagcttccacctttttcctagatatattttttcgtttgattatgaatttagttgtacttgacaataaaataatgcgcttgtaagattatgcaattttcgttgcattacctccctctcttttcatttgtcagtttttttctactccctccgtttcatattttaagtcgtttgacttttttcctagtcagactttgttaagtttgacaaagtttatagtaaaatttagcaatatctaaaatattaaattggtttcatcaaatctaacattgaatatattttgataatatgttttttttgtgttgaaaatacttctatatttttctataaacttagtcaaacttggaagaaatttcagtaagaaaaaagtcaaacggcttataatatgaaacggagggagtaaatttttttttgcattaattaaaattcatgtattttcctccttttttggtttttgttgtgcgatatattacagtgtagagactctgaagctcatatggttggcattggttgatatgataacgtgtcacgttgtgagatgcagacccccgttatgggggttgattgaatgaagtggatagcactcttataactttcttgctcgaagtttcttgataccaaggcaagtagcttataagaagatttggactttggtggcataaatggatgaactaactggcggcttcaacgtggtgttgattacaggtgcttttattgctaactggagcagtttgtgtagtttctttctttctctttgagaattctctcagtttcttgggcaggctcttgagctaccaagtatgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggtaacattattcctatttgattcttgttagaaattctatatgttgaacatacttgcctcaatgggaaacttggttgcgtctcctctctgaaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcaggtactgtatctctaagcatgtttagtatcttctgtgataaaatctttctccagtgtgttaacttttgcattctttttcttgcatgcttgcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattaggtaatttgatccagtgatcaatacatgttccacaaacaattgaagaaagagtttgacatatttttctcatatttccagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgaggtaaggcccctactctaatggttagttccaatttattggttcatttctctgtgtttgataagattttgcctctgctgtgcatttcaatattatatctaaactgtgctgcttatagcttatcaaactactcttgctcataaagatgcaacttttatgcagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgagtcataataaatgataacatattattgttctgtggactgaaaatctgaaatcctcctttcttgagaaatacaacctcaattttatggtgatattgacaagaaggaagagcgcatataatttaaagttaaaacacggaaaataatagaagtatgtgagcagaacgcaggagcaaggggtggaactcataacaagttcaaacttcatgcatgttcataattccccaatgcaagtatggcaagaaaaatagacaacttttaaaagtatgctaatggaactgcatgcattgtgtaaacttgcttttgtcaagcatgttggaacatactatatatttgctgctgatgatgaaaaatttgcttgagtactcttctattgctaatggacgcttaccattccctgttttgtcttcaaatgcatcaggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactggttggtaaaatcagcaatggaattttctatcgtgaattttttataattattgtaacattaattactcccatgtctttttattctgttgagataccatgtgacaatgtctacaattgcatattttgcattctttggtagatcagttcccttcttaagatccccaaagggaaactggagtatcaataacttactccctcaaataaaaaaaaagtgatactttaggcataaacacattcttcaaaacaaaactttgaatagcagtttcttttgtaatatattaataaagcatggaaaaattataatgtgaagaatgtacattcagggacaaatatactcatcatttttgcatggttgatcttaatgtccatagagagattgatcagaccaaaagtttatttttgacaggagtgagtatggattataataggagtgaatggcaatatgctgtgaaagtgagcattttatatttgtttactactagcatttggtgacaaagcaaatgatgtatattgtaaatcaaagtaaacaagtagaacatgtcctgtggcgacaagacataacctacaaattaaagttagcctgattgtttattttactgggagcacattttttctagattctagagtacatattattaaaaaaggtgtgatatgtttgcaaattactttttaatggaagtttgtaaattactttgaaacttgcattcagtttttttgcttcttaaaaaacatagtgcatatgggctttatgagctcatactatgcatgtttttctgcattggagtgggacctttatctttttcttgcaactcggatgctaatgcttattataaacacaagttactaacatgtgcttgggtcattcagcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcataggtaagctcttatgatcctgtccactgttgtaccatgtgtactttccaaatgattgctaacatgctatttttcgtcagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcgggtatatcatacatctctctggctgttgtttcagtgctatcaatcaagatgacacatcatgcaatattatcataacagcacattatgtgttttgaatattttattgattgtcatagctttgttttatgttttagcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcaggtatgcagtgtttcctgcacctacttgctacctattctgtatttgatgtattcaatttcctttttatcagttttctcgaaatatatgctaataggcttgtcactaaacggctacattatccagactctgttctgaatttttttttcattaagtactgttagctgccagattacatcatcatgactaaagtagtttagctacaatattgcaaagctagatttgtctagcctctattgttttgaatagaaaaggcaggtatgcagtgtttcctgcacctacttgctacctattctgtatttgatgtattcaatttcctttttatcagttttctcgaaatatatgctaataggcttgtcactaaacggctacattatccagactctgttctgaatttttttttcattaagtactgttagctgccagattacatcatcatgactaaagtagtttagctacaatattgcaaagctagatttgtctagcctctattgttttgaatagaaaattaaatcttttggtcatttttttgtaaccttttaggagtctctgtatgcttgtcgtcaacatgctatgataagtttatatcagttcttttggagctatgatgtattcttgtatattttatgactgctaggatattatgttaaaactgttttaaaggtatcatcaaacaagttaatatagatattgctacctaccaaagttggaatttgattatgtcgtaaaatatatgcaccctacatgttttagcttcatattgacttatattttcatatgtgctccttcaactgcacaaagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggtatggtcattcctgcttataagataccagaataatgctatgaagaccgtccttggttgtcagcattcttatacacaatgataatctttgtttttcatgggtaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggtcaagatattgttattgttatatttacatacttgttatcatatttaaaagcttcatattgtatgtacctttgaaaaaaaaagttccatactgtagcttcttttgttctcatttcttggtttcaaagcatcagcatcattaccgatataaaatatacccatttcctcagtcatcttccccgactcttgtatctttcacaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagaggtaacaccttcattcatgaggtccttaggtgcaaatcccagtgaagaataatcagttctactttgtatggttcatctcgcttactgaccggttcatctcgggatcaattacgagcagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaaggcatgtcctcttctctgatctcactcacgttcatgccaaacttactcatcatcatccatgatccatcatcctgactgaatcttgatatgcagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctgacaaggccacaccatgactgtcactgcaagcagctgctgtaggatcagcgagtaggaaaggatgacttgccggccagtttcctgagatgtgatttaagactgatattagctgatgttcccaagcatgatacggggcttgctcccaagatgtgatttttactttacttttggttaatgtgctaccgctgacattagatgttcaagcatatgaaaactgcttgtgctgtaacttgtatgtctgtaacgtaaaaatctttccatatggtaaccggggaagcagaggtatatccttttctgctgttgtatctatttccagtttcctgctgaacttgcaaatgaatattcagcaatttccgttttgctc</dnaseqindica>|<br />
Link = [http://www.ncbi.nlm.nih.gov/nuccore/NM_001048755.1 RefSeq:Os01g0182600]|<br />
}}<br />
[[Category:Genes]]<br />
[[Category:Japonica mRNA]]<br />
[[Category:Oryza Sativa Japonica Group]]<br />
[[Category:Japonica Genes]]<br />
[[Category:Japonica Chromosome 1]]<br />
[[Category:Chromosome 1]]</div>Mikahttps://ngdc.cncb.ac.cn/ricewiki/index.php?title=Os01g0182600&diff=183860Os01g01826002014-06-11T04:29:05Z<p>Mika: /* Evolution */</p>
<hr />
<div>The rice gene of ''Gigantea(GI)'' is solo ortholog of ''GI'' in the Arabidopsis circadian system and related to photoperiodic control of flowering.<br />
<br />
==Annotated Information==<br />
===Function===<br />
*''OsGI'',a ''GI'' ortholog, regulates flowering time in response to photoperiodic conditions. Mutations in this gene delay flowering in inductive SDs (short-days),whereas flowering is marginally induced under LDs(long-days).In comparison, over expression of ''OsGI'' in transgenic rice causes late flowering in both SDs and LDs.In addition, ''OsGI'' has function to regulate ''Hd1'', a gene that also shows circadian rhythm<ref name="ref1" />. ''Os-GI'' affects gene expression of approximately 75% of genes among 27,201 genes and is required for strong amplitudes and proper phase-setting of global gene expression under natural field conditions<ref name="ref2" />. <br />
*The cDNA of ''OsGI'' firstly found in ''se5'' mutant that is completely photoperiod-insensitive, the expression patterns of rice and Arabidopsis ''GI'' gene are both photoperiodically regulated and are very similar under LD and SD conditions('''Figure 1''')<ref name="ref3" />. [[File:Limeijia2.png|frame|'''Figure 1''' Possible roles of isolated genes in the photoperiodic regulation of flowering in rice, including ''OsGI'', may function to inhibit flowering under LD conditions.(from reference <ref name="ref3" />).]]''GI'' is required for phase-dependent ''CO''transcription through the interaction with FKF1, which leads to cycling DNA binding with one finger (CDF) degradation and consequent derepression of CO. In addition, it has been known that GI protein interacts with ZEITLUPE (ZTL) F-box protein to control the diurnal rhythm of TOC1 protein in regulation of the Arabidopsis circadian clock<ref name="ref4" />. <br />
*''Os-GI'' also has function to maintain bioactive GA level through the regulation of the GA-deactivating enzyme genes in rice, ''Os-GI'' represses major ''OsGA2ox'' genes in order to maintain a proper bioactive GA level, ''osgi-1'' plants showed a semi-dwarf phenotype with elevated ''OsGA2ox'' gene expression('''Figure 2''') <ref name="ref4" /> .[[File:Limeijia1.png|frame|'''Figure 2''' WT rice and osgi-1 plants grown under natural field conditions and ''osgi-1'' plant exhibits Semi-dwarf phenotype(from reference <ref name="ref4" />).]]<br />
<br />
===Expression===<br />
*''GI'' encodes a nuclear protein that defines the proper amplitude and period length of circadian rhythms. ''GI'' expression is regulated by the circadian clock, with a peak in transcript levels at 8 to 10 h after dawn. The ''GI'' protein acts between the circadian oscillator and ''CO'' to promote flowering by increasing ''CO'' and ''FT'' mRNA abundance<ref name="ref1" />. The expression of the ''OsGI'' messenger RNA was circadian-controlled and that its temporal expression pattern was very similar to that of ''GI'' under both SD and LD conditions. Decreased ''OsGI'' expression led to early flowering under LD conditions and late flowering under SD conditions relative to the wild type, and that this early flowering phenotype under LD conditions was the opposite of that of the Arabidopsis ''gi'' mutants, confirming the opposite role of the ''GI'' gene in flowering under LD conditions in rice and Arabidopsis<ref name="ref5" />.<br />
<br />
===Mutation===<br />
*The single ''osgi-1'' mutation extended the time to flowering only under SD conditions. Under LD conditions the single ''hd1'' mutation induced early flowering, but ''osgi-1'' plants exhibited no phenotypic change<ref name="ref4" /><br />
<br />
===Evolution===<br />
* There is some evolutionary conservation between rice circadian clock–related genes and those associated with Arabidopsis circadian clocks, where the circadian clock consists of interlocked subloops, it is also possible that ''Os-GI'' is a member of a subloop in rice circadian clocks<ref name="ref4" />.But there is a divergence in the photoperiodic pathways of these two species arises from the molecular mechanisms controlling the expression of ''FT/Hd3a'' by ''CO/Hd1(Se1)'' ('''Figure 3''')<ref name="ref5" />[[File:Limeijia3.png|frame|'''Figure 3'''The molecular mechanisms for the regulation of ''GI/OsGI'' expression, and the regulation of ''CO/Hd1(Se1)'' expression by ''GI/OsGI'', might be conserved between ''Arabidopsis'' and rice. Both the ''GI–CO'' and ''OsGI–Hd1(Se1)'' pathways are activated under LD conditions; the former promotes the flowering of ''Arabidopsis'', whereas the latter inhibits the flowering of rice. The divergence in the photoperiodic pathways of these two species arises from the molecular mechanisms controlling the expression of ''FT/Hd3a'' by ''CO/Hd1(Se1)''(from reference <ref name="ref5" />).]]<br />
<br />
==Labs working on this gene==<br />
*Photosynthesis and Photobiology Research Unit, National Institute of Agrobiological Sciences, 2-1-2 Kannondai, Tsukuba 305-8602, Japan<br />
*Laboratory of Plant Molecular Genetics, Nara Institute of Science and Technology, 8916-5 Takayama, Ikoma 630-0101, Japan<br />
*Plant Genomics Research Unit, National Institute of Agrobiological Sciences, 2-1-2 Kannondai, Tsukuba 305-8602, Japan<br />
<br />
==Knowledge Extension==<br />
*The rice orthologues of the two key regulators of flowering time in Arabidopsis, ''CO'' and ''FT'', were recently shown to be important in the photoperiodic control of flowering in rice. The rice ''Hd1(Se1)''gene, which is an orthologue of the Arabidopsis ''CO'' gene, was required for the suppression of flowering under LD conditions and for the promotion of flowering under SD conditions. Furthermore, the rice ''Hd3a'' gene, an orthologue of the Arabidopsis ''FT'' gene, was shown to be an activator of flowering in rice. It was previously demonstrated that ''CO'' controls the flowering time by integrating signals from the circadian clock and light to regulate ''FT'' expression in Arabidopsis<ref name="ref5" />.Unlike Arabidopsis ''FT'', expression of ''Hd3a'' mRNA is more or less opposite to that of ''Hd1'' under inductive short-day conditions<ref name="ref6" />.<br />
<br />
==References==<br />
<references><br />
<ref name="ref1">Song Lim Kim, Shinyoung Lee, Hyo Jung Kim, Hong Gil Nam, Gynheung An（2007）''OsMADS51'' is a short-day flowering promoter that functions upstream of ''Ehd1'', ''OsMADS14'', and ''Hd3a''.''Plant Physiology'' 145(4): 1484-1494.</ref><br />
<ref name="ref2">Hironori Itoh, Takeshi Izawa (2011) A study of phytohormone biosynthetic gene expression using a circadian clock-related mutant in rice. ''Plant Signaling & Behavior'' 6(12): 1932-1936.</ref> <br />
<ref name="ref3">Ryousuke Hayama, Takeshi Izawa, Ko Shimamoto (2002) Isolation of rice genes possibly involved in the photoperiodic control of flowering by a<br />
fluorescent differential display method. ''Plant and Cell Physiology'' 43(5): 494-504.</ref> <br />
<ref name="ref4">Takeshi Izawa, Motohiro Mihara, Yuji Suzuki, Meenu Gupta, Hironori Itoh, et al. (2011) ''Os-GIGANTEA'' confers robust diurnal rhythms on the global transcriptome of rice in the field. ''The Plant Cell'' 23(5): 1741-1755.</ref> <br />
<ref name="ref5">Ryosuke Hayama, Shuji Yokoi, Shojiro Tamaki, Masahiro Yano, Ko Shimamoto (2003) Adaptation of photoperiodic control pathways produces short-day flowering in rice. ''Nature'' 422(6933): 719-722.</ref> <br />
<ref name="ref6">Bong-Soo Shin, Jeong-Hyun Lee, Jeong-Hwan Lee, Hyun-Joo Jeong, Choong-Hyo Yun, Jeong-Kook Kim (2004) Circadian regulation of rice (''Oryza sativa L.'') ''CONSTANS-like'' gene transcripts. ''Molecules and Cells'' 17(1): 10-16.</ref><br />
<br />
==Structured Information==<br />
{{JaponicaGene|<br />
GeneName = Os01g0182600|<br />
Description = GIGANTEA protein|<br />
Version = NM_001048755.1 GI:115434923 GeneID:4325329|<br />
Length = 9083 bp|<br />
Definition = Oryza sativa Japonica Group Os01g0182600, complete gene.|<br />
Source = Oryza sativa Japonica Group<br />
<br />
ORGANISM Oryza sativa Japonica Group<br />
Eukaryota; Viridiplantae; Streptophyta; Embryophyta; Tracheophyta;<br />
Spermatophyta; Magnoliophyta; Liliopsida; Poales; Poaceae; BEP<br />
clade; Ehrhartoideae; Oryzeae; Oryza.<br />
|<br />
Chromosome = [[:category:Japonica Chromosome 1|Chromosome 1]]|<br />
AP = Chromosome 1:4328362..4337444|<br />
CDS = 4328725..4329030,4329123..4329215,4329333..4329551,4329752..4329820,4329925..4331421<br>,4332220..4332429,4332563..4332713,4332791..4333031,4333898..4333956<br>,4334379..4334653,4334813..4334955,4335034..4335088,4335181..4335258<br>,4335351..4335437|<br />
GCID = <gbrowseImage1><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage1>|<br />
GSID = <gbrowseImage2><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage2>|<br />
CDNA = <cdnaseq>atgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactgcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcatagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcggcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctga</cdnaseq>|<br />
AA = <aaseq>MSASNEKWIDGLQFSSLFWPPPQDSQQKQAQILAYVEYFGQFTA DSEQFPEDIAQLIQSCYPSKEKRLVDEVLATFVLHHPEHGHAVVHPILSRIIDGTLSY DRNGFPFMSFISLFSHTSEKEYSEQWALACGEILRVLTHYNRPIFKVDHQHSEAECSS TSDQASSCESMEKRANGSPRNEPDRKPLRPLSPWITDILLAAPLGIRSDYFRWCGGVM GKYAAGGELKPPTTAYSRGSGKHPQLMPSTPRWAVANGAGVILSVCDEEVARYETANL TAAAVPALLLPPPTTPLDEHLVAGLPPLEPYARLFHRYYAIATPSATQRLLFGLLEAP PSWAPDALDAAVQLVELLRAAEDYDSGMRLPKNWMHLHFLRAIGTAMSMRAGIAADTS AALLFRILSQPTLLFPPLRHAEGVELHHEPLGGYVSSYKRQLEVPASEATIDATAQGI ASMLCAHGPDVEWRICTIWEAAYGLLPLSSSAVDLPEIVVAAPLQPPTLSWSLYLPLL KVFEYLPRGSPSEACLMRIFVATVEAILRRTFPSETSEQSRKPRSQSKNLAVAELRTM IHSLFVESCASMDLASRLLFVVLTVCVSHQALPGGSKRPTGSDNHSSEEVTNDSRLTN GRNRCKKRQGPVATFDSYVLAAVCALSCELQLFPFISKNGNHSNLKDSIKIVIPGKTT GISNELHNSISSAILHTRRILGILEALFSLKPSSVGTSWSYSSNEIVAAAMVAAHVSE LFRRSRPCLNALSALKQCKWDAEISTRASSLYHLIDLHGKTVTSIVNKAEPLEAHLTL TPVKKDEPPIEEKNINSSDGGALEKKDASRSHRKNGFARPLLKCAEDVILNGDVASTS GKAIASLQVEASDLANFLTMDRNGGYRGSQTLLRSVLSEKQELCFSVVSLLWQKLIAS PEMQMSAESTSAHQGWRKVVDALCDIVSASPTKASAAIVLQAEKDLQPWIARDDEQGQ KMWRVNQRIVKLIAELMRNHDSPEALVILASASDLLLRATDGMLVDGEACTLPQLELL EVTARAVHLIVEWGDSGVSVADGLSNLLKCRLSTTIRCLSHPSAHVRALSMSVLRDIL NSGQINSSKLIQGEHRNGIQSPTYQCLAASIINWQADVERCIEWEAHSRRATGLTLAF LTAAAKELGCPLTC</aaseq>|<br />
DNA = <dnaseqindica>8415..8720#8230..8322#7894..8112#7625..7693#6024..7520#5016..5225#4732..4882#4414..4654#3489..3547#2792..3066#2490..2632#2357..2411#2187..2264#2008..2094#ccgagccgtttctctccctctctctctcctcctccttcccctgcgcctcctccaccgcccgtgtcgccgccgcgcgcgtctatcccctcgccgccgccgccgtctacgccgcgtgaggtccggattggttgaaaggctgatcgaggtaatgcttttttatttttcccgtgcggttttagtttttctgtgctgtgcttggtggttttttctttaaaactatttttaggctggttttgatgatggcctttctaaggttgtcgtgtatggggggcgaggtggttcgggtggatttgtggtgttatgttgaggggatttttgtttggggggagagagttttggtggtagatgcggttgggattttgaagacgtgaaagggaaattagttttgaaatttgacggcgaggttgagctgattagtattgtaataattcgtggcatcggatagatcaacgcgagatggatgcttattactgcgaatttcaggaggtttttgtggtttttttaggcgagtagcgttgattctagctggattttgctgcttcgtggaggcgtttagatgtggtgcatacggctgtggatgtaatttgtgcggggataattactgaaaattaattcgcagtaagctagctctgctttaatattttttcccctgaaacagtaacaagggatcccttcctcatttcgttaacagtggaagtgaatacagacaccaaccgatgagcagagacacctgtaattaaacaccaaaagaagaacctgttttgttcctatatattttccatttctatgctttaggtaatatactttctggttcggtgtcatagtacccgaaatagaatctttgcttgccaaaaaaagaaagaataacaagatagaacaacatcattaaattctgtcttagacaatacatttataatcgaccaacaaaaacagaccgatgagtgacgtggaataacaaatgaagtataaatgtcacgaaaaacacatctattgattttgtgtgataagccataggagcctacgaggaaccaagtgaatggttctgtacaccacatccattagcattaccataaatttgcctatttctatgatgccttccttaacatatttatcgacatgtgagcttgcaccataactaaatgttatggtaaactttaccttataaatgattgcaacatgtaccttttcatttcgataattaaaaacgagaaagcttccacctttttcctagatatattttttcgtttgattatgaatttagttgtacttgacaataaaataatgcgcttgtaagattatgcaattttcgttgcattacctccctctcttttcatttgtcagtttttttctactccctccgtttcatattttaagtcgtttgacttttttcctagtcagactttgttaagtttgacaaagtttatagtaaaatttagcaatatctaaaatattaaattggtttcatcaaatctaacattgaatatattttgataatatgttttttttgtgttgaaaatacttctatatttttctataaacttagtcaaacttggaagaaatttcagtaagaaaaaagtcaaacggcttataatatgaaacggagggagtaaatttttttttgcattaattaaaattcatgtattttcctccttttttggtttttgttgtgcgatatattacagtgtagagactctgaagctcatatggttggcattggttgatatgataacgtgtcacgttgtgagatgcagacccccgttatgggggttgattgaatgaagtggatagcactcttataactttcttgctcgaagtttcttgataccaaggcaagtagcttataagaagatttggactttggtggcataaatggatgaactaactggcggcttcaacgtggtgttgattacaggtgcttttattgctaactggagcagtttgtgtagtttctttctttctctttgagaattctctcagtttcttgggcaggctcttgagctaccaagtatgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggtaacattattcctatttgattcttgttagaaattctatatgttgaacatacttgcctcaatgggaaacttggttgcgtctcctctctgaaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcaggtactgtatctctaagcatgtttagtatcttctgtgataaaatctttctccagtgtgttaacttttgcattctttttcttgcatgcttgcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattaggtaatttgatccagtgatcaatacatgttccacaaacaattgaagaaagagtttgacatatttttctcatatttccagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgaggtaaggcccctactctaatggttagttccaatttattggttcatttctctgtgtttgataagattttgcctctgctgtgcatttcaatattatatctaaactgtgctgcttatagcttatcaaactactcttgctcataaagatgcaacttttatgcagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgagtcataataaatgataacatattattgttctgtggactgaaaatctgaaatcctcctttcttgagaaatacaacctcaattttatggtgatattgacaagaaggaagagcgcatataatttaaagttaaaacacggaaaataatagaagtatgtgagcagaacgcaggagcaaggggtggaactcataacaagttcaaacttcatgcatgttcataattccccaatgcaagtatggcaagaaaaatagacaacttttaaaagtatgctaatggaactgcatgcattgtgtaaacttgcttttgtcaagcatgttggaacatactatatatttgctgctgatgatgaaaaatttgcttgagtactcttctattgctaatggacgcttaccattccctgttttgtcttcaaatgcatcaggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactggttggtaaaatcagcaatggaattttctatcgtgaattttttataattattgtaacattaattactcccatgtctttttattctgttgagataccatgtgacaatgtctacaattgcatattttgcattctttggtagatcagttcccttcttaagatccccaaagggaaactggagtatcaataacttactccctcaaataaaaaaaaagtgatactttaggcataaacacattcttcaaaacaaaactttgaatagcagtttcttttgtaatatattaataaagcatggaaaaattataatgtgaagaatgtacattcagggacaaatatactcatcatttttgcatggttgatcttaatgtccatagagagattgatcagaccaaaagtttatttttgacaggagtgagtatggattataataggagtgaatggcaatatgctgtgaaagtgagcattttatatttgtttactactagcatttggtgacaaagcaaatgatgtatattgtaaatcaaagtaaacaagtagaacatgtcctgtggcgacaagacataacctacaaattaaagttagcctgattgtttattttactgggagcacattttttctagattctagagtacatattattaaaaaaggtgtgatatgtttgcaaattactttttaatggaagtttgtaaattactttgaaacttgcattcagtttttttgcttcttaaaaaacatagtgcatatgggctttatgagctcatactatgcatgtttttctgcattggagtgggacctttatctttttcttgcaactcggatgctaatgcttattataaacacaagttactaacatgtgcttgggtcattcagcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcataggtaagctcttatgatcctgtccactgttgtaccatgtgtactttccaaatgattgctaacatgctatttttcgtcagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtac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/>
Link = [http://www.ncbi.nlm.nih.gov/nuccore/NM_001048755.1 RefSeq:Os01g0182600]|<br />
}}<br />
[[Category:Genes]]<br />
[[Category:Japonica mRNA]]<br />
[[Category:Oryza Sativa Japonica Group]]<br />
[[Category:Japonica Genes]]<br />
[[Category:Japonica Chromosome 1]]<br />
[[Category:Chromosome 1]]</div>Mikahttps://ngdc.cncb.ac.cn/ricewiki/index.php?title=Os01g0182600&diff=183859Os01g01826002014-06-11T04:28:22Z<p>Mika: /* Evolution */</p>
<hr />
<div>The rice gene of ''Gigantea(GI)'' is solo ortholog of ''GI'' in the Arabidopsis circadian system and related to photoperiodic control of flowering.<br />
<br />
==Annotated Information==<br />
===Function===<br />
*''OsGI'',a ''GI'' ortholog, regulates flowering time in response to photoperiodic conditions. Mutations in this gene delay flowering in inductive SDs (short-days),whereas flowering is marginally induced under LDs(long-days).In comparison, over expression of ''OsGI'' in transgenic rice causes late flowering in both SDs and LDs.In addition, ''OsGI'' has function to regulate ''Hd1'', a gene that also shows circadian rhythm<ref name="ref1" />. ''Os-GI'' affects gene expression of approximately 75% of genes among 27,201 genes and is required for strong amplitudes and proper phase-setting of global gene expression under natural field conditions<ref name="ref2" />. <br />
*The cDNA of ''OsGI'' firstly found in ''se5'' mutant that is completely photoperiod-insensitive, the expression patterns of rice and Arabidopsis ''GI'' gene are both photoperiodically regulated and are very similar under LD and SD conditions('''Figure 1''')<ref name="ref3" />. [[File:Limeijia2.png|frame|'''Figure 1''' Possible roles of isolated genes in the photoperiodic regulation of flowering in rice, including ''OsGI'', may function to inhibit flowering under LD conditions.(from reference <ref name="ref3" />).]]''GI'' is required for phase-dependent ''CO''transcription through the interaction with FKF1, which leads to cycling DNA binding with one finger (CDF) degradation and consequent derepression of CO. In addition, it has been known that GI protein interacts with ZEITLUPE (ZTL) F-box protein to control the diurnal rhythm of TOC1 protein in regulation of the Arabidopsis circadian clock<ref name="ref4" />. <br />
*''Os-GI'' also has function to maintain bioactive GA level through the regulation of the GA-deactivating enzyme genes in rice, ''Os-GI'' represses major ''OsGA2ox'' genes in order to maintain a proper bioactive GA level, ''osgi-1'' plants showed a semi-dwarf phenotype with elevated ''OsGA2ox'' gene expression('''Figure 2''') <ref name="ref4" /> .[[File:Limeijia1.png|frame|'''Figure 2''' WT rice and osgi-1 plants grown under natural field conditions and ''osgi-1'' plant exhibits Semi-dwarf phenotype(from reference <ref name="ref4" />).]]<br />
<br />
===Expression===<br />
*''GI'' encodes a nuclear protein that defines the proper amplitude and period length of circadian rhythms. ''GI'' expression is regulated by the circadian clock, with a peak in transcript levels at 8 to 10 h after dawn. The ''GI'' protein acts between the circadian oscillator and ''CO'' to promote flowering by increasing ''CO'' and ''FT'' mRNA abundance<ref name="ref1" />. The expression of the ''OsGI'' messenger RNA was circadian-controlled and that its temporal expression pattern was very similar to that of ''GI'' under both SD and LD conditions. Decreased ''OsGI'' expression led to early flowering under LD conditions and late flowering under SD conditions relative to the wild type, and that this early flowering phenotype under LD conditions was the opposite of that of the Arabidopsis ''gi'' mutants, confirming the opposite role of the ''GI'' gene in flowering under LD conditions in rice and Arabidopsis<ref name="ref5" />.<br />
<br />
===Mutation===<br />
*The single ''osgi-1'' mutation extended the time to flowering only under SD conditions. Under LD conditions the single ''hd1'' mutation induced early flowering, but ''osgi-1'' plants exhibited no phenotypic change<ref name="ref4" /><br />
<br />
===Evolution===<br />
* There is some evolutionary conservation between rice circadian clock–related genes and those associated with Arabidopsis circadian clocks, where the circadian clock consists of interlocked subloops, it is also possible that ''Os-GI'' is a member of a subloop in rice circadian clocks<ref name="ref4" />.But there is a divergence in the photoperiodic pathways of these two species arises from the molecular mechanisms controlling the expression of ''FT/Hd3a'' by ''CO/Hd1(Se1)'' ('''Figure3''')<ref name="ref5" />[[File:Limeijia3.png|frame|'''Figure 3'''The molecular mechanisms for the regulation of ''GI/OsGI'' expression, and the regulation of ''CO/Hd1(Se1)'' expression by ''GI/OsGI'', might be conserved between ''Arabidopsis'' and rice. Both the ''GI–CO'' and ''OsGI–Hd1(Se1)'' pathways are activated under LD conditions; the former promotes the flowering of ''Arabidopsis'', whereas the latter inhibits the flowering of rice. The divergence in the photoperiodic pathways of these two species arises from the molecular mechanisms controlling the expression of ''FT/Hd3a'' by ''CO/Hd1(Se1)''(from reference <ref name="ref5" />).]]<br />
<br />
==Labs working on this gene==<br />
*Photosynthesis and Photobiology Research Unit, National Institute of Agrobiological Sciences, 2-1-2 Kannondai, Tsukuba 305-8602, Japan<br />
*Laboratory of Plant Molecular Genetics, Nara Institute of Science and Technology, 8916-5 Takayama, Ikoma 630-0101, Japan<br />
*Plant Genomics Research Unit, National Institute of Agrobiological Sciences, 2-1-2 Kannondai, Tsukuba 305-8602, Japan<br />
<br />
==Knowledge Extension==<br />
*The rice orthologues of the two key regulators of flowering time in Arabidopsis, ''CO'' and ''FT'', were recently shown to be important in the photoperiodic control of flowering in rice. The rice ''Hd1(Se1)''gene, which is an orthologue of the Arabidopsis ''CO'' gene, was required for the suppression of flowering under LD conditions and for the promotion of flowering under SD conditions. Furthermore, the rice ''Hd3a'' gene, an orthologue of the Arabidopsis ''FT'' gene, was shown to be an activator of flowering in rice. It was previously demonstrated that ''CO'' controls the flowering time by integrating signals from the circadian clock and light to regulate ''FT'' expression in Arabidopsis<ref name="ref5" />.Unlike Arabidopsis ''FT'', expression of ''Hd3a'' mRNA is more or less opposite to that of ''Hd1'' under inductive short-day conditions<ref name="ref6" />.<br />
<br />
==References==<br />
<references><br />
<ref name="ref1">Song Lim Kim, Shinyoung Lee, Hyo Jung Kim, Hong Gil Nam, Gynheung An（2007）''OsMADS51'' is a short-day flowering promoter that functions upstream of ''Ehd1'', ''OsMADS14'', and ''Hd3a''.''Plant Physiology'' 145(4): 1484-1494.</ref><br />
<ref name="ref2">Hironori Itoh, Takeshi Izawa (2011) A study of phytohormone biosynthetic gene expression using a circadian clock-related mutant in rice. ''Plant Signaling & Behavior'' 6(12): 1932-1936.</ref> <br />
<ref name="ref3">Ryousuke Hayama, Takeshi Izawa, Ko Shimamoto (2002) Isolation of rice genes possibly involved in the photoperiodic control of flowering by a<br />
fluorescent differential display method. ''Plant and Cell Physiology'' 43(5): 494-504.</ref> <br />
<ref name="ref4">Takeshi Izawa, Motohiro Mihara, Yuji Suzuki, Meenu Gupta, Hironori Itoh, et al. (2011) ''Os-GIGANTEA'' confers robust diurnal rhythms on the global transcriptome of rice in the field. ''The Plant Cell'' 23(5): 1741-1755.</ref> <br />
<ref name="ref5">Ryosuke Hayama, Shuji Yokoi, Shojiro Tamaki, Masahiro Yano, Ko Shimamoto (2003) Adaptation of photoperiodic control pathways produces short-day flowering in rice. ''Nature'' 422(6933): 719-722.</ref> <br />
<ref name="ref6">Bong-Soo Shin, Jeong-Hyun Lee, Jeong-Hwan Lee, Hyun-Joo Jeong, Choong-Hyo Yun, Jeong-Kook Kim (2004) Circadian regulation of rice (''Oryza sativa L.'') ''CONSTANS-like'' gene transcripts. ''Molecules and Cells'' 17(1): 10-16.</ref><br />
<br />
==Structured Information==<br />
{{JaponicaGene|<br />
GeneName = Os01g0182600|<br />
Description = GIGANTEA protein|<br />
Version = NM_001048755.1 GI:115434923 GeneID:4325329|<br />
Length = 9083 bp|<br />
Definition = Oryza sativa Japonica Group Os01g0182600, complete gene.|<br />
Source = Oryza sativa Japonica Group<br />
<br />
ORGANISM Oryza sativa Japonica Group<br />
Eukaryota; Viridiplantae; Streptophyta; Embryophyta; Tracheophyta;<br />
Spermatophyta; Magnoliophyta; Liliopsida; Poales; Poaceae; BEP<br />
clade; Ehrhartoideae; Oryzeae; Oryza.<br />
|<br />
Chromosome = [[:category:Japonica Chromosome 1|Chromosome 1]]|<br />
AP = Chromosome 1:4328362..4337444|<br />
CDS = 4328725..4329030,4329123..4329215,4329333..4329551,4329752..4329820,4329925..4331421<br>,4332220..4332429,4332563..4332713,4332791..4333031,4333898..4333956<br>,4334379..4334653,4334813..4334955,4335034..4335088,4335181..4335258<br>,4335351..4335437|<br />
GCID = <gbrowseImage1><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage1>|<br />
GSID = <gbrowseImage2><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage2>|<br />
CDNA = <cdnaseq>atgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactgcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcatagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcggcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctga</cdnaseq>|<br />
AA = <aaseq>MSASNEKWIDGLQFSSLFWPPPQDSQQKQAQILAYVEYFGQFTA DSEQFPEDIAQLIQSCYPSKEKRLVDEVLATFVLHHPEHGHAVVHPILSRIIDGTLSY DRNGFPFMSFISLFSHTSEKEYSEQWALACGEILRVLTHYNRPIFKVDHQHSEAECSS TSDQASSCESMEKRANGSPRNEPDRKPLRPLSPWITDILLAAPLGIRSDYFRWCGGVM GKYAAGGELKPPTTAYSRGSGKHPQLMPSTPRWAVANGAGVILSVCDEEVARYETANL TAAAVPALLLPPPTTPLDEHLVAGLPPLEPYARLFHRYYAIATPSATQRLLFGLLEAP PSWAPDALDAAVQLVELLRAAEDYDSGMRLPKNWMHLHFLRAIGTAMSMRAGIAADTS AALLFRILSQPTLLFPPLRHAEGVELHHEPLGGYVSSYKRQLEVPASEATIDATAQGI ASMLCAHGPDVEWRICTIWEAAYGLLPLSSSAVDLPEIVVAAPLQPPTLSWSLYLPLL KVFEYLPRGSPSEACLMRIFVATVEAILRRTFPSETSEQSRKPRSQSKNLAVAELRTM IHSLFVESCASMDLASRLLFVVLTVCVSHQALPGGSKRPTGSDNHSSEEVTNDSRLTN GRNRCKKRQGPVATFDSYVLAAVCALSCELQLFPFISKNGNHSNLKDSIKIVIPGKTT GISNELHNSISSAILHTRRILGILEALFSLKPSSVGTSWSYSSNEIVAAAMVAAHVSE LFRRSRPCLNALSALKQCKWDAEISTRASSLYHLIDLHGKTVTSIVNKAEPLEAHLTL TPVKKDEPPIEEKNINSSDGGALEKKDASRSHRKNGFARPLLKCAEDVILNGDVASTS GKAIASLQVEASDLANFLTMDRNGGYRGSQTLLRSVLSEKQELCFSVVSLLWQKLIAS PEMQMSAESTSAHQGWRKVVDALCDIVSASPTKASAAIVLQAEKDLQPWIARDDEQGQ KMWRVNQRIVKLIAELMRNHDSPEALVILASASDLLLRATDGMLVDGEACTLPQLELL EVTARAVHLIVEWGDSGVSVADGLSNLLKCRLSTTIRCLSHPSAHVRALSMSVLRDIL NSGQINSSKLIQGEHRNGIQSPTYQCLAASIINWQADVERCIEWEAHSRRATGLTLAF LTAAAKELGCPLTC</aaseq>|<br />
DNA = <dnaseqindica>8415..8720#8230..8322#7894..8112#7625..7693#6024..7520#5016..5225#4732..4882#4414..4654#3489..3547#2792..3066#2490..2632#2357..2411#2187..2264#2008..2094#ccgagccgtttctctccctctctctctcctcctccttcccctgcgcctcctccaccgcccgtgtcgccgccgcgcgcgtctatcccctcgccgccgccgccgtctacgccgcgtgaggtccggattggttgaaaggctgatcgaggtaatgcttttttatttttcccgtgcggttttagtttttctgtgctgtgcttggtggttttttctttaaaactatttttaggctggttttgatgatggcctttctaaggttgtcgtgtatggggggcgaggtggttcgggtggatttgtggtgttatgttgaggggatttttgtttggggggagagagttttggtggtagatgcggttgggattttgaagacgtgaaagggaaattagttttgaaatttgacggcgaggttgagctgattagtattgtaataattcgtggcatcggatagatcaacgcgagatggatgcttattactgcgaatttcaggaggtttttgtggtttttttaggcgagtagcgttgattctagctggattttgctgcttcgtggaggcgtttagatgtggtgcatacggctgtggatgtaatttgtgcggggataattactgaaaattaattcgcagtaagctagctctgctttaatattttttcccctgaaacagtaacaagggatcccttcctcatttcgttaacagtggaagtgaatacagacaccaaccgatgagcagagacacctgtaattaaacaccaaaagaagaacctgttttgttcctatatattttccatttctatgctttaggtaatatactttctggttcggtgtcatagtacccgaaatagaatctttgcttgccaaaaaaagaaagaataacaagatagaacaacatcattaaattctgtcttagacaatacatttataatcgaccaacaaaaacagaccgatgagtgacgtggaataacaaatgaagtataaatgtcacgaaaaacacatctattgattttgtgtgataagccataggagcctacgaggaaccaagtgaatggttctgtacaccacatccattagcattaccataaatttgcctatttctatgatgccttccttaacatatttatcgacatgtgagcttgcaccataactaaatgttatggtaaactttaccttataaatgattgcaacatgtaccttttcatttcgataattaaaaacgagaaagcttccacctttttcctagatatattttttcgtttgattatgaatttagttgtacttgacaataaaataatgcgcttgtaagattatgcaattttcgttgcattacctccctctcttttcatttgtcagtttttttctactccctccgtttcatattttaagtcgtttgacttttttcctagtcagactttgttaagtttgacaaagtttatagtaaaatttagcaatatctaaaatattaaattggtttcatcaaatctaacattgaatatattttgataatatgttttttttgtgttgaaaatacttctatatttttctataaacttagtcaaacttggaagaaatttcagtaagaaaaaagtcaaacggcttataatatgaaacggagggagtaaatttttttttgcattaattaaaattcatgtattttcctccttttttggtttttgttgtgcgatatattacagtgtagagactctgaagctcatatggttggcattggttgatatgataacgtgtcacgttgtgagatgcagacccccgttatgggggttgattgaatgaagtggatagcactcttataactttcttgctcgaagtttcttgataccaaggcaagtagcttataagaagatttggactttggtggcataaatggatgaactaactggcggcttcaacgtggtgttgattacaggtgcttttattgctaactggagcagtttgtgtagtttctttctttctctttgagaattctctcagtttcttgggcaggctcttgagctaccaagtatgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggtaacattattcctatttgattcttgttagaaattctatatgttgaacatacttgcctcaatgggaaacttggttgcgtctcctctctgaaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcaggtactgtatctctaagcatgtttagtatcttctgtgataaaatctttctccagtgtgttaacttttgcattctttttcttgcatgcttgcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattaggtaatttgatccagtgatcaatacatgttccacaaacaattgaagaaagagtttgacatatttttctcatatttccagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgaggtaaggcccctactctaatggttagttccaatttattggttcatttctctgtgtttgataagattttgcctctgctgtgcatttcaatattatatctaaactgtgctgcttatagcttatcaaactactcttgctcataaagatgcaacttttatgcagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgagtcataataaatgataacatattattgttctgtggactgaaaatctgaaatcctcctttcttgagaaatacaacctcaattttatggtgatattgacaagaaggaagagcgcatataatttaaagttaaaacacggaaaataatagaagtatgtgagcagaacgcaggagcaaggggtggaactcataacaagttcaaacttcatgcatgttcataattccccaatgcaagtatggcaagaaaaatagacaacttttaaaagtatgctaatggaactgcatgcattgtgtaaacttgcttttgtcaagcatgttggaacatactatatatttgctgctgatgatgaaaaatttgcttgagtactcttctattgctaatggacgcttaccattccctgttttgtcttcaaatgcatcaggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactggttggtaaaatcagcaatggaattttctatcgtgaattttttataattattgtaacattaattactcccatgtctttttattctgttgagataccatgtgacaatgtctacaattgcatattttgcattctttggtagatcagttcccttcttaagatccccaaagggaaactggagtatcaataacttactccctcaaataaaaaaaaagtgatactttaggcataaacacattcttcaaaacaaaactttgaatagcagtttcttttgtaatatattaataaagcatggaaaaattataatgtgaagaatgtacattcagggacaaatatactcatcatttttgcatggttgatcttaatgtccatagagagattgatcagaccaaaagtttatttttgacaggagtgagtatggattataataggagtgaatggcaatatgctgtgaaagtgagcattttatatttgtttactactagcatttggtgacaaagcaaatgatgtatattgtaaatcaaagtaaacaagtagaacatgtcctgtggcgacaagacataacctacaaattaaagttagcctgattgtttattttactgggagcacattttttctagattctagagtacatattattaaaaaaggtgtgatatgtttgcaaattactttttaatggaagtttgtaaattactttgaaacttgcattcagtttttttgcttcttaaaaaacatagtgcatatgggctttatgagctcatactatgcatgtttttctgcattggagtgggacctttatctttttcttgcaactcggatgctaatgcttattataaacacaagttactaacatgtgcttgggtcattcagcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcataggtaagctcttatgatcctgtccactgttgtaccatgtgtactttccaaatgattgctaacatgctatttttcgtcagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcgggtatatcatacatctctctggctgttgtttcagtgctatcaatcaagatgacacatcatgcaatattatcataacagcacattatgtgttttgaatattttattgattgtcatagctttgttttatgttttagcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcaggtatgcagtgtttcctgcacctacttgctacctattctgtatttgatgtattcaatttcctttttatcagttttctcgaaatatatgctaataggcttgtcactaaacggctacattatccagactctgttctgaatttttttttcattaagtactgttagctgccagattacatcatcatgactaaagtagtttagctacaatattgcaaagctagatttgtctagcctctattgttttgaatagaaaaggcaggtatgcagtgtttcctgcacctacttgctacctattctgtatttgatgtattcaatttcctttttatcagttttctcgaaatatatgctaataggcttgtcactaaacggctacattatccagactctgttctgaatttttttttcattaagtactgttagctgccagattacatcatcatgactaaagtagtttagctacaatattgcaaagctagatttgtctagcctctattgttttgaatagaaaattaaatcttttggtcatttttttgtaaccttttaggagtctctgtatgcttgtcgtcaacatgctatgataagtttatatcagttcttttggagctatgatgtattcttgtatattttatgactgctaggatattatgttaaaactgttttaaaggtatcatcaaacaagttaatatagatattgctacctaccaaagttggaatttgattatgtcgtaaaatatatgcaccctacatgttttagcttcatattgacttatattttcatatgtgctccttcaactgcacaaagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggtatggtcattcctgcttataagataccagaataatgctatgaagaccgtccttggttgtcagcattcttatacacaatgataatctttgtttttcatgggtaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggtcaagatattgttattgttatatttacatacttgttatcatatttaaaagcttcatattgtatgtacctttgaaaaaaaaagttccatactgtagcttcttttgttctcatttcttggtttcaaagcatcagcatcattaccgatataaaatatacccatttcctcagtcatcttccccgactcttgtatctttcacaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagaggtaacaccttcattcatgaggtccttaggtgcaaatcccagtgaagaataatcagttctactttgtatggttcatctcgcttactgaccggttcatctcgggatcaattacgagcagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaaggcatgtcctcttctctgatctcactcacgttcatgccaaacttactcatcatcatccatgatccatcatcctgactgaatcttgatatgcagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctgacaaggccacaccatgactgtcactgcaagcagctgctgtaggatcagcgagtaggaaaggatgacttgccggccagtttcctgagatgtgatttaagactgatattagctgatgttcccaagcatgatacggggcttgctcccaagatgtgatttttactttacttttggttaatgtgctaccgctgacattagatgttcaagcatatgaaaactgcttgtgctgtaacttgtatgtctgtaacgtaaaaatctttccatatggtaaccggggaagcagaggtatatccttttctgctgttgtatctatttccagtttcctgctgaacttgcaaatgaatattcagcaatttccgttttgctc</dnaseqindica>|<br />
Link = [http://www.ncbi.nlm.nih.gov/nuccore/NM_001048755.1 RefSeq:Os01g0182600]|<br />
}}<br />
[[Category:Genes]]<br />
[[Category:Japonica mRNA]]<br />
[[Category:Oryza Sativa Japonica Group]]<br />
[[Category:Japonica Genes]]<br />
[[Category:Japonica Chromosome 1]]<br />
[[Category:Chromosome 1]]</div>Mikahttps://ngdc.cncb.ac.cn/ricewiki/index.php?title=Os01g0182600&diff=183858Os01g01826002014-06-11T04:27:51Z<p>Mika: /* Function */</p>
<hr />
<div>The rice gene of ''Gigantea(GI)'' is solo ortholog of ''GI'' in the Arabidopsis circadian system and related to photoperiodic control of flowering.<br />
<br />
==Annotated Information==<br />
===Function===<br />
*''OsGI'',a ''GI'' ortholog, regulates flowering time in response to photoperiodic conditions. Mutations in this gene delay flowering in inductive SDs (short-days),whereas flowering is marginally induced under LDs(long-days).In comparison, over expression of ''OsGI'' in transgenic rice causes late flowering in both SDs and LDs.In addition, ''OsGI'' has function to regulate ''Hd1'', a gene that also shows circadian rhythm<ref name="ref1" />. ''Os-GI'' affects gene expression of approximately 75% of genes among 27,201 genes and is required for strong amplitudes and proper phase-setting of global gene expression under natural field conditions<ref name="ref2" />. <br />
*The cDNA of ''OsGI'' firstly found in ''se5'' mutant that is completely photoperiod-insensitive, the expression patterns of rice and Arabidopsis ''GI'' gene are both photoperiodically regulated and are very similar under LD and SD conditions('''Figure 1''')<ref name="ref3" />. [[File:Limeijia2.png|frame|'''Figure 1''' Possible roles of isolated genes in the photoperiodic regulation of flowering in rice, including ''OsGI'', may function to inhibit flowering under LD conditions.(from reference <ref name="ref3" />).]]''GI'' is required for phase-dependent ''CO''transcription through the interaction with FKF1, which leads to cycling DNA binding with one finger (CDF) degradation and consequent derepression of CO. In addition, it has been known that GI protein interacts with ZEITLUPE (ZTL) F-box protein to control the diurnal rhythm of TOC1 protein in regulation of the Arabidopsis circadian clock<ref name="ref4" />. <br />
*''Os-GI'' also has function to maintain bioactive GA level through the regulation of the GA-deactivating enzyme genes in rice, ''Os-GI'' represses major ''OsGA2ox'' genes in order to maintain a proper bioactive GA level, ''osgi-1'' plants showed a semi-dwarf phenotype with elevated ''OsGA2ox'' gene expression('''Figure 2''') <ref name="ref4" /> .[[File:Limeijia1.png|frame|'''Figure 2''' WT rice and osgi-1 plants grown under natural field conditions and ''osgi-1'' plant exhibits Semi-dwarf phenotype(from reference <ref name="ref4" />).]]<br />
<br />
===Expression===<br />
*''GI'' encodes a nuclear protein that defines the proper amplitude and period length of circadian rhythms. ''GI'' expression is regulated by the circadian clock, with a peak in transcript levels at 8 to 10 h after dawn. The ''GI'' protein acts between the circadian oscillator and ''CO'' to promote flowering by increasing ''CO'' and ''FT'' mRNA abundance<ref name="ref1" />. The expression of the ''OsGI'' messenger RNA was circadian-controlled and that its temporal expression pattern was very similar to that of ''GI'' under both SD and LD conditions. Decreased ''OsGI'' expression led to early flowering under LD conditions and late flowering under SD conditions relative to the wild type, and that this early flowering phenotype under LD conditions was the opposite of that of the Arabidopsis ''gi'' mutants, confirming the opposite role of the ''GI'' gene in flowering under LD conditions in rice and Arabidopsis<ref name="ref5" />.<br />
<br />
===Mutation===<br />
*The single ''osgi-1'' mutation extended the time to flowering only under SD conditions. Under LD conditions the single ''hd1'' mutation induced early flowering, but ''osgi-1'' plants exhibited no phenotypic change<ref name="ref4" /><br />
<br />
===Evolution===<br />
* There is some evolutionary conservation between rice circadian clock–related genes and those associated with Arabidopsis circadian clocks, where the circadian clock consists of interlocked subloops, it is also possible that ''Os-GI'' is a member of a subloop in rice circadian clocks<ref name="ref4" />.But there is a divergence in the photoperiodic pathways of these two species arises from the molecular mechanisms controlling the expression of ''FT/Hd3a'' by ''CO/Hd1(Se1)'' ['''Figure3''']<ref name="ref5" />[[File:Limeijia3.png|frame|'''Figure 3'''The molecular mechanisms for the regulation of ''GI/OsGI'' expression, and the regulation of ''CO/Hd1(Se1)'' expression by ''GI/OsGI'', might be conserved between ''Arabidopsis'' and rice. Both the ''GI–CO'' and ''OsGI–Hd1(Se1)'' pathways are activated under LD conditions; the former promotes the flowering of ''Arabidopsis'', whereas the latter inhibits the flowering of rice. The divergence in the photoperiodic pathways of these two species arises from the molecular mechanisms controlling the expression of ''FT/Hd3a'' by ''CO/Hd1(Se1)''(from reference <ref name="ref5" />).]]<br />
<br />
==Labs working on this gene==<br />
*Photosynthesis and Photobiology Research Unit, National Institute of Agrobiological Sciences, 2-1-2 Kannondai, Tsukuba 305-8602, Japan<br />
*Laboratory of Plant Molecular Genetics, Nara Institute of Science and Technology, 8916-5 Takayama, Ikoma 630-0101, Japan<br />
*Plant Genomics Research Unit, National Institute of Agrobiological Sciences, 2-1-2 Kannondai, Tsukuba 305-8602, Japan<br />
<br />
==Knowledge Extension==<br />
*The rice orthologues of the two key regulators of flowering time in Arabidopsis, ''CO'' and ''FT'', were recently shown to be important in the photoperiodic control of flowering in rice. The rice ''Hd1(Se1)''gene, which is an orthologue of the Arabidopsis ''CO'' gene, was required for the suppression of flowering under LD conditions and for the promotion of flowering under SD conditions. Furthermore, the rice ''Hd3a'' gene, an orthologue of the Arabidopsis ''FT'' gene, was shown to be an activator of flowering in rice. It was previously demonstrated that ''CO'' controls the flowering time by integrating signals from the circadian clock and light to regulate ''FT'' expression in Arabidopsis<ref name="ref5" />.Unlike Arabidopsis ''FT'', expression of ''Hd3a'' mRNA is more or less opposite to that of ''Hd1'' under inductive short-day conditions<ref name="ref6" />.<br />
<br />
==References==<br />
<references><br />
<ref name="ref1">Song Lim Kim, Shinyoung Lee, Hyo Jung Kim, Hong Gil Nam, Gynheung An（2007）''OsMADS51'' is a short-day flowering promoter that functions upstream of ''Ehd1'', ''OsMADS14'', and ''Hd3a''.''Plant Physiology'' 145(4): 1484-1494.</ref><br />
<ref name="ref2">Hironori Itoh, Takeshi Izawa (2011) A study of phytohormone biosynthetic gene expression using a circadian clock-related mutant in rice. ''Plant Signaling & Behavior'' 6(12): 1932-1936.</ref> <br />
<ref name="ref3">Ryousuke Hayama, Takeshi Izawa, Ko Shimamoto (2002) Isolation of rice genes possibly involved in the photoperiodic control of flowering by a<br />
fluorescent differential display method. ''Plant and Cell Physiology'' 43(5): 494-504.</ref> <br />
<ref name="ref4">Takeshi Izawa, Motohiro Mihara, Yuji Suzuki, Meenu Gupta, Hironori Itoh, et al. (2011) ''Os-GIGANTEA'' confers robust diurnal rhythms on the global transcriptome of rice in the field. ''The Plant Cell'' 23(5): 1741-1755.</ref> <br />
<ref name="ref5">Ryosuke Hayama, Shuji Yokoi, Shojiro Tamaki, Masahiro Yano, Ko Shimamoto (2003) Adaptation of photoperiodic control pathways produces short-day flowering in rice. ''Nature'' 422(6933): 719-722.</ref> <br />
<ref name="ref6">Bong-Soo Shin, Jeong-Hyun Lee, Jeong-Hwan Lee, Hyun-Joo Jeong, Choong-Hyo Yun, Jeong-Kook Kim (2004) Circadian regulation of rice (''Oryza sativa L.'') ''CONSTANS-like'' gene transcripts. ''Molecules and Cells'' 17(1): 10-16.</ref><br />
<br />
==Structured Information==<br />
{{JaponicaGene|<br />
GeneName = Os01g0182600|<br />
Description = GIGANTEA protein|<br />
Version = NM_001048755.1 GI:115434923 GeneID:4325329|<br />
Length = 9083 bp|<br />
Definition = Oryza sativa Japonica Group Os01g0182600, complete gene.|<br />
Source = Oryza sativa Japonica Group<br />
<br />
ORGANISM Oryza sativa Japonica Group<br />
Eukaryota; Viridiplantae; Streptophyta; Embryophyta; Tracheophyta;<br />
Spermatophyta; Magnoliophyta; Liliopsida; Poales; Poaceae; BEP<br />
clade; Ehrhartoideae; Oryzeae; Oryza.<br />
|<br />
Chromosome = [[:category:Japonica Chromosome 1|Chromosome 1]]|<br />
AP = Chromosome 1:4328362..4337444|<br />
CDS = 4328725..4329030,4329123..4329215,4329333..4329551,4329752..4329820,4329925..4331421<br>,4332220..4332429,4332563..4332713,4332791..4333031,4333898..4333956<br>,4334379..4334653,4334813..4334955,4335034..4335088,4335181..4335258<br>,4335351..4335437|<br />
GCID = <gbrowseImage1><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage1>|<br />
GSID = <gbrowseImage2><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage2>|<br />
CDNA = <cdnaseq>atgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactgcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcatagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcggcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctga</cdnaseq>|<br />
AA = <aaseq>MSASNEKWIDGLQFSSLFWPPPQDSQQKQAQILAYVEYFGQFTA DSEQFPEDIAQLIQSCYPSKEKRLVDEVLATFVLHHPEHGHAVVHPILSRIIDGTLSY DRNGFPFMSFISLFSHTSEKEYSEQWALACGEILRVLTHYNRPIFKVDHQHSEAECSS TSDQASSCESMEKRANGSPRNEPDRKPLRPLSPWITDILLAAPLGIRSDYFRWCGGVM GKYAAGGELKPPTTAYSRGSGKHPQLMPSTPRWAVANGAGVILSVCDEEVARYETANL TAAAVPALLLPPPTTPLDEHLVAGLPPLEPYARLFHRYYAIATPSATQRLLFGLLEAP PSWAPDALDAAVQLVELLRAAEDYDSGMRLPKNWMHLHFLRAIGTAMSMRAGIAADTS AALLFRILSQPTLLFPPLRHAEGVELHHEPLGGYVSSYKRQLEVPASEATIDATAQGI ASMLCAHGPDVEWRICTIWEAAYGLLPLSSSAVDLPEIVVAAPLQPPTLSWSLYLPLL KVFEYLPRGSPSEACLMRIFVATVEAILRRTFPSETSEQSRKPRSQSKNLAVAELRTM IHSLFVESCASMDLASRLLFVVLTVCVSHQALPGGSKRPTGSDNHSSEEVTNDSRLTN GRNRCKKRQGPVATFDSYVLAAVCALSCELQLFPFISKNGNHSNLKDSIKIVIPGKTT GISNELHNSISSAILHTRRILGILEALFSLKPSSVGTSWSYSSNEIVAAAMVAAHVSE LFRRSRPCLNALSALKQCKWDAEISTRASSLYHLIDLHGKTVTSIVNKAEPLEAHLTL TPVKKDEPPIEEKNINSSDGGALEKKDASRSHRKNGFARPLLKCAEDVILNGDVASTS GKAIASLQVEASDLANFLTMDRNGGYRGSQTLLRSVLSEKQELCFSVVSLLWQKLIAS PEMQMSAESTSAHQGWRKVVDALCDIVSASPTKASAAIVLQAEKDLQPWIARDDEQGQ KMWRVNQRIVKLIAELMRNHDSPEALVILASASDLLLRATDGMLVDGEACTLPQLELL EVTARAVHLIVEWGDSGVSVADGLSNLLKCRLSTTIRCLSHPSAHVRALSMSVLRDIL NSGQINSSKLIQGEHRNGIQSPTYQCLAASIINWQADVERCIEWEAHSRRATGLTLAF LTAAAKELGCPLTC</aaseq>|<br />
DNA = <dnaseqindica>8415..8720#8230..8322#7894..8112#7625..7693#6024..7520#5016..5225#4732..4882#4414..4654#3489..3547#2792..3066#2490..2632#2357..2411#2187..2264#2008..2094#ccgagccgtttctctccctctctctctcctcctccttcccctgcgcctcctccaccgcccgtgtcgccgccgcgcgcgtctatcccctcgccgccgccgccgtctacgccgcgtgaggtccggattggttgaaaggctgatcgaggtaatgcttttttatttttcccgtgcggttttagtttttctgtgctgtgcttggtggttttttctttaaaactatttttaggctggttttgatgatggcctttctaaggttgtcgtgtatggggggcgaggtggttcgggtggatttgtggtgttatgttgaggggatttttgtttggggggagagagttttggtggtagatgcggttgggattttgaagacgtgaaagggaaattagttttgaaatttgacggcgaggttgagctgattagtattgtaataattcgtggcatcggatagatcaacgcgagatggatgcttattactgcgaatttcaggaggtttttgtggtttttttaggcgagtagcgttgattctagctggattttgctgcttcgtggaggcgtttagatgtggtgcatacggctgtggatgtaatttgtgcggggataattactgaaaattaattcgcagtaagctagctctgctttaatattttttcccctgaaacagtaacaagggatcccttcctcatttcgttaacagtggaagtgaatacagacaccaaccgatgagcagagacacctgtaattaaacaccaaaagaagaacctgttttgttcctatatattttccatttctatgctttaggtaatatactttctggttcggtgtcatagtacccgaaatagaatctttgcttgccaaaaaaagaaagaataacaagatagaacaacatcattaaattctgtcttagacaatacatttataatcgaccaacaaaaacagaccgatgagtgacgtggaataacaaatgaagtataaatgtcacgaaaaacacatctattgattttgtgtgataagccataggagcctacgaggaaccaagtgaatggttctgtacaccacatccattagcattaccataaatttgcctatttctatgatgccttccttaacatatttatcgacatgtgagcttgcaccataactaaatgttatggtaaactttaccttataaatgattgcaacatgtaccttttcatttcgataattaaaaacgagaaagcttccacctttttcctagatatattttttcgtttgattatgaatttagttgtacttgacaataaaataatgcgcttgtaagattatgcaattttcgttgcattacctccctctcttttcatttgtcagtttttttctactccctccgtttcatattttaagtcgtttgacttttttcctagtcagactttgttaagtttgacaaagtttatagtaaaatttagcaatatctaaaatattaaattggtttcatcaaatctaacattgaatatattttgataatatgttttttttgtgttgaaaatacttctatatttttctataaacttagtcaaacttggaagaaatttcagtaagaaaaaagtcaaacggcttataatatgaaacggagggagtaaatttttttttgcattaattaaaattcatgtattttcctccttttttggtttttgttgtgcgatatattacagtgtagagactctgaagctcatatggttggcattggttgatatgataacgtgtcacgttgtgagatgcagacccccgttatgggggttgattgaatgaagtggatagcactcttataactttcttgctcgaagtttcttgataccaaggcaagtagcttataagaagatttggactttggtggcataaatggatgaactaactggcggcttcaacgtggtgttgattacaggtgcttttattgctaactggagcagtttgtgtagtttctttctttctctttgagaattctctcagtttcttgggcaggctcttgagctaccaagtatgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggtaacattattcctatttgattcttgttagaaattctatatgttgaacatacttgcctcaatgggaaacttggttgcgtctcctctctgaaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcaggtactgtatctctaagcatgtttagtatcttctgtgataaaatctttctccagtgtgttaacttttgcattctttttcttgcatgcttgcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattaggtaatttgatccagtgatcaatacatgttccacaaacaattgaagaaagagtttgacatatttttctcatatttccagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgaggtaaggcccctactctaatggttagttccaatttattggttcatttctctgtgtttgataagattttgcctctgctgtgcatttcaatattatatctaaactgtgctgcttatagcttatcaaactactcttgctcataaagatgcaacttttatgcagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgagtcataataaatgataacatattattgttctgtggactgaaaatctgaaatcctcctttcttgagaaatacaacctcaattttatggtgatattgacaagaaggaagagcgcatataatttaaagttaaaacacggaaaataatagaagtatgtgagcagaacgcaggagcaaggggtggaactcataacaagttcaaacttcatgcatgttcataattccccaatgcaagtatggcaagaaaaatagacaacttttaaaagtatgctaatggaactgcatgcattgtgtaaacttgcttttgtcaagcatgttggaacatactatatatttgctgctgatgatgaaaaatttgcttgagtactcttctattgctaatggacgcttaccattccctgttttgtcttcaaatgcatcaggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactggttggtaaaatcagcaatggaattttctatcgtgaattttttataattattgtaacattaattactcccatgtctttttattctgttgagataccatgtgacaatgtctacaattgcatattttgcattctttggtagatcagttcccttcttaagatccccaaagggaaactggagtatcaataacttactccctcaaataaaaaaaaagtgatactttaggcataaacacattcttcaaaacaaaactttgaatagcagtttcttttgtaatatattaataaagcatggaaaaattataatgtgaagaatgtacattcagggacaaatatactcatcatttttgcatggttgatcttaatgtccatagagagattgatcagaccaaaagtttatttttgacaggagtgagtatggattataataggagtgaatggcaatatgctgtgaaagtgagcattttatatttgtttactactagcatttggtgacaaagcaaatgatgtatattgtaaatcaaagtaaacaagtagaacatgtcctgtggcgacaagacataacctacaaattaaagttagcctgattgtttattttactgggagcacattttttctagattctagagtacatattattaaaaaaggtgtgatatgtttgcaaattactttttaatggaagtttgtaaattactttgaaacttgcattcagtttttttgcttcttaaaaaacatagtgcatatgggctttatgagctcatactatgcatgtttttctgcattggagtgggacctttatctttttcttgcaactcggatgctaatgcttattataaacacaagttactaacatgtgcttgggtcattcagcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcataggtaagctcttatgatcctgtccactgttgtaccatgtgtactttccaaatgattgctaacatgctatttttcgtcagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtac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/>
Link = [http://www.ncbi.nlm.nih.gov/nuccore/NM_001048755.1 RefSeq:Os01g0182600]|<br />
}}<br />
[[Category:Genes]]<br />
[[Category:Japonica mRNA]]<br />
[[Category:Oryza Sativa Japonica Group]]<br />
[[Category:Japonica Genes]]<br />
[[Category:Japonica Chromosome 1]]<br />
[[Category:Chromosome 1]]</div>Mikahttps://ngdc.cncb.ac.cn/ricewiki/index.php?title=Os01g0182600&diff=183857Os01g01826002014-06-11T04:26:06Z<p>Mika: /* Evolution */</p>
<hr />
<div>The rice gene of ''Gigantea(GI)'' is solo ortholog of ''GI'' in the Arabidopsis circadian system and related to photoperiodic control of flowering.<br />
<br />
==Annotated Information==<br />
===Function===<br />
*''OsGI'',a ''GI'' ortholog, regulates flowering time in response to photoperiodic conditions. Mutations in this gene delay flowering in inductive SDs (short-days),whereas flowering is marginally induced under LDs(long-days).In comparison, over expression of ''OsGI'' in transgenic rice causes late flowering in both SDs and LDs.In addition, ''OsGI'' has function to regulate ''Hd1'', a gene that also shows circadian rhythm<ref name="ref1" />. ''Os-GI'' affects gene expression of approximately 75% of genes among 27,201 genes and is required for strong amplitudes and proper phase-setting of global gene expression under natural field conditions<ref name="ref2" />. <br />
*The cDNA of ''OsGI'' firstly found in ''se5'' mutant that is completely photoperiod-insensitive, the expression patterns of rice and Arabidopsis ''GI'' gene are both photoperiodically regulated and are very similar under LD and SD conditions<ref name="ref3" />. [[File:Limeijia2.png|frame|'''Figure 1''' Possible roles of isolated genes in the photoperiodic regulation of flowering in rice, including ''OsGI'', may function to inhibit flowering under LD conditions.(from reference <ref name="ref3" />).]]''GI'' is required for phase-dependent ''CO''transcription through the interaction with FKF1, which leads to cycling DNA binding with one finger (CDF) degradation and consequent derepression of CO. In addition, it has been known that GI protein interacts with ZEITLUPE (ZTL) F-box protein to control the diurnal rhythm of TOC1 protein in regulation of the Arabidopsis circadian clock<ref name="ref4" />. <br />
*''Os-GI'' also has function to maintain bioactive GA level through the regulation of the GA-deactivating enzyme genes in rice, ''Os-GI'' represses major ''OsGA2ox'' genes in order to maintain a proper bioactive GA level, ''osgi-1'' plants showed a semi-dwarf phenotype with elevated ''OsGA2ox'' gene expression <ref name="ref4" /> .[[File:Limeijia1.png|frame|'''Figure 2''' WT rice and osgi-1 plants grown under natural field conditions and ''osgi-1'' plant exhibits Semi-dwarf phenotype(from reference <ref name="ref4" />).]]<br />
<br />
===Expression===<br />
*''GI'' encodes a nuclear protein that defines the proper amplitude and period length of circadian rhythms. ''GI'' expression is regulated by the circadian clock, with a peak in transcript levels at 8 to 10 h after dawn. The ''GI'' protein acts between the circadian oscillator and ''CO'' to promote flowering by increasing ''CO'' and ''FT'' mRNA abundance<ref name="ref1" />. The expression of the ''OsGI'' messenger RNA was circadian-controlled and that its temporal expression pattern was very similar to that of ''GI'' under both SD and LD conditions. Decreased ''OsGI'' expression led to early flowering under LD conditions and late flowering under SD conditions relative to the wild type, and that this early flowering phenotype under LD conditions was the opposite of that of the Arabidopsis ''gi'' mutants, confirming the opposite role of the ''GI'' gene in flowering under LD conditions in rice and Arabidopsis<ref name="ref5" />.<br />
<br />
===Mutation===<br />
*The single ''osgi-1'' mutation extended the time to flowering only under SD conditions. Under LD conditions the single ''hd1'' mutation induced early flowering, but ''osgi-1'' plants exhibited no phenotypic change<ref name="ref4" /><br />
<br />
===Evolution===<br />
* There is some evolutionary conservation between rice circadian clock–related genes and those associated with Arabidopsis circadian clocks, where the circadian clock consists of interlocked subloops, it is also possible that ''Os-GI'' is a member of a subloop in rice circadian clocks<ref name="ref4" />.But there is a divergence in the photoperiodic pathways of these two species arises from the molecular mechanisms controlling the expression of ''FT/Hd3a'' by ''CO/Hd1(Se1)'' ['''Figure3''']<ref name="ref5" />[[File:Limeijia3.png|frame|'''Figure 3'''The molecular mechanisms for the regulation of ''GI/OsGI'' expression, and the regulation of ''CO/Hd1(Se1)'' expression by ''GI/OsGI'', might be conserved between ''Arabidopsis'' and rice. Both the ''GI–CO'' and ''OsGI–Hd1(Se1)'' pathways are activated under LD conditions; the former promotes the flowering of ''Arabidopsis'', whereas the latter inhibits the flowering of rice. The divergence in the photoperiodic pathways of these two species arises from the molecular mechanisms controlling the expression of ''FT/Hd3a'' by ''CO/Hd1(Se1)''(from reference <ref name="ref5" />).]]<br />
<br />
==Labs working on this gene==<br />
*Photosynthesis and Photobiology Research Unit, National Institute of Agrobiological Sciences, 2-1-2 Kannondai, Tsukuba 305-8602, Japan<br />
*Laboratory of Plant Molecular Genetics, Nara Institute of Science and Technology, 8916-5 Takayama, Ikoma 630-0101, Japan<br />
*Plant Genomics Research Unit, National Institute of Agrobiological Sciences, 2-1-2 Kannondai, Tsukuba 305-8602, Japan<br />
<br />
==Knowledge Extension==<br />
*The rice orthologues of the two key regulators of flowering time in Arabidopsis, ''CO'' and ''FT'', were recently shown to be important in the photoperiodic control of flowering in rice. The rice ''Hd1(Se1)''gene, which is an orthologue of the Arabidopsis ''CO'' gene, was required for the suppression of flowering under LD conditions and for the promotion of flowering under SD conditions. Furthermore, the rice ''Hd3a'' gene, an orthologue of the Arabidopsis ''FT'' gene, was shown to be an activator of flowering in rice. It was previously demonstrated that ''CO'' controls the flowering time by integrating signals from the circadian clock and light to regulate ''FT'' expression in Arabidopsis<ref name="ref5" />.Unlike Arabidopsis ''FT'', expression of ''Hd3a'' mRNA is more or less opposite to that of ''Hd1'' under inductive short-day conditions<ref name="ref6" />.<br />
<br />
==References==<br />
<references><br />
<ref name="ref1">Song Lim Kim, Shinyoung Lee, Hyo Jung Kim, Hong Gil Nam, Gynheung An（2007）''OsMADS51'' is a short-day flowering promoter that functions upstream of ''Ehd1'', ''OsMADS14'', and ''Hd3a''.''Plant Physiology'' 145(4): 1484-1494.</ref><br />
<ref name="ref2">Hironori Itoh, Takeshi Izawa (2011) A study of phytohormone biosynthetic gene expression using a circadian clock-related mutant in rice. ''Plant Signaling & Behavior'' 6(12): 1932-1936.</ref> <br />
<ref name="ref3">Ryousuke Hayama, Takeshi Izawa, Ko Shimamoto (2002) Isolation of rice genes possibly involved in the photoperiodic control of flowering by a<br />
fluorescent differential display method. ''Plant and Cell Physiology'' 43(5): 494-504.</ref> <br />
<ref name="ref4">Takeshi Izawa, Motohiro Mihara, Yuji Suzuki, Meenu Gupta, Hironori Itoh, et al. (2011) ''Os-GIGANTEA'' confers robust diurnal rhythms on the global transcriptome of rice in the field. ''The Plant Cell'' 23(5): 1741-1755.</ref> <br />
<ref name="ref5">Ryosuke Hayama, Shuji Yokoi, Shojiro Tamaki, Masahiro Yano, Ko Shimamoto (2003) Adaptation of photoperiodic control pathways produces short-day flowering in rice. ''Nature'' 422(6933): 719-722.</ref> <br />
<ref name="ref6">Bong-Soo Shin, Jeong-Hyun Lee, Jeong-Hwan Lee, Hyun-Joo Jeong, Choong-Hyo Yun, Jeong-Kook Kim (2004) Circadian regulation of rice (''Oryza sativa L.'') ''CONSTANS-like'' gene transcripts. ''Molecules and Cells'' 17(1): 10-16.</ref><br />
<br />
==Structured Information==<br />
{{JaponicaGene|<br />
GeneName = Os01g0182600|<br />
Description = GIGANTEA protein|<br />
Version = NM_001048755.1 GI:115434923 GeneID:4325329|<br />
Length = 9083 bp|<br />
Definition = Oryza sativa Japonica Group Os01g0182600, complete gene.|<br />
Source = Oryza sativa Japonica Group<br />
<br />
ORGANISM Oryza sativa Japonica Group<br />
Eukaryota; Viridiplantae; Streptophyta; Embryophyta; Tracheophyta;<br />
Spermatophyta; Magnoliophyta; Liliopsida; Poales; Poaceae; BEP<br />
clade; Ehrhartoideae; Oryzeae; Oryza.<br />
|<br />
Chromosome = [[:category:Japonica Chromosome 1|Chromosome 1]]|<br />
AP = Chromosome 1:4328362..4337444|<br />
CDS = 4328725..4329030,4329123..4329215,4329333..4329551,4329752..4329820,4329925..4331421<br>,4332220..4332429,4332563..4332713,4332791..4333031,4333898..4333956<br>,4334379..4334653,4334813..4334955,4335034..4335088,4335181..4335258<br>,4335351..4335437|<br />
GCID = <gbrowseImage1><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage1>|<br />
GSID = <gbrowseImage2><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage2>|<br />
CDNA = <cdnaseq>atgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactgcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcatagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcggcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctga</cdnaseq>|<br />
AA = <aaseq>MSASNEKWIDGLQFSSLFWPPPQDSQQKQAQILAYVEYFGQFTA DSEQFPEDIAQLIQSCYPSKEKRLVDEVLATFVLHHPEHGHAVVHPILSRIIDGTLSY DRNGFPFMSFISLFSHTSEKEYSEQWALACGEILRVLTHYNRPIFKVDHQHSEAECSS TSDQASSCESMEKRANGSPRNEPDRKPLRPLSPWITDILLAAPLGIRSDYFRWCGGVM GKYAAGGELKPPTTAYSRGSGKHPQLMPSTPRWAVANGAGVILSVCDEEVARYETANL TAAAVPALLLPPPTTPLDEHLVAGLPPLEPYARLFHRYYAIATPSATQRLLFGLLEAP PSWAPDALDAAVQLVELLRAAEDYDSGMRLPKNWMHLHFLRAIGTAMSMRAGIAADTS AALLFRILSQPTLLFPPLRHAEGVELHHEPLGGYVSSYKRQLEVPASEATIDATAQGI ASMLCAHGPDVEWRICTIWEAAYGLLPLSSSAVDLPEIVVAAPLQPPTLSWSLYLPLL KVFEYLPRGSPSEACLMRIFVATVEAILRRTFPSETSEQSRKPRSQSKNLAVAELRTM IHSLFVESCASMDLASRLLFVVLTVCVSHQALPGGSKRPTGSDNHSSEEVTNDSRLTN GRNRCKKRQGPVATFDSYVLAAVCALSCELQLFPFISKNGNHSNLKDSIKIVIPGKTT GISNELHNSISSAILHTRRILGILEALFSLKPSSVGTSWSYSSNEIVAAAMVAAHVSE LFRRSRPCLNALSALKQCKWDAEISTRASSLYHLIDLHGKTVTSIVNKAEPLEAHLTL TPVKKDEPPIEEKNINSSDGGALEKKDASRSHRKNGFARPLLKCAEDVILNGDVASTS GKAIASLQVEASDLANFLTMDRNGGYRGSQTLLRSVLSEKQELCFSVVSLLWQKLIAS PEMQMSAESTSAHQGWRKVVDALCDIVSASPTKASAAIVLQAEKDLQPWIARDDEQGQ KMWRVNQRIVKLIAELMRNHDSPEALVILASASDLLLRATDGMLVDGEACTLPQLELL EVTARAVHLIVEWGDSGVSVADGLSNLLKCRLSTTIRCLSHPSAHVRALSMSVLRDIL NSGQINSSKLIQGEHRNGIQSPTYQCLAASIINWQADVERCIEWEAHSRRATGLTLAF LTAAAKELGCPLTC</aaseq>|<br />
DNA = <dnaseqindica>8415..8720#8230..8322#7894..8112#7625..7693#6024..7520#5016..5225#4732..4882#4414..4654#3489..3547#2792..3066#2490..2632#2357..2411#2187..2264#2008..2094#ccgagccgtttctctccctctctctctcctcctccttcccctgcgcctcctccaccgcccgtgtcgccgccgcgcgcgtctatcccctcgccgccgccgccgtctacgccgcgtgaggtccggattggttgaaaggctgatcgaggtaatgcttttttatttttcccgtgcggttttagtttttctgtgctgtgcttggtggttttttctttaaaactatttttaggctggttttgatgatggcctttctaaggttgtcgtgtatggggggcgaggtggttcgggtggatttgtggtgttatgttgaggggatttttgtttggggggagagagttttggtggtagatgcggttgggattttgaagacgtgaaagggaaattagttttgaaatttgacggcgaggttgagctgattagtattgtaataattcgtggcatcggatagatcaacgcgagatggatgcttattactgcgaatttcaggaggtttttgtggtttttttaggcgagtagcgttgattctagctggattttgctgcttcgtggaggcgtttagatgtggtgcatacggctgtggatgtaatttgtgcggggataattactgaaaattaattcgcagtaagctagctctgctttaatattttttcccctgaaacagtaacaagggatcccttcctcatttcgttaacagtggaagtgaatacagacaccaaccgatgagcagagacacctgtaattaaacaccaaaagaagaacctgttttgttcctatatattttccatttctatgctttaggtaatatactttctggttcggtgtcatagtacccgaaatagaatctttgcttgccaaaaaaagaaagaataacaagatagaacaacatcattaaattctgtcttagacaatacatttataatcgaccaacaaaaacagaccgatgagtgacgtggaataacaaatgaagtataaatgtcacgaaaaacacatctattgattttgtgtgataagccataggagcctacgaggaaccaagtgaatggttctgtacaccacatccattagcattaccataaatttgcctatttctatgatgccttccttaacatatttatcgacatgtgagcttgcaccataactaaatgttatggtaaactttaccttataaatgattgcaacatgtaccttttcatttcgataattaaaaacgagaaagcttccacctttttcctagatatattttttcgtttgattatgaatttagttgtacttgacaataaaataatgcgcttgtaagattatgcaattttcgttgcattacctccctctcttttcatttgtcagtttttttctactccctccgtttcatattttaagtcgtttgacttttttcctagtcagactttgttaagtttgacaaagtttatagtaaaatttagcaatatctaaaatattaaattggtttcatcaaatctaacattgaatatattttgataatatgttttttttgtgttgaaaatacttctatatttttctataaacttagtcaaacttggaagaaatttcagtaagaaaaaagtcaaacggcttataatatgaaacggagggagtaaatttttttttgcattaattaaaattcatgtattttcctccttttttggtttttgttgtgcgatatattacagtgtagagactctgaagctcatatggttggcattggttgatatgataacgtgtcacgttgtgagatgcagacccccgttatgggggttgattgaatgaagtggatagcactcttataactttcttgctcgaagtttcttgataccaaggcaagtagcttataagaagatttggactttggtggcataaatggatgaactaactggcggcttcaacgtggtgttgattacaggtgcttttattgctaactggagcagtttgtgtagtttctttctttctctttgagaattctctcagtttcttgggcaggctcttgagctaccaagtatgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggtaacattattcctatttgattcttgttagaaattctatatgttgaacatacttgcctcaatgggaaacttggttgcgtctcctctctgaaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcaggtactgtatctctaagcatgtttagtatcttctgtgataaaatctttctccagtgtgttaacttttgcattctttttcttgcatgcttgcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattaggtaatttgatccagtgatcaatacatgttccacaaacaattgaagaaagagtttgacatatttttctcatatttccagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgaggtaaggcccctactctaatggttagttccaatttattggttcatttctctgtgtttgataagattttgcctctgctgtgcatttcaatattatatctaaactgtgctgcttatagcttatcaaactactcttgctcataaagatgcaacttttatgcagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgagtcataataaatgataacatattattgttctgtggactgaaaatctgaaatcctcctttcttgagaaatacaacctcaattttatggtgatattgacaagaaggaagagcgcatataatttaaagttaaaacacggaaaataatagaagtatgtgagcagaacgcaggagcaaggggtggaactcataacaagttcaaacttcatgcatgttcataattccccaatgcaagtatggcaagaaaaatagacaacttttaaaagtatgctaatggaactgcatgcattgtgtaaacttgcttttgtcaagcatgttggaacatactatatatttgctgctgatgatgaaaaatttgcttgagtactcttctattgctaatggacgcttaccattccctgttttgtcttcaaatgcatcaggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactggttggtaaaatcagcaatggaattttctatcgtgaattttttataattattgtaacattaattactcccatgtctttttattctgttgagataccatgtgacaatgtctacaattgcatattttgcattctttggtagatcagttcccttcttaagatccccaaagggaaactggagtatcaataacttactccctcaaataaaaaaaaagtgatactttaggcataaacacattcttcaaaacaaaactttgaatagcagtttcttttgtaatatattaataaagcatggaaaaattataatgtgaagaatgtacattcagggacaaatatactcatcatttttgcatggttgatcttaatgtccatagagagattgatcagaccaaaagtttatttttgacaggagtgagtatggattataataggagtgaatggcaatatgctgtgaaagtgagcattttatatttgtttactactagcatttggtgacaaagcaaatgatgtatattgtaaatcaaagtaaacaagtagaacatgtcctgtggcgacaagacataacctacaaattaaagttagcctgattgtttattttactgggagcacattttttctagattctagagtacatattattaaaaaaggtgtgatatgtttgcaaattactttttaatggaagtttgtaaattactttgaaacttgcattcagtttttttgcttcttaaaaaacatagtgcatatgggctttatgagctcatactatgcatgtttttctgcattggagtgggacctttatctttttcttgcaactcggatgctaatgcttattataaacacaagttactaacatgtgcttgggtcattcagcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcataggtaagctcttatgatcctgtccactgttgtaccatgtgtactttccaaatgattgctaacatgctatttttcgtcagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcgggtatatcatacatctctctggctgttgtttcagtgctatcaatcaagatgacacatcatgcaatattatcataacagcacattatgtgttttgaatattttattgattgtcatagctttgttttatgttttagcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcaggtatgcagtgtttcctgcacctacttgctacctattctgtatttgatgtattcaatttcctttttatcagttttctcgaaatatatgctaataggcttgtcactaaacggctacattatccagactctgttctgaatttttttttcattaagtactgttagctgccagattacatcatcatgactaaagtagtttagctacaatattgcaaagctagatttgtctagcctctattgttttgaatagaaaaggcaggtatgcagtgtttcctgcacctacttgctacctattctgtatttgatgtattcaatttcctttttatcagttttctcgaaatatatgctaataggcttgtcactaaacggctacattatccagactctgttctgaatttttttttcattaagtactgttagctgccagattacatcatcatgactaaagtagtttagctacaatattgcaaagctagatttgtctagcctctattgttttgaatagaaaattaaatcttttggtcatttttttgtaaccttttaggagtctctgtatgcttgtcgtcaacatgctatgataagtttatatcagttcttttggagctatgatgtattcttgtatattttatgactgctaggatattatgttaaaactgttttaaaggtatcatcaaacaagttaatatagatattgctacctaccaaagttggaatttgattatgtcgtaaaatatatgcaccctacatgttttagcttcatattgacttatattttcatatgtgctccttcaactgcacaaagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggtatggtcattcctgcttataagataccagaataatgctatgaagaccgtccttggttgtcagcattcttatacacaatgataatctttgtttttcatgggtaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggtcaagatattgttattgttatatttacatacttgttatcatatttaaaagcttcatattgtatgtacctttgaaaaaaaaagttccatactgtagcttcttttgttctcatttcttggtttcaaagcatcagcatcattaccgatataaaatatacccatttcctcagtcatcttccccgactcttgtatctttcacaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagaggtaacaccttcattcatgaggtccttaggtgcaaatcccagtgaagaataatcagttctactttgtatggttcatctcgcttactgaccggttcatctcgggatcaattacgagcagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaaggcatgtcctcttctctgatctcactcacgttcatgccaaacttactcatcatcatccatgatccatcatcctgactgaatcttgatatgcagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctgacaaggccacaccatgactgtcactgcaagcagctgctgtaggatcagcgagtaggaaaggatgacttgccggccagtttcctgagatgtgatttaagactgatattagctgatgttcccaagcatgatacggggcttgctcccaagatgtgatttttactttacttttggttaatgtgctaccgctgacattagatgttcaagcatatgaaaactgcttgtgctgtaacttgtatgtctgtaacgtaaaaatctttccatatggtaaccggggaagcagaggtatatccttttctgctgttgtatctatttccagtttcctgctgaacttgcaaatgaatattcagcaatttccgttttgctc</dnaseqindica>|<br />
Link = [http://www.ncbi.nlm.nih.gov/nuccore/NM_001048755.1 RefSeq:Os01g0182600]|<br />
}}<br />
[[Category:Genes]]<br />
[[Category:Japonica mRNA]]<br />
[[Category:Oryza Sativa Japonica Group]]<br />
[[Category:Japonica Genes]]<br />
[[Category:Japonica Chromosome 1]]<br />
[[Category:Chromosome 1]]</div>Mikahttps://ngdc.cncb.ac.cn/ricewiki/index.php?title=File:Limeijia3.png&diff=183856File:Limeijia3.png2014-06-11T04:23:57Z<p>Mika: The molecular mechanisms for the regulation of GI/OsGIexpression, and the regulation of CO/Hd1(Se1) expression by GI/OsGI, might be conserved between
Arabidopsisand rice. Both theGI–COandOsGI–Hd1(Se1) pathways are activated under
LD conditions; the fo</p>
<hr />
<div>The molecular mechanisms for the regulation of GI/OsGIexpression, and the regulation of CO/Hd1(Se1) expression by GI/OsGI, might be conserved between<br />
Arabidopsisand rice. Both theGI–COandOsGI–Hd1(Se1) pathways are activated under<br />
LD conditions; the former promotes the flowering ofArabidopsis, whereas the latter<br />
inhibits the flowering of rice. The divergence in the photoperiodic pathways of these two species arises from the molecular mechanisms controlling the expression ofFT/Hd3a by CO/Hd1(Se1)</div>Mikahttps://ngdc.cncb.ac.cn/ricewiki/index.php?title=Os01g0182600&diff=183855Os01g01826002014-06-11T04:22:57Z<p>Mika: /* Evolution */</p>
<hr />
<div>The rice gene of ''Gigantea(GI)'' is solo ortholog of ''GI'' in the Arabidopsis circadian system and related to photoperiodic control of flowering.<br />
<br />
==Annotated Information==<br />
===Function===<br />
*''OsGI'',a ''GI'' ortholog, regulates flowering time in response to photoperiodic conditions. Mutations in this gene delay flowering in inductive SDs (short-days),whereas flowering is marginally induced under LDs(long-days).In comparison, over expression of ''OsGI'' in transgenic rice causes late flowering in both SDs and LDs.In addition, ''OsGI'' has function to regulate ''Hd1'', a gene that also shows circadian rhythm<ref name="ref1" />. ''Os-GI'' affects gene expression of approximately 75% of genes among 27,201 genes and is required for strong amplitudes and proper phase-setting of global gene expression under natural field conditions<ref name="ref2" />. <br />
*The cDNA of ''OsGI'' firstly found in ''se5'' mutant that is completely photoperiod-insensitive, the expression patterns of rice and Arabidopsis ''GI'' gene are both photoperiodically regulated and are very similar under LD and SD conditions<ref name="ref3" />. [[File:Limeijia2.png|frame|'''Figure 1''' Possible roles of isolated genes in the photoperiodic regulation of flowering in rice, including ''OsGI'', may function to inhibit flowering under LD conditions.(from reference <ref name="ref3" />).]]''GI'' is required for phase-dependent ''CO''transcription through the interaction with FKF1, which leads to cycling DNA binding with one finger (CDF) degradation and consequent derepression of CO. In addition, it has been known that GI protein interacts with ZEITLUPE (ZTL) F-box protein to control the diurnal rhythm of TOC1 protein in regulation of the Arabidopsis circadian clock<ref name="ref4" />. <br />
*''Os-GI'' also has function to maintain bioactive GA level through the regulation of the GA-deactivating enzyme genes in rice, ''Os-GI'' represses major ''OsGA2ox'' genes in order to maintain a proper bioactive GA level, ''osgi-1'' plants showed a semi-dwarf phenotype with elevated ''OsGA2ox'' gene expression <ref name="ref4" /> .[[File:Limeijia1.png|frame|'''Figure 2''' WT rice and osgi-1 plants grown under natural field conditions and ''osgi-1'' plant exhibits Semi-dwarf phenotype(from reference <ref name="ref4" />).]]<br />
<br />
===Expression===<br />
*''GI'' encodes a nuclear protein that defines the proper amplitude and period length of circadian rhythms. ''GI'' expression is regulated by the circadian clock, with a peak in transcript levels at 8 to 10 h after dawn. The ''GI'' protein acts between the circadian oscillator and ''CO'' to promote flowering by increasing ''CO'' and ''FT'' mRNA abundance<ref name="ref1" />. The expression of the ''OsGI'' messenger RNA was circadian-controlled and that its temporal expression pattern was very similar to that of ''GI'' under both SD and LD conditions. Decreased ''OsGI'' expression led to early flowering under LD conditions and late flowering under SD conditions relative to the wild type, and that this early flowering phenotype under LD conditions was the opposite of that of the Arabidopsis ''gi'' mutants, confirming the opposite role of the ''GI'' gene in flowering under LD conditions in rice and Arabidopsis<ref name="ref5" />.<br />
<br />
===Mutation===<br />
*The single ''osgi-1'' mutation extended the time to flowering only under SD conditions. Under LD conditions the single ''hd1'' mutation induced early flowering, but ''osgi-1'' plants exhibited no phenotypic change<ref name="ref4" /><br />
<br />
===Evolution===<br />
* There is some evolutionary conservation between rice circadian clock–related genes and those associated with Arabidopsis circadian clocks, where the circadian clock consists of interlocked subloops, it is also possible that ''Os-GI'' is a member of a subloop in rice circadian clocks<ref name="ref4" />.But there is a divergence in the photoperiodic pathways of these two species arises from the molecular mechanisms controlling the expression of ''FT/Hd3a'' by ''CO/Hd1(Se1)''<ref name="ref5" />[[File:Limeijia3.png|frame|'''Figure 3'''The molecular mechanisms for the regulation of ''GI/OsGI'' expression, and the regulation of ''CO/Hd1(Se1)'' expression by ''GI/OsGI'', might be conserved between ''Arabidopsis'' and rice. Both the ''GI–CO'' and ''OsGI–Hd1(Se1)'' pathways are activated under LD conditions; the former promotes the flowering of ''Arabidopsis'', whereas the latter inhibits the flowering of rice. The divergence in the photoperiodic pathways of these two species arises from the molecular mechanisms controlling the expression of ''FT/Hd3a'' by ''CO/Hd1(Se1)''(from reference <ref name="ref5" />).]]<br />
<br />
==Labs working on this gene==<br />
*Photosynthesis and Photobiology Research Unit, National Institute of Agrobiological Sciences, 2-1-2 Kannondai, Tsukuba 305-8602, Japan<br />
*Laboratory of Plant Molecular Genetics, Nara Institute of Science and Technology, 8916-5 Takayama, Ikoma 630-0101, Japan<br />
*Plant Genomics Research Unit, National Institute of Agrobiological Sciences, 2-1-2 Kannondai, Tsukuba 305-8602, Japan<br />
<br />
==Knowledge Extension==<br />
*The rice orthologues of the two key regulators of flowering time in Arabidopsis, ''CO'' and ''FT'', were recently shown to be important in the photoperiodic control of flowering in rice. The rice ''Hd1(Se1)''gene, which is an orthologue of the Arabidopsis ''CO'' gene, was required for the suppression of flowering under LD conditions and for the promotion of flowering under SD conditions. Furthermore, the rice ''Hd3a'' gene, an orthologue of the Arabidopsis ''FT'' gene, was shown to be an activator of flowering in rice. It was previously demonstrated that ''CO'' controls the flowering time by integrating signals from the circadian clock and light to regulate ''FT'' expression in Arabidopsis<ref name="ref5" />.Unlike Arabidopsis ''FT'', expression of ''Hd3a'' mRNA is more or less opposite to that of ''Hd1'' under inductive short-day conditions<ref name="ref6" />.<br />
<br />
==References==<br />
<references><br />
<ref name="ref1">Song Lim Kim, Shinyoung Lee, Hyo Jung Kim, Hong Gil Nam, Gynheung An（2007）''OsMADS51'' is a short-day flowering promoter that functions upstream of ''Ehd1'', ''OsMADS14'', and ''Hd3a''.''Plant Physiology'' 145(4): 1484-1494.</ref><br />
<ref name="ref2">Hironori Itoh, Takeshi Izawa (2011) A study of phytohormone biosynthetic gene expression using a circadian clock-related mutant in rice. ''Plant Signaling & Behavior'' 6(12): 1932-1936.</ref> <br />
<ref name="ref3">Ryousuke Hayama, Takeshi Izawa, Ko Shimamoto (2002) Isolation of rice genes possibly involved in the photoperiodic control of flowering by a<br />
fluorescent differential display method. ''Plant and Cell Physiology'' 43(5): 494-504.</ref> <br />
<ref name="ref4">Takeshi Izawa, Motohiro Mihara, Yuji Suzuki, Meenu Gupta, Hironori Itoh, et al. (2011) ''Os-GIGANTEA'' confers robust diurnal rhythms on the global transcriptome of rice in the field. ''The Plant Cell'' 23(5): 1741-1755.</ref> <br />
<ref name="ref5">Ryosuke Hayama, Shuji Yokoi, Shojiro Tamaki, Masahiro Yano, Ko Shimamoto (2003) Adaptation of photoperiodic control pathways produces short-day flowering in rice. ''Nature'' 422(6933): 719-722.</ref> <br />
<ref name="ref6">Bong-Soo Shin, Jeong-Hyun Lee, Jeong-Hwan Lee, Hyun-Joo Jeong, Choong-Hyo Yun, Jeong-Kook Kim (2004) Circadian regulation of rice (''Oryza sativa L.'') ''CONSTANS-like'' gene transcripts. ''Molecules and Cells'' 17(1): 10-16.</ref><br />
<br />
==Structured Information==<br />
{{JaponicaGene|<br />
GeneName = Os01g0182600|<br />
Description = GIGANTEA protein|<br />
Version = NM_001048755.1 GI:115434923 GeneID:4325329|<br />
Length = 9083 bp|<br />
Definition = Oryza sativa Japonica Group Os01g0182600, complete gene.|<br />
Source = Oryza sativa Japonica Group<br />
<br />
ORGANISM Oryza sativa Japonica Group<br />
Eukaryota; Viridiplantae; Streptophyta; Embryophyta; Tracheophyta;<br />
Spermatophyta; Magnoliophyta; Liliopsida; Poales; Poaceae; BEP<br />
clade; Ehrhartoideae; Oryzeae; Oryza.<br />
|<br />
Chromosome = [[:category:Japonica Chromosome 1|Chromosome 1]]|<br />
AP = Chromosome 1:4328362..4337444|<br />
CDS = 4328725..4329030,4329123..4329215,4329333..4329551,4329752..4329820,4329925..4331421<br>,4332220..4332429,4332563..4332713,4332791..4333031,4333898..4333956<br>,4334379..4334653,4334813..4334955,4335034..4335088,4335181..4335258<br>,4335351..4335437|<br />
GCID = <gbrowseImage1><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage1>|<br />
GSID = <gbrowseImage2><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage2>|<br />
CDNA = <cdnaseq>atgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactgcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcatagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcggcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctga</cdnaseq>|<br />
AA = <aaseq>MSASNEKWIDGLQFSSLFWPPPQDSQQKQAQILAYVEYFGQFTA DSEQFPEDIAQLIQSCYPSKEKRLVDEVLATFVLHHPEHGHAVVHPILSRIIDGTLSY DRNGFPFMSFISLFSHTSEKEYSEQWALACGEILRVLTHYNRPIFKVDHQHSEAECSS TSDQASSCESMEKRANGSPRNEPDRKPLRPLSPWITDILLAAPLGIRSDYFRWCGGVM GKYAAGGELKPPTTAYSRGSGKHPQLMPSTPRWAVANGAGVILSVCDEEVARYETANL TAAAVPALLLPPPTTPLDEHLVAGLPPLEPYARLFHRYYAIATPSATQRLLFGLLEAP PSWAPDALDAAVQLVELLRAAEDYDSGMRLPKNWMHLHFLRAIGTAMSMRAGIAADTS AALLFRILSQPTLLFPPLRHAEGVELHHEPLGGYVSSYKRQLEVPASEATIDATAQGI ASMLCAHGPDVEWRICTIWEAAYGLLPLSSSAVDLPEIVVAAPLQPPTLSWSLYLPLL KVFEYLPRGSPSEACLMRIFVATVEAILRRTFPSETSEQSRKPRSQSKNLAVAELRTM IHSLFVESCASMDLASRLLFVVLTVCVSHQALPGGSKRPTGSDNHSSEEVTNDSRLTN GRNRCKKRQGPVATFDSYVLAAVCALSCELQLFPFISKNGNHSNLKDSIKIVIPGKTT GISNELHNSISSAILHTRRILGILEALFSLKPSSVGTSWSYSSNEIVAAAMVAAHVSE LFRRSRPCLNALSALKQCKWDAEISTRASSLYHLIDLHGKTVTSIVNKAEPLEAHLTL TPVKKDEPPIEEKNINSSDGGALEKKDASRSHRKNGFARPLLKCAEDVILNGDVASTS GKAIASLQVEASDLANFLTMDRNGGYRGSQTLLRSVLSEKQELCFSVVSLLWQKLIAS PEMQMSAESTSAHQGWRKVVDALCDIVSASPTKASAAIVLQAEKDLQPWIARDDEQGQ KMWRVNQRIVKLIAELMRNHDSPEALVILASASDLLLRATDGMLVDGEACTLPQLELL EVTARAVHLIVEWGDSGVSVADGLSNLLKCRLSTTIRCLSHPSAHVRALSMSVLRDIL NSGQINSSKLIQGEHRNGIQSPTYQCLAASIINWQADVERCIEWEAHSRRATGLTLAF LTAAAKELGCPLTC</aaseq>|<br />
DNA = <dnaseqindica>8415..8720#8230..8322#7894..8112#7625..7693#6024..7520#5016..5225#4732..4882#4414..4654#3489..3547#2792..3066#2490..2632#2357..2411#2187..2264#2008..2094#ccgagccgtttctctccctctctctctcctcctccttcccctgcgcctcctccaccgcccgtgtcgccgccgcgcgcgtctatcccctcgccgccgccgccgtctacgccgcgtgaggtccggattggttgaaaggctgatcgaggtaatgcttttttatttttcccgtgcggttttagtttttctgtgctgtgcttggtggttttttctttaaaactatttttaggctggttttgatgatggcctttctaaggttgtcgtgtatggggggcgaggtggttcgggtggatttgtggtgttatgttgaggggatttttgtttggggggagagagttttggtggtagatgcggttgggattttgaagacgtgaaagggaaattagttttgaaatttgacggcgaggttgagctgattagtattgtaataattcgtggcatcggatagatcaacgcgagatggatgcttattactgcgaatttcaggaggtttttgtggtttttttaggcgagtagcgttgattctagctggattttgctgcttcgtggaggcgtttagatgtggtgcatacggctgtggatgtaatttgtgcggggataattactgaaaattaattcgcagtaagctagctctgctttaatattttttcccctgaaacagtaacaagggatcccttcctcatttcgttaacagtggaagtgaatacagacaccaaccgatgagcagagacacctgtaattaaacaccaaaagaagaacctgttttgttcctatatattttccatttctatgctttaggtaatatactttctggttcggtgtcatagtacccgaaatagaatctttgcttgccaaaaaaagaaagaataacaagatagaacaacatcattaaattctgtcttagacaatacatttataatcgaccaacaaaaacagaccgatgagtgacgtggaataacaaatgaagtataaatgtcacgaaaaacacatctattgattttgtgtgataagccataggagcctacgaggaaccaagtgaatggttctgtacaccacatccattagcattaccataaatttgcctatttctatgatgccttccttaacatatttatcgacatgtgagcttgcaccataactaaatgttatggtaaactttaccttataaatgattgcaacatgtaccttttcatttcgataattaaaaacgagaaagcttccacctttttcctagatatattttttcgtttgattatgaatttagttgtacttgacaataaaataatgcgcttgtaagattatgcaattttcgttgcattacctccctctcttttcatttgtcagtttttttctactccctccgtttcatattttaagtcgtttgacttttttcctagtcagactttgttaagtttgacaaagtttatagtaaaatttagcaatatctaaaatattaaattggtttcatcaaatctaacattgaatatattttgataatatgttttttttgtgttgaaaatacttctatatttttctataaacttagtcaaacttggaagaaatttcagtaagaaaaaagtcaaacggcttataatatgaaacggagggagtaaatttttttttgcattaattaaaattcatgtattttcctccttttttggtttttgttgtgcgatatattacagtgtagagactctgaagctcatatggttggcattggttgatatgataacgtgtcacgttgtgagatgcagacccccgttatgggggttgattgaatgaagtggatagcactcttataactttcttgctcgaagtttcttgataccaaggcaagtagcttataagaagatttggactttggtggcataaatggatgaactaactggcggcttcaacgtggtgttgattacaggtgcttttattgctaactggagcagtttgtgtagtttctttctttctctttgagaattctctcagtttcttgggcaggctcttgagctaccaagtatgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggtaacattattcctatttgattcttgttagaaattctatatgttgaacatacttgcctcaatgggaaacttggttgcgtctcctctctgaaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcaggtactgtatctctaagcatgtttagtatcttctgtgataaaatctttctccagtgtgttaacttttgcattctttttcttgcatgcttgcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattaggtaatttgatccagtgatcaatacatgttccacaaacaattgaagaaagagtttgacatatttttctcatatttccagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgaggtaaggcccctactctaatggttagttccaatttattggttcatttctctgtgtttgataagattttgcctctgctgtgcatttcaatattatatctaaactgtgctgcttatagcttatcaaactactcttgctcataaagatgcaacttttatgcagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgagtcataataaatgataacatattattgttctgtggactgaaaatctgaaatcctcctttcttgagaaatacaacctcaattttatggtgatattgacaagaaggaagagcgcatataatttaaagttaaaacacggaaaataatagaagtatgtgagcagaacgcaggagcaaggggtggaactcataacaagttcaaacttcatgcatgttcataattccccaatgcaagtatggcaagaaaaatagacaacttttaaaagtatgctaatggaactgcatgcattgtgtaaacttgcttttgtcaagcatgttggaacatactatatatttgctgctgatgatgaaaaatttgcttgagtactcttctattgctaatggacgcttaccattccctgttttgtcttcaaatgcatcaggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactggttggtaaaatcagcaatggaattttctatcgtgaattttttataattattgtaacattaattactcccatgtctttttattctgttgagataccatgtgacaatgtctacaattgcatattttgcattctttggtagatcagttcccttcttaagatccccaaagggaaactggagtatcaataacttactccctcaaataaaaaaaaagtgatactttaggcataaacacattcttcaaaacaaaactttgaatagcagtttcttttgtaatatattaataaagcatggaaaaattataatgtgaagaatgtacattcagggacaaatatactcatcatttttgcatggttgatcttaatgtccatagagagattgatcagaccaaaagtttatttttgacaggagtgagtatggattataataggagtgaatggcaatatgctgtgaaagtgagcattttatatttgtttactactagcatttggtgacaaagcaaatgatgtatattgtaaatcaaagtaaacaagtagaacatgtcctgtggcgacaagacataacctacaaattaaagttagcctgattgtttattttactgggagcacattttttctagattctagagtacatattattaaaaaaggtgtgatatgtttgcaaattactttttaatggaagtttgtaaattactttgaaacttgcattcagtttttttgcttcttaaaaaacatagtgcatatgggctttatgagctcatactatgcatgtttttctgcattggagtgggacctttatctttttcttgcaactcggatgctaatgcttattataaacacaagttactaacatgtgcttgggtcattcagcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcataggtaagctcttatgatcctgtccactgttgtaccatgtgtactttccaaatgattgctaacatgctatttttcgtcagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtac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/>
Link = [http://www.ncbi.nlm.nih.gov/nuccore/NM_001048755.1 RefSeq:Os01g0182600]|<br />
}}<br />
[[Category:Genes]]<br />
[[Category:Japonica mRNA]]<br />
[[Category:Oryza Sativa Japonica Group]]<br />
[[Category:Japonica Genes]]<br />
[[Category:Japonica Chromosome 1]]<br />
[[Category:Chromosome 1]]</div>Mikahttps://ngdc.cncb.ac.cn/ricewiki/index.php?title=File:Limeijia2.png&diff=183854File:Limeijia2.png2014-06-11T04:04:37Z<p>Mika: Possible roles of isolated genes in the photoperiodic regulation of flowering in rice, including OsGI, may function to inhibit flowering under LD conditions.</p>
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<div>Possible roles of isolated genes in the photoperiodic regulation of flowering in rice, including OsGI, may function to inhibit flowering under LD conditions.</div>Mikahttps://ngdc.cncb.ac.cn/ricewiki/index.php?title=Os01g0182600&diff=183853Os01g01826002014-06-11T04:03:51Z<p>Mika: /* Function */</p>
<hr />
<div>The rice gene of ''Gigantea(GI)'' is solo ortholog of ''GI'' in the Arabidopsis circadian system and related to photoperiodic control of flowering.<br />
<br />
==Annotated Information==<br />
===Function===<br />
*''OsGI'',a ''GI'' ortholog, regulates flowering time in response to photoperiodic conditions. Mutations in this gene delay flowering in inductive SDs (short-days),whereas flowering is marginally induced under LDs(long-days).In comparison, over expression of ''OsGI'' in transgenic rice causes late flowering in both SDs and LDs.In addition, ''OsGI'' has function to regulate ''Hd1'', a gene that also shows circadian rhythm<ref name="ref1" />. ''Os-GI'' affects gene expression of approximately 75% of genes among 27,201 genes and is required for strong amplitudes and proper phase-setting of global gene expression under natural field conditions<ref name="ref2" />. <br />
*The cDNA of ''OsGI'' firstly found in ''se5'' mutant that is completely photoperiod-insensitive, the expression patterns of rice and Arabidopsis ''GI'' gene are both photoperiodically regulated and are very similar under LD and SD conditions<ref name="ref3" />. [[File:Limeijia2.png|frame|'''Figure 1''' Possible roles of isolated genes in the photoperiodic regulation of flowering in rice, including ''OsGI'', may function to inhibit flowering under LD conditions.(from reference <ref name="ref3" />).]]''GI'' is required for phase-dependent ''CO''transcription through the interaction with FKF1, which leads to cycling DNA binding with one finger (CDF) degradation and consequent derepression of CO. In addition, it has been known that GI protein interacts with ZEITLUPE (ZTL) F-box protein to control the diurnal rhythm of TOC1 protein in regulation of the Arabidopsis circadian clock<ref name="ref4" />. <br />
*''Os-GI'' also has function to maintain bioactive GA level through the regulation of the GA-deactivating enzyme genes in rice, ''Os-GI'' represses major ''OsGA2ox'' genes in order to maintain a proper bioactive GA level, ''osgi-1'' plants showed a semi-dwarf phenotype with elevated ''OsGA2ox'' gene expression <ref name="ref4" /> .[[File:Limeijia1.png|frame|'''Figure 2''' WT rice and osgi-1 plants grown under natural field conditions and ''osgi-1'' plant exhibits Semi-dwarf phenotype(from reference <ref name="ref4" />).]]<br />
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===Expression===<br />
*''GI'' encodes a nuclear protein that defines the proper amplitude and period length of circadian rhythms. ''GI'' expression is regulated by the circadian clock, with a peak in transcript levels at 8 to 10 h after dawn. The ''GI'' protein acts between the circadian oscillator and ''CO'' to promote flowering by increasing ''CO'' and ''FT'' mRNA abundance<ref name="ref1" />. The expression of the ''OsGI'' messenger RNA was circadian-controlled and that its temporal expression pattern was very similar to that of ''GI'' under both SD and LD conditions. Decreased ''OsGI'' expression led to early flowering under LD conditions and late flowering under SD conditions relative to the wild type, and that this early flowering phenotype under LD conditions was the opposite of that of the Arabidopsis ''gi'' mutants, confirming the opposite role of the ''GI'' gene in flowering under LD conditions in rice and Arabidopsis<ref name="ref5" />.<br />
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===Mutation===<br />
*The single ''osgi-1'' mutation extended the time to flowering only under SD conditions. Under LD conditions the single ''hd1'' mutation induced early flowering, but ''osgi-1'' plants exhibited no phenotypic change<ref name="ref4" /><br />
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===Evolution===<br />
* There is some evolutionary conservation between rice circadian clock–related genes and those associated with Arabidopsis circadian clocks, where the circadian clock consists of interlocked subloops, it is also possible that ''Os-GI'' is a member of a subloop in rice circadian clocks<ref name="ref4" />.But there is a divergence in the photoperiodic pathways of these two species arises from the molecular mechanisms controlling the expression of ''FT/Hd3a'' by ''CO/Hd1(Se1)''<ref name="ref5" />(figure 3)<br />
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==Labs working on this gene==<br />
*Photosynthesis and Photobiology Research Unit, National Institute of Agrobiological Sciences, 2-1-2 Kannondai, Tsukuba 305-8602, Japan<br />
*Laboratory of Plant Molecular Genetics, Nara Institute of Science and Technology, 8916-5 Takayama, Ikoma 630-0101, Japan<br />
*Plant Genomics Research Unit, National Institute of Agrobiological Sciences, 2-1-2 Kannondai, Tsukuba 305-8602, Japan<br />
<br />
==Knowledge Extension==<br />
*The rice orthologues of the two key regulators of flowering time in Arabidopsis, ''CO'' and ''FT'', were recently shown to be important in the photoperiodic control of flowering in rice. The rice ''Hd1(Se1)''gene, which is an orthologue of the Arabidopsis ''CO'' gene, was required for the suppression of flowering under LD conditions and for the promotion of flowering under SD conditions. Furthermore, the rice ''Hd3a'' gene, an orthologue of the Arabidopsis ''FT'' gene, was shown to be an activator of flowering in rice. It was previously demonstrated that ''CO'' controls the flowering time by integrating signals from the circadian clock and light to regulate ''FT'' expression in Arabidopsis<ref name="ref5" />.Unlike Arabidopsis ''FT'', expression of ''Hd3a'' mRNA is more or less opposite to that of ''Hd1'' under inductive short-day conditions<ref name="ref6" />.<br />
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==References==<br />
<references><br />
<ref name="ref1">Song Lim Kim, Shinyoung Lee, Hyo Jung Kim, Hong Gil Nam, Gynheung An（2007）''OsMADS51'' is a short-day flowering promoter that functions upstream of ''Ehd1'', ''OsMADS14'', and ''Hd3a''.''Plant Physiology'' 145(4): 1484-1494.</ref><br />
<ref name="ref2">Hironori Itoh, Takeshi Izawa (2011) A study of phytohormone biosynthetic gene expression using a circadian clock-related mutant in rice. ''Plant Signaling & Behavior'' 6(12): 1932-1936.</ref> <br />
<ref name="ref3">Ryousuke Hayama, Takeshi Izawa, Ko Shimamoto (2002) Isolation of rice genes possibly involved in the photoperiodic control of flowering by a<br />
fluorescent differential display method. ''Plant and Cell Physiology'' 43(5): 494-504.</ref> <br />
<ref name="ref4">Takeshi Izawa, Motohiro Mihara, Yuji Suzuki, Meenu Gupta, Hironori Itoh, et al. (2011) ''Os-GIGANTEA'' confers robust diurnal rhythms on the global transcriptome of rice in the field. ''The Plant Cell'' 23(5): 1741-1755.</ref> <br />
<ref name="ref5">Ryosuke Hayama, Shuji Yokoi, Shojiro Tamaki, Masahiro Yano, Ko Shimamoto (2003) Adaptation of photoperiodic control pathways produces short-day flowering in rice. ''Nature'' 422(6933): 719-722.</ref> <br />
<ref name="ref6">Bong-Soo Shin, Jeong-Hyun Lee, Jeong-Hwan Lee, Hyun-Joo Jeong, Choong-Hyo Yun, Jeong-Kook Kim (2004) Circadian regulation of rice (''Oryza sativa L.'') ''CONSTANS-like'' gene transcripts. ''Molecules and Cells'' 17(1): 10-16.</ref><br />
<br />
==Structured Information==<br />
{{JaponicaGene|<br />
GeneName = Os01g0182600|<br />
Description = GIGANTEA protein|<br />
Version = NM_001048755.1 GI:115434923 GeneID:4325329|<br />
Length = 9083 bp|<br />
Definition = Oryza sativa Japonica Group Os01g0182600, complete gene.|<br />
Source = Oryza sativa Japonica Group<br />
<br />
ORGANISM Oryza sativa Japonica Group<br />
Eukaryota; Viridiplantae; Streptophyta; Embryophyta; Tracheophyta;<br />
Spermatophyta; Magnoliophyta; Liliopsida; Poales; Poaceae; BEP<br />
clade; Ehrhartoideae; Oryzeae; Oryza.<br />
|<br />
Chromosome = [[:category:Japonica Chromosome 1|Chromosome 1]]|<br />
AP = Chromosome 1:4328362..4337444|<br />
CDS = 4328725..4329030,4329123..4329215,4329333..4329551,4329752..4329820,4329925..4331421<br>,4332220..4332429,4332563..4332713,4332791..4333031,4333898..4333956<br>,4334379..4334653,4334813..4334955,4335034..4335088,4335181..4335258<br>,4335351..4335437|<br />
GCID = <gbrowseImage1><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage1>|<br />
GSID = <gbrowseImage2><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage2>|<br />
CDNA = <cdnaseq>atgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactgcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcatagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcggcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctga</cdnaseq>|<br />
AA = <aaseq>MSASNEKWIDGLQFSSLFWPPPQDSQQKQAQILAYVEYFGQFTA DSEQFPEDIAQLIQSCYPSKEKRLVDEVLATFVLHHPEHGHAVVHPILSRIIDGTLSY DRNGFPFMSFISLFSHTSEKEYSEQWALACGEILRVLTHYNRPIFKVDHQHSEAECSS TSDQASSCESMEKRANGSPRNEPDRKPLRPLSPWITDILLAAPLGIRSDYFRWCGGVM GKYAAGGELKPPTTAYSRGSGKHPQLMPSTPRWAVANGAGVILSVCDEEVARYETANL TAAAVPALLLPPPTTPLDEHLVAGLPPLEPYARLFHRYYAIATPSATQRLLFGLLEAP PSWAPDALDAAVQLVELLRAAEDYDSGMRLPKNWMHLHFLRAIGTAMSMRAGIAADTS AALLFRILSQPTLLFPPLRHAEGVELHHEPLGGYVSSYKRQLEVPASEATIDATAQGI ASMLCAHGPDVEWRICTIWEAAYGLLPLSSSAVDLPEIVVAAPLQPPTLSWSLYLPLL KVFEYLPRGSPSEACLMRIFVATVEAILRRTFPSETSEQSRKPRSQSKNLAVAELRTM IHSLFVESCASMDLASRLLFVVLTVCVSHQALPGGSKRPTGSDNHSSEEVTNDSRLTN GRNRCKKRQGPVATFDSYVLAAVCALSCELQLFPFISKNGNHSNLKDSIKIVIPGKTT GISNELHNSISSAILHTRRILGILEALFSLKPSSVGTSWSYSSNEIVAAAMVAAHVSE LFRRSRPCLNALSALKQCKWDAEISTRASSLYHLIDLHGKTVTSIVNKAEPLEAHLTL TPVKKDEPPIEEKNINSSDGGALEKKDASRSHRKNGFARPLLKCAEDVILNGDVASTS GKAIASLQVEASDLANFLTMDRNGGYRGSQTLLRSVLSEKQELCFSVVSLLWQKLIAS PEMQMSAESTSAHQGWRKVVDALCDIVSASPTKASAAIVLQAEKDLQPWIARDDEQGQ KMWRVNQRIVKLIAELMRNHDSPEALVILASASDLLLRATDGMLVDGEACTLPQLELL EVTARAVHLIVEWGDSGVSVADGLSNLLKCRLSTTIRCLSHPSAHVRALSMSVLRDIL NSGQINSSKLIQGEHRNGIQSPTYQCLAASIINWQADVERCIEWEAHSRRATGLTLAF LTAAAKELGCPLTC</aaseq>|<br />
DNA = <dnaseqindica>8415..8720#8230..8322#7894..8112#7625..7693#6024..7520#5016..5225#4732..4882#4414..4654#3489..3547#2792..3066#2490..2632#2357..2411#2187..2264#2008..2094#ccgagccgtttctctccctctctctctcctcctccttcccctgcgcctcctccaccgcccgtgtcgccgccgcgcgcgtctatcccctcgccgccgccgccgtctacgccgcgtgaggtccggattggttgaaaggctgatcgaggtaatgcttttttatttttcccgtgcggttttagtttttctgtgctgtgcttggtggttttttctttaaaactatttttaggctggttttgatgatggcctttctaaggttgtcgtgtatggggggcgaggtggttcgggtggatttgtggtgttatgttgaggggatttttgtttggggggagagagttttggtggtagatgcggttgggattttgaagacgtgaaagggaaattagttttgaaatttgacggcgaggttgagctgattagtattgtaataattcgtggcatcggatagatcaacgcgagatggatgcttattactgcgaatttcaggaggtttttgtggtttttttaggcgagtagcgttgattctagctggattttgctgcttcgtggaggcgtttagatgtggtgcatacggctgtggatgtaatttgtgcggggataattactgaaaattaattcgcagtaagctagctctgctttaatattttttcccctgaaacagtaacaagggatcccttcctcatttcgttaacagtggaagtgaatacagacaccaaccgatgagcagagacacctgtaattaaacaccaaaagaagaacctgttttgttcctatatattttccatttctatgctttaggtaatatactttctggttcggtgtcatagtacccgaaatagaatctttgcttgccaaaaaaagaaagaataacaagatagaacaacatcattaaattctgtcttagacaatacatttataatcgaccaacaaaaacagaccgatgagtgacgtggaataacaaatgaagtataaatgtcacgaaaaacacatctattgattttgtgtgataagccataggagcctacgaggaaccaagtgaatggttctgtacaccacatccattagcattaccataaatttgcctatttctatgatgccttccttaacatatttatcgacatgtgagcttgcaccataactaaatgttatggtaaactttaccttataaatgattgcaacatgtaccttttcatttcgataattaaaaacgagaaagcttccacctttttcctagatatattttttcgtttgattatgaatttagttgtacttgacaataaaataatgcgcttgtaagattatgcaattttcgttgcattacctccctctcttttcatttgtcagtttttttctactccctccgtttcatattttaagtcgtttgacttttttcctagtcagactttgttaagtttgacaaagtttatagtaaaatttagcaatatctaaaatattaaattggtttcatcaaatctaacattgaatatattttgataatatgttttttttgtgttgaaaatacttctatatttttctataaacttagtcaaacttggaagaaatttcagtaagaaaaaagtcaaacggcttataatatgaaacggagggagtaaatttttttttgcattaattaaaattcatgtattttcctccttttttggtttttgttgtgcgatatattacagtgtagagactctgaagctcatatggttggcattggttgatatgataacgtgtcacgttgtgagatgcagacccccgttatgggggttgattgaatgaagtggatagcactcttataactttcttgctcgaagtttcttgataccaaggcaagtagcttataagaagatttggactttggtggcataaatggatgaactaactggcggcttcaacgtggtgttgattacaggtgcttttattgctaactggagcagtttgtgtagtttctttctttctctttgagaattctctcagtttcttgggcaggctcttgagctaccaagtatgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggtaacattattcctatttgattcttgttagaaattctatatgttgaacatacttgcctcaatgggaaacttggttgcgtctcctctctgaaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcaggtactgtatctctaagcatgtttagtatcttctgtgataaaatctttctccagtgtgttaacttttgcattctttttcttgcatgcttgcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattaggtaatttgatccagtgatcaatacatgttccacaaacaattgaagaaagagtttgacatatttttctcatatttccagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgaggtaaggcccctactctaatggttagttccaatttattggttcatttctctgtgtttgataagattttgcctctgctgtgcatttcaatattatatctaaactgtgctgcttatagcttatcaaactactcttgctcataaagatgcaacttttatgcagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgagtcataataaatgataacatattattgttctgtggactgaaaatctgaaatcctcctttcttgagaaatacaacctcaattttatggtgatattgacaagaaggaagagcgcatataatttaaagttaaaacacggaaaataatagaagtatgtgagcagaacgcaggagcaaggggtggaactcataacaagttcaaacttcatgcatgttcataattccccaatgcaagtatggcaagaaaaatagacaacttttaaaagtatgctaatggaactgcatgcattgtgtaaacttgcttttgtcaagcatgttggaacatactatatatttgctgctgatgatgaaaaatttgcttgagtactcttctattgctaatggacgcttaccattccctgttttgtcttcaaatgcatcaggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactggttggtaaaatcagcaatggaattttctatcgtgaattttttataattattgtaacattaattactcccatgtctttttattctgttgagataccatgtgacaatgtctacaattgcatattttgcattctttggtagatcagttcccttcttaagatccccaaagggaaactggagtatcaataacttactccctcaaataaaaaaaaagtgatactttaggcataaacacattcttcaaaacaaaactttgaatagcagtttcttttgtaatatattaataaagcatggaaaaattataatgtgaagaatgtacattcagggacaaatatactcatcatttttgcatggttgatcttaatgtccatagagagattgatcagaccaaaagtttatttttgacaggagtgagtatggattataataggagtgaatggcaatatgctgtgaaagtgagcattttatatttgtttactactagcatttggtgacaaagcaaatgatgtatattgtaaatcaaagtaaacaagtagaacatgtcctgtggcgacaagacataacctacaaattaaagttagcctgattgtttattttactgggagcacattttttctagattctagagtacatattattaaaaaaggtgtgatatgtttgcaaattactttttaatggaagtttgtaaattactttgaaacttgcattcagtttttttgcttcttaaaaaacatagtgcatatgggctttatgagctcatactatgcatgtttttctgcattggagtgggacctttatctttttcttgcaactcggatgctaatgcttattataaacacaagttactaacatgtgcttgggtcattcagcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcataggtaagctcttatgatcctgtccactgttgtaccatgtgtactttccaaatgattgctaacatgctatttttcgtcagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcgggtatatcatacatctctctggctgttgtttcagtgctatcaatcaagatgacacatcatgcaatattatcataacagcacattatgtgttttgaatattttattgattgtcatagctttgttttatgttttagcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcaggtatgcagtgtttcctgcacctacttgctacctattctgtatttgatgtattcaatttcctttttatcagttttctcgaaatatatgctaataggcttgtcactaaacggctacattatccagactctgttctgaatttttttttcattaagtactgttagctgccagattacatcatcatgactaaagtagtttagctacaatattgcaaagctagatttgtctagcctctattgttttgaatagaaaaggcaggtatgcagtgtttcctgcacctacttgctacctattctgtatttgatgtattcaatttcctttttatcagttttctcgaaatatatgctaataggcttgtcactaaacggctacattatccagactctgttctgaatttttttttcattaagtactgttagctgccagattacatcatcatgactaaagtagtttagctacaatattgcaaagctagatttgtctagcctctattgttttgaatagaaaattaaatcttttggtcatttttttgtaaccttttaggagtctctgtatgcttgtcgtcaacatgctatgataagtttatatcagttcttttggagctatgatgtattcttgtatattttatgactgctaggatattatgttaaaactgttttaaaggtatcatcaaacaagttaatatagatattgctacctaccaaagttggaatttgattatgtcgtaaaatatatgcaccctacatgttttagcttcatattgacttatattttcatatgtgctccttcaactgcacaaagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggtatggtcattcctgcttataagataccagaataatgctatgaagaccgtccttggttgtcagcattcttatacacaatgataatctttgtttttcatgggtaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggtcaagatattgttattgttatatttacatacttgttatcatatttaaaagcttcatattgtatgtacctttgaaaaaaaaagttccatactgtagcttcttttgttctcatttcttggtttcaaagcatcagcatcattaccgatataaaatatacccatttcctcagtcatcttccccgactcttgtatctttcacaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagaggtaacaccttcattcatgaggtccttaggtgcaaatcccagtgaagaataatcagttctactttgtatggttcatctcgcttactgaccggttcatctcgggatcaattacgagcagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaaggcatgtcctcttctctgatctcactcacgttcatgccaaacttactcatcatcatccatgatccatcatcctgactgaatcttgatatgcagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctgacaaggccacaccatgactgtcactgcaagcagctgctgtaggatcagcgagtaggaaaggatgacttgccggccagtttcctgagatgtgatttaagactgatattagctgatgttcccaagcatgatacggggcttgctcccaagatgtgatttttactttacttttggttaatgtgctaccgctgacattagatgttcaagcatatgaaaactgcttgtgctgtaacttgtatgtctgtaacgtaaaaatctttccatatggtaaccggggaagcagaggtatatccttttctgctgttgtatctatttccagtttcctgctgaacttgcaaatgaatattcagcaatttccgttttgctc</dnaseqindica>|<br />
Link = [http://www.ncbi.nlm.nih.gov/nuccore/NM_001048755.1 RefSeq:Os01g0182600]|<br />
}}<br />
[[Category:Genes]]<br />
[[Category:Japonica mRNA]]<br />
[[Category:Oryza Sativa Japonica Group]]<br />
[[Category:Japonica Genes]]<br />
[[Category:Japonica Chromosome 1]]<br />
[[Category:Chromosome 1]]</div>Mikahttps://ngdc.cncb.ac.cn/ricewiki/index.php?title=Os01g0182600&diff=183852Os01g01826002014-06-11T04:02:08Z<p>Mika: /* Function */</p>
<hr />
<div>The rice gene of ''Gigantea(GI)'' is solo ortholog of ''GI'' in the Arabidopsis circadian system and related to photoperiodic control of flowering.<br />
<br />
==Annotated Information==<br />
===Function===<br />
*''OsGI'',a ''GI'' ortholog, regulates flowering time in response to photoperiodic conditions. Mutations in this gene delay flowering in inductive SDs (short-days),whereas flowering is marginally induced under LDs(long-days).In comparison, over expression of ''OsGI'' in transgenic rice causes late flowering in both SDs and LDs.In addition, ''OsGI'' has function to regulate ''Hd1'', a gene that also shows circadian rhythm<ref name="ref1" />. ''Os-GI'' affects gene expression of approximately 75% of genes among 27,201 genes and is required for strong amplitudes and proper phase-setting of global gene expression under natural field conditions<ref name="ref2" />. <br />
*The cDNA of ''OsGI'' firstly found in ''se5'' mutant that is completely photoperiod-insensitive, the expression patterns of rice and Arabidopsis ''GI'' gene are both photoperiodically regulated and are very similar under LD and SD conditions<ref name="ref3" />. [[File:Limeijia1.png|frame|'''Figure 1''' Possible roles of isolated genes in the photoperiodic regulation of flowering in rice, including ''OsGI'', may function to inhibit flowering under LD conditions.(from reference <ref name="ref3" />).]]''GI'' is required for phase-dependent ''CO''transcription through the interaction with FKF1, which leads to cycling DNA binding with one finger (CDF) degradation and consequent derepression of CO. In addition, it has been known that GI protein interacts with ZEITLUPE (ZTL) F-box protein to control the diurnal rhythm of TOC1 protein in regulation of the Arabidopsis circadian clock<ref name="ref4" />. <br />
*''Os-GI'' also has function to maintain bioactive GA level through the regulation of the GA-deactivating enzyme genes in rice, ''Os-GI'' represses major ''OsGA2ox'' genes in order to maintain a proper bioactive GA level, ''osgi-1'' plants showed a semi-dwarf phenotype with elevated ''OsGA2ox'' gene expression <ref name="ref4" /> .[[File:Limeijia1.png|frame|'''Figure 2''' WT rice and osgi-1 plants grown under natural field conditions and ''osgi-1'' plant exhibits Semi-dwarf phenotype(from reference <ref name="ref4" />).]]<br />
<br />
===Expression===<br />
*''GI'' encodes a nuclear protein that defines the proper amplitude and period length of circadian rhythms. ''GI'' expression is regulated by the circadian clock, with a peak in transcript levels at 8 to 10 h after dawn. The ''GI'' protein acts between the circadian oscillator and ''CO'' to promote flowering by increasing ''CO'' and ''FT'' mRNA abundance<ref name="ref1" />. The expression of the ''OsGI'' messenger RNA was circadian-controlled and that its temporal expression pattern was very similar to that of ''GI'' under both SD and LD conditions. Decreased ''OsGI'' expression led to early flowering under LD conditions and late flowering under SD conditions relative to the wild type, and that this early flowering phenotype under LD conditions was the opposite of that of the Arabidopsis ''gi'' mutants, confirming the opposite role of the ''GI'' gene in flowering under LD conditions in rice and Arabidopsis<ref name="ref5" />.<br />
<br />
===Mutation===<br />
*The single ''osgi-1'' mutation extended the time to flowering only under SD conditions. Under LD conditions the single ''hd1'' mutation induced early flowering, but ''osgi-1'' plants exhibited no phenotypic change<ref name="ref4" /><br />
<br />
===Evolution===<br />
* There is some evolutionary conservation between rice circadian clock–related genes and those associated with Arabidopsis circadian clocks, where the circadian clock consists of interlocked subloops, it is also possible that ''Os-GI'' is a member of a subloop in rice circadian clocks<ref name="ref4" />.But there is a divergence in the photoperiodic pathways of these two species arises from the molecular mechanisms controlling the expression of ''FT/Hd3a'' by ''CO/Hd1(Se1)''<ref name="ref5" />(figure 3)<br />
<br />
==Labs working on this gene==<br />
*Photosynthesis and Photobiology Research Unit, National Institute of Agrobiological Sciences, 2-1-2 Kannondai, Tsukuba 305-8602, Japan<br />
*Laboratory of Plant Molecular Genetics, Nara Institute of Science and Technology, 8916-5 Takayama, Ikoma 630-0101, Japan<br />
*Plant Genomics Research Unit, National Institute of Agrobiological Sciences, 2-1-2 Kannondai, Tsukuba 305-8602, Japan<br />
<br />
==Knowledge Extension==<br />
*The rice orthologues of the two key regulators of flowering time in Arabidopsis, ''CO'' and ''FT'', were recently shown to be important in the photoperiodic control of flowering in rice. The rice ''Hd1(Se1)''gene, which is an orthologue of the Arabidopsis ''CO'' gene, was required for the suppression of flowering under LD conditions and for the promotion of flowering under SD conditions. Furthermore, the rice ''Hd3a'' gene, an orthologue of the Arabidopsis ''FT'' gene, was shown to be an activator of flowering in rice. It was previously demonstrated that ''CO'' controls the flowering time by integrating signals from the circadian clock and light to regulate ''FT'' expression in Arabidopsis<ref name="ref5" />.Unlike Arabidopsis ''FT'', expression of ''Hd3a'' mRNA is more or less opposite to that of ''Hd1'' under inductive short-day conditions<ref name="ref6" />.<br />
<br />
==References==<br />
<references><br />
<ref name="ref1">Song Lim Kim, Shinyoung Lee, Hyo Jung Kim, Hong Gil Nam, Gynheung An（2007）''OsMADS51'' is a short-day flowering promoter that functions upstream of ''Ehd1'', ''OsMADS14'', and ''Hd3a''.''Plant Physiology'' 145(4): 1484-1494.</ref><br />
<ref name="ref2">Hironori Itoh, Takeshi Izawa (2011) A study of phytohormone biosynthetic gene expression using a circadian clock-related mutant in rice. ''Plant Signaling & Behavior'' 6(12): 1932-1936.</ref> <br />
<ref name="ref3">Ryousuke Hayama, Takeshi Izawa, Ko Shimamoto (2002) Isolation of rice genes possibly involved in the photoperiodic control of flowering by a<br />
fluorescent differential display method. ''Plant and Cell Physiology'' 43(5): 494-504.</ref> <br />
<ref name="ref4">Takeshi Izawa, Motohiro Mihara, Yuji Suzuki, Meenu Gupta, Hironori Itoh, et al. (2011) ''Os-GIGANTEA'' confers robust diurnal rhythms on the global transcriptome of rice in the field. ''The Plant Cell'' 23(5): 1741-1755.</ref> <br />
<ref name="ref5">Ryosuke Hayama, Shuji Yokoi, Shojiro Tamaki, Masahiro Yano, Ko Shimamoto (2003) Adaptation of photoperiodic control pathways produces short-day flowering in rice. ''Nature'' 422(6933): 719-722.</ref> <br />
<ref name="ref6">Bong-Soo Shin, Jeong-Hyun Lee, Jeong-Hwan Lee, Hyun-Joo Jeong, Choong-Hyo Yun, Jeong-Kook Kim (2004) Circadian regulation of rice (''Oryza sativa L.'') ''CONSTANS-like'' gene transcripts. ''Molecules and Cells'' 17(1): 10-16.</ref><br />
<br />
==Structured Information==<br />
{{JaponicaGene|<br />
GeneName = Os01g0182600|<br />
Description = GIGANTEA protein|<br />
Version = NM_001048755.1 GI:115434923 GeneID:4325329|<br />
Length = 9083 bp|<br />
Definition = Oryza sativa Japonica Group Os01g0182600, complete gene.|<br />
Source = Oryza sativa Japonica Group<br />
<br />
ORGANISM Oryza sativa Japonica Group<br />
Eukaryota; Viridiplantae; Streptophyta; Embryophyta; Tracheophyta;<br />
Spermatophyta; Magnoliophyta; Liliopsida; Poales; Poaceae; BEP<br />
clade; Ehrhartoideae; Oryzeae; Oryza.<br />
|<br />
Chromosome = [[:category:Japonica Chromosome 1|Chromosome 1]]|<br />
AP = Chromosome 1:4328362..4337444|<br />
CDS = 4328725..4329030,4329123..4329215,4329333..4329551,4329752..4329820,4329925..4331421<br>,4332220..4332429,4332563..4332713,4332791..4333031,4333898..4333956<br>,4334379..4334653,4334813..4334955,4335034..4335088,4335181..4335258<br>,4335351..4335437|<br />
GCID = <gbrowseImage1><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage1>|<br />
GSID = <gbrowseImage2><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage2>|<br />
CDNA = <cdnaseq>atgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactgcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcatagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcggcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctga</cdnaseq>|<br />
AA = <aaseq>MSASNEKWIDGLQFSSLFWPPPQDSQQKQAQILAYVEYFGQFTA DSEQFPEDIAQLIQSCYPSKEKRLVDEVLATFVLHHPEHGHAVVHPILSRIIDGTLSY DRNGFPFMSFISLFSHTSEKEYSEQWALACGEILRVLTHYNRPIFKVDHQHSEAECSS TSDQASSCESMEKRANGSPRNEPDRKPLRPLSPWITDILLAAPLGIRSDYFRWCGGVM GKYAAGGELKPPTTAYSRGSGKHPQLMPSTPRWAVANGAGVILSVCDEEVARYETANL TAAAVPALLLPPPTTPLDEHLVAGLPPLEPYARLFHRYYAIATPSATQRLLFGLLEAP PSWAPDALDAAVQLVELLRAAEDYDSGMRLPKNWMHLHFLRAIGTAMSMRAGIAADTS AALLFRILSQPTLLFPPLRHAEGVELHHEPLGGYVSSYKRQLEVPASEATIDATAQGI ASMLCAHGPDVEWRICTIWEAAYGLLPLSSSAVDLPEIVVAAPLQPPTLSWSLYLPLL KVFEYLPRGSPSEACLMRIFVATVEAILRRTFPSETSEQSRKPRSQSKNLAVAELRTM IHSLFVESCASMDLASRLLFVVLTVCVSHQALPGGSKRPTGSDNHSSEEVTNDSRLTN GRNRCKKRQGPVATFDSYVLAAVCALSCELQLFPFISKNGNHSNLKDSIKIVIPGKTT GISNELHNSISSAILHTRRILGILEALFSLKPSSVGTSWSYSSNEIVAAAMVAAHVSE LFRRSRPCLNALSALKQCKWDAEISTRASSLYHLIDLHGKTVTSIVNKAEPLEAHLTL TPVKKDEPPIEEKNINSSDGGALEKKDASRSHRKNGFARPLLKCAEDVILNGDVASTS GKAIASLQVEASDLANFLTMDRNGGYRGSQTLLRSVLSEKQELCFSVVSLLWQKLIAS PEMQMSAESTSAHQGWRKVVDALCDIVSASPTKASAAIVLQAEKDLQPWIARDDEQGQ KMWRVNQRIVKLIAELMRNHDSPEALVILASASDLLLRATDGMLVDGEACTLPQLELL EVTARAVHLIVEWGDSGVSVADGLSNLLKCRLSTTIRCLSHPSAHVRALSMSVLRDIL NSGQINSSKLIQGEHRNGIQSPTYQCLAASIINWQADVERCIEWEAHSRRATGLTLAF LTAAAKELGCPLTC</aaseq>|<br />
DNA = <dnaseqindica>8415..8720#8230..8322#7894..8112#7625..7693#6024..7520#5016..5225#4732..4882#4414..4654#3489..3547#2792..3066#2490..2632#2357..2411#2187..2264#2008..2094#ccgagccgtttctctccctctctctctcctcctccttcccctgcgcctcctccaccgcccgtgtcgccgccgcgcgcgtctatcccctcgccgccgccgccgtctacgccgcgtgaggtccggattggttgaaaggctgatcgaggtaatgcttttttatttttcccgtgcggttttagtttttctgtgctgtgcttggtggttttttctttaaaactatttttaggctggttttgatgatggcctttctaaggttgtcgtgtatggggggcgaggtggttcgggtggatttgtggtgttatgttgaggggatttttgtttggggggagagagttttggtggtagatgcggttgggattttgaagacgtgaaagggaaattagttttgaaatttgacggcgaggttgagctgattagtattgtaataattcgtggcatcggatagatcaacgcgagatggatgcttattactgcgaatttcaggaggtttttgtggtttttttaggcgagtagcgttgattctagctggattttgctgcttcgtggaggcgtttagatgtggtgcatacggctgtggatgtaatttgtgcggggataattactgaaaattaattcgcagtaagctagctctgctttaatattttttcccctgaaacagtaacaagggatcccttcctcatttcgttaacagtggaagtgaatacagacaccaaccgatgagcagagacacctgtaattaaacaccaaaagaagaacctgttttgttcctatatattttccatttctatgctttaggtaatatactttctggttcggtgtcatagtacccgaaatagaatctttgcttgccaaaaaaagaaagaataacaagatagaacaacatcattaaattctgtcttagacaatacatttataatcgaccaacaaaaacagaccgatgagtgacgtggaataacaaatgaagtataaatgtcacgaaaaacacatctattgattttgtgtgataagccataggagcctacgaggaaccaagtgaatggttctgtacaccacatccattagcattaccataaatttgcctatttctatgatgccttccttaacatatttatcgacatgtgagcttgcaccataactaaatgttatggtaaactttaccttataaatgattgcaacatgtaccttttcatttcgataattaaaaacgagaaagcttccacctttttcctagatatattttttcgtttgattatgaatttagttgtacttgacaataaaataatgcgcttgtaagattatgcaattttcgttgcattacctccctctcttttcatttgtcagtttttttctactccctccgtttcatattttaagtcgtttgacttttttcctagtcagactttgttaagtttgacaaagtttatagtaaaatttagcaatatctaaaatattaaattggtttcatcaaatctaacattgaatatattttgataatatgttttttttgtgttgaaaatacttctatatttttctataaacttagtcaaacttggaagaaatttcagtaagaaaaaagtcaaacggcttataatatgaaacggagggagtaaatttttttttgcattaattaaaattcatgtattttcctccttttttggtttttgttgtgcgatatattacagtgtagagactctgaagctcatatggttggcattggttgatatgataacgtgtcacgttgtgagatgcagacccccgttatgggggttgattgaatgaagtggatagcactcttataactttcttgctcgaagtttcttgataccaaggcaagtagcttataagaagatttggactttggtggcataaatggatgaactaactggcggcttcaacgtggtgttgattacaggtgcttttattgctaactggagcagtttgtgtagtttctttctttctctttgagaattctctcagtttcttgggcaggctcttgagctaccaagtatgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggtaacattattcctatttgattcttgttagaaattctatatgttgaacatacttgcctcaatgggaaacttggttgcgtctcctctctgaaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcaggtactgtatctctaagcatgtttagtatcttctgtgataaaatctttctccagtgtgttaacttttgcattctttttcttgcatgcttgcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattaggtaatttgatccagtgatcaatacatgttccacaaacaattgaagaaagagtttgacatatttttctcatatttccagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgaggtaaggcccctactctaatggttagttccaatttattggttcatttctctgtgtttgataagattttgcctctgctgtgcatttcaatattatatctaaactgtgctgcttatagcttatcaaactactcttgctcataaagatgcaacttttatgcagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgagtcataataaatgataacatattattgttctgtggactgaaaatctgaaatcctcctttcttgagaaatacaacctcaattttatggtgatattgacaagaaggaagagcgcatataatttaaagttaaaacacggaaaataatagaagtatgtgagcagaacgcaggagcaaggggtggaactcataacaagttcaaacttcatgcatgttcataattccccaatgcaagtatggcaagaaaaatagacaacttttaaaagtatgctaatggaactgcatgcattgtgtaaacttgcttttgtcaagcatgttggaacatactatatatttgctgctgatgatgaaaaatttgcttgagtactcttctattgctaatggacgcttaccattccctgttttgtcttcaaatgcatcaggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactggttggtaaaatcagcaatggaattttctatcgtgaattttttataattattgtaacattaattactcccatgtctttttattctgttgagataccatgtgacaatgtctacaattgcatattttgcattctttggtagatcagttcccttcttaagatccccaaagggaaactggagtatcaataacttactccctcaaataaaaaaaaagtgatactttaggcataaacacattcttcaaaacaaaactttgaatagcagtttcttttgtaatatattaataaagcatggaaaaattataatgtgaagaatgtacattcagggacaaatatactcatcatttttgcatggttgatcttaatgtccatagagagattgatcagaccaaaagtttatttttgacaggagtgagtatggattataataggagtgaatggcaatatgctgtgaaagtgagcattttatatttgtttactactagcatttggtgacaaagcaaatgatgtatattgtaaatcaaagtaaacaagtagaacatgtcctgtggcgacaagacataacctacaaattaaagttagcctgattgtttattttactgggagcacattttttctagattctagagtacatattattaaaaaaggtgtgatatgtttgcaaattactttttaatggaagtttgtaaattactttgaaacttgcattcagtttttttgcttcttaaaaaacatagtgcatatgggctttatgagctcatactatgcatgtttttctgcattggagtgggacctttatctttttcttgcaactcggatgctaatgcttattataaacacaagttactaacatgtgcttgggtcattcagcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcataggtaagctcttatgatcctgtccactgttgtaccatgtgtactttccaaatgattgctaacatgctatttttcgtcagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtac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/>
Link = [http://www.ncbi.nlm.nih.gov/nuccore/NM_001048755.1 RefSeq:Os01g0182600]|<br />
}}<br />
[[Category:Genes]]<br />
[[Category:Japonica mRNA]]<br />
[[Category:Oryza Sativa Japonica Group]]<br />
[[Category:Japonica Genes]]<br />
[[Category:Japonica Chromosome 1]]<br />
[[Category:Chromosome 1]]</div>Mikahttps://ngdc.cncb.ac.cn/ricewiki/index.php?title=Os01g0182600&diff=183833Os01g01826002014-06-11T03:23:28Z<p>Mika: /* Function */</p>
<hr />
<div>The rice gene of ''Gigantea(GI)'' is solo ortholog of ''GI'' in the Arabidopsis circadian system and related to photoperiodic control of flowering.<br />
<br />
==Annotated Information==<br />
===Function===<br />
*''OsGI'',a ''GI'' ortholog, regulates flowering time in response to photoperiodic conditions. Mutations in this gene delay flowering in inductive SDs (short-days),whereas flowering is marginally induced under LDs(long-days).In comparison, over expression of ''OsGI'' in transgenic rice causes late flowering in both SDs and LDs.In addition, ''OsGI'' has function to regulate ''Hd1'', a gene that also shows circadian rhythm<ref name="ref1" />. ''Os-GI'' affects gene expression of approximately 75% of genes among 27,201 genes and is required for strong amplitudes and proper phase-setting of global gene expression under natural field conditions<ref name="ref2" />. <br />
*The cDNA of ''OsGI'' firstly found in ''se5'' mutant that is completely photoperiod-insensitive, the expression patterns of rice and Arabidopsis ''GI'' gene are both photoperiodically regulated and are very similar under LD and SD conditions(FIG1)<ref name="ref3" />. ''GI'' is required for phase-dependent ''CO''transcription through the interaction with FKF1, which leads to cycling DNA binding with one finger (CDF) degradation and consequent derepression of CO. In addition, it has been known that GI protein interacts with ZEITLUPE (ZTL) F-box protein to control the diurnal rhythm of TOC1 protein in regulation of the Arabidopsis circadian clock<ref name="ref4" />. <br />
*''Os-GI'' also has function to maintain bioactive GA level through the regulation of the GA-deactivating enzyme genes in rice, ''Os-GI'' represses major ''OsGA2ox'' genes in order to maintain a proper bioactive GA level, ''osgi-1'' plants showed a semi-dwarf phenotype with elevated ''OsGA2ox'' gene expression <ref name="ref4" /> .[[File:Limeijia1.png|frame|'''Figure 2''' WT rice and osgi-1 plants grown under natural field conditions and osgi-1plant exhibits Semi-dwarf phenotype(from reference <ref name="ref4" />).]]<br />
<br />
===Expression===<br />
*''GI'' encodes a nuclear protein that defines the proper amplitude and period length of circadian rhythms. ''GI'' expression is regulated by the circadian clock, with a peak in transcript levels at 8 to 10 h after dawn. The ''GI'' protein acts between the circadian oscillator and ''CO'' to promote flowering by increasing ''CO'' and ''FT'' mRNA abundance<ref name="ref1" />. The expression of the ''OsGI'' messenger RNA was circadian-controlled and that its temporal expression pattern was very similar to that of ''GI'' under both SD and LD conditions. Decreased ''OsGI'' expression led to early flowering under LD conditions and late flowering under SD conditions relative to the wild type, and that this early flowering phenotype under LD conditions was the opposite of that of the Arabidopsis ''gi'' mutants, confirming the opposite role of the ''GI'' gene in flowering under LD conditions in rice and Arabidopsis<ref name="ref5" />.<br />
<br />
===Mutation===<br />
*The single ''osgi-1'' mutation extended the time to flowering only under SD conditions. Under LD conditions the single ''hd1'' mutation induced early flowering, but ''osgi-1'' plants exhibited no phenotypic change<ref name="ref4" /><br />
<br />
===Evolution===<br />
* There is some evolutionary conservation between rice circadian clock–related genes and those associated with Arabidopsis circadian clocks, where the circadian clock consists of interlocked subloops, it is also possible that ''Os-GI'' is a member of a subloop in rice circadian clocks<ref name="ref4" />.But there is a divergence in the photoperiodic pathways of these two species arises from the molecular mechanisms controlling the expression of ''FT/Hd3a'' by ''CO/Hd1(Se1)''<ref name="ref5" />(figure 3)<br />
<br />
==Labs working on this gene==<br />
*Photosynthesis and Photobiology Research Unit, National Institute of Agrobiological Sciences, 2-1-2 Kannondai, Tsukuba 305-8602, Japan<br />
*Laboratory of Plant Molecular Genetics, Nara Institute of Science and Technology, 8916-5 Takayama, Ikoma 630-0101, Japan<br />
*Plant Genomics Research Unit, National Institute of Agrobiological Sciences, 2-1-2 Kannondai, Tsukuba 305-8602, Japan<br />
<br />
==Knowledge Extension==<br />
*The rice orthologues of the two key regulators of flowering time in Arabidopsis, ''CO'' and ''FT'', were recently shown to be important in the photoperiodic control of flowering in rice. The rice ''Hd1(Se1)''gene, which is an orthologue of the Arabidopsis ''CO'' gene, was required for the suppression of flowering under LD conditions and for the promotion of flowering under SD conditions. Furthermore, the rice ''Hd3a'' gene, an orthologue of the Arabidopsis ''FT'' gene, was shown to be an activator of flowering in rice. It was previously demonstrated that ''CO'' controls the flowering time by integrating signals from the circadian clock and light to regulate ''FT'' expression in Arabidopsis<ref name="ref5" />.Unlike Arabidopsis ''FT'', expression of ''Hd3a'' mRNA is more or less opposite to that of ''Hd1'' under inductive short-day conditions<ref name="ref6" />.<br />
<br />
==References==<br />
<references><br />
<ref name="ref1">Song Lim Kim, Shinyoung Lee, Hyo Jung Kim, Hong Gil Nam, Gynheung An（2007）''OsMADS51'' is a short-day flowering promoter that functions upstream of ''Ehd1'', ''OsMADS14'', and ''Hd3a''.''Plant Physiology'' 145(4): 1484-1494.</ref><br />
<ref name="ref2">Hironori Itoh, Takeshi Izawa (2011) A study of phytohormone biosynthetic gene expression using a circadian clock-related mutant in rice. ''Plant Signaling & Behavior'' 6(12): 1932-1936.</ref> <br />
<ref name="ref3">Ryousuke Hayama, Takeshi Izawa, Ko Shimamoto (2002) Isolation of rice genes possibly involved in the photoperiodic control of flowering by a<br />
fluorescent differential display method. ''Plant and Cell Physiology'' 43(5): 494-504.</ref> <br />
<ref name="ref4">Takeshi Izawa, Motohiro Mihara, Yuji Suzuki, Meenu Gupta, Hironori Itoh, et al. (2011) ''Os-GIGANTEA'' confers robust diurnal rhythms on the global transcriptome of rice in the field. ''The Plant Cell'' 23(5): 1741-1755.</ref> <br />
<ref name="ref5">Ryosuke Hayama, Shuji Yokoi, Shojiro Tamaki, Masahiro Yano, Ko Shimamoto (2003) Adaptation of photoperiodic control pathways produces short-day flowering in rice. ''Nature'' 422(6933): 719-722.</ref> <br />
<ref name="ref6">Bong-Soo Shin, Jeong-Hyun Lee, Jeong-Hwan Lee, Hyun-Joo Jeong, Choong-Hyo Yun, Jeong-Kook Kim (2004) Circadian regulation of rice (''Oryza sativa L.'') ''CONSTANS-like'' gene transcripts. ''Molecules and Cells'' 17(1): 10-16.</ref><br />
<br />
==Structured Information==<br />
{{JaponicaGene|<br />
GeneName = Os01g0182600|<br />
Description = GIGANTEA protein|<br />
Version = NM_001048755.1 GI:115434923 GeneID:4325329|<br />
Length = 9083 bp|<br />
Definition = Oryza sativa Japonica Group Os01g0182600, complete gene.|<br />
Source = Oryza sativa Japonica Group<br />
<br />
ORGANISM Oryza sativa Japonica Group<br />
Eukaryota; Viridiplantae; Streptophyta; Embryophyta; Tracheophyta;<br />
Spermatophyta; Magnoliophyta; Liliopsida; Poales; Poaceae; BEP<br />
clade; Ehrhartoideae; Oryzeae; Oryza.<br />
|<br />
Chromosome = [[:category:Japonica Chromosome 1|Chromosome 1]]|<br />
AP = Chromosome 1:4328362..4337444|<br />
CDS = 4328725..4329030,4329123..4329215,4329333..4329551,4329752..4329820,4329925..4331421<br>,4332220..4332429,4332563..4332713,4332791..4333031,4333898..4333956<br>,4334379..4334653,4334813..4334955,4335034..4335088,4335181..4335258<br>,4335351..4335437|<br />
GCID = <gbrowseImage1><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage1>|<br />
GSID = <gbrowseImage2><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage2>|<br />
CDNA = <cdnaseq>atgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactgcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcatagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcggcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctga</cdnaseq>|<br />
AA = <aaseq>MSASNEKWIDGLQFSSLFWPPPQDSQQKQAQILAYVEYFGQFTA DSEQFPEDIAQLIQSCYPSKEKRLVDEVLATFVLHHPEHGHAVVHPILSRIIDGTLSY DRNGFPFMSFISLFSHTSEKEYSEQWALACGEILRVLTHYNRPIFKVDHQHSEAECSS TSDQASSCESMEKRANGSPRNEPDRKPLRPLSPWITDILLAAPLGIRSDYFRWCGGVM GKYAAGGELKPPTTAYSRGSGKHPQLMPSTPRWAVANGAGVILSVCDEEVARYETANL TAAAVPALLLPPPTTPLDEHLVAGLPPLEPYARLFHRYYAIATPSATQRLLFGLLEAP PSWAPDALDAAVQLVELLRAAEDYDSGMRLPKNWMHLHFLRAIGTAMSMRAGIAADTS AALLFRILSQPTLLFPPLRHAEGVELHHEPLGGYVSSYKRQLEVPASEATIDATAQGI ASMLCAHGPDVEWRICTIWEAAYGLLPLSSSAVDLPEIVVAAPLQPPTLSWSLYLPLL KVFEYLPRGSPSEACLMRIFVATVEAILRRTFPSETSEQSRKPRSQSKNLAVAELRTM IHSLFVESCASMDLASRLLFVVLTVCVSHQALPGGSKRPTGSDNHSSEEVTNDSRLTN GRNRCKKRQGPVATFDSYVLAAVCALSCELQLFPFISKNGNHSNLKDSIKIVIPGKTT GISNELHNSISSAILHTRRILGILEALFSLKPSSVGTSWSYSSNEIVAAAMVAAHVSE LFRRSRPCLNALSALKQCKWDAEISTRASSLYHLIDLHGKTVTSIVNKAEPLEAHLTL TPVKKDEPPIEEKNINSSDGGALEKKDASRSHRKNGFARPLLKCAEDVILNGDVASTS GKAIASLQVEASDLANFLTMDRNGGYRGSQTLLRSVLSEKQELCFSVVSLLWQKLIAS PEMQMSAESTSAHQGWRKVVDALCDIVSASPTKASAAIVLQAEKDLQPWIARDDEQGQ KMWRVNQRIVKLIAELMRNHDSPEALVILASASDLLLRATDGMLVDGEACTLPQLELL EVTARAVHLIVEWGDSGVSVADGLSNLLKCRLSTTIRCLSHPSAHVRALSMSVLRDIL NSGQINSSKLIQGEHRNGIQSPTYQCLAASIINWQADVERCIEWEAHSRRATGLTLAF LTAAAKELGCPLTC</aaseq>|<br />
DNA = <dnaseqindica>8415..8720#8230..8322#7894..8112#7625..7693#6024..7520#5016..5225#4732..4882#4414..4654#3489..3547#2792..3066#2490..2632#2357..2411#2187..2264#2008..2094#ccgagccgtttctctccctctctctctcctcctccttcccctgcgcctcctccaccgcccgtgtcgccgccgcgcgcgtctatcccctcgccgccgccgccgtctacgccgcgtgaggtccggattggttgaaaggctgatcgaggtaatgcttttttatttttcccgtgcggttttagtttttctgtgctgtgcttggtggttttttctttaaaactatttttaggctggttttgatgatggcctttctaaggttgtcgtgtatggggggcgaggtggttcgggtggatttgtggtgttatgttgaggggatttttgtttggggggagagagttttggtggtagatgcggttgggattttgaagacgtgaaagggaaattagttttgaaatttgacggcgaggttgagctgattagtattgtaataattcgtggcatcggatagatcaacgcgagatggatgcttattactgcgaatttcaggaggtttttgtggtttttttaggcgagtagcgttgattctagctggattttgctgcttcgtggaggcgtttagatgtggtgcatacggctgtggatgtaatttgtgcggggataattactgaaaattaattcgcagtaagctagctctgctttaatattttttcccctgaaacagtaacaagggatcccttcctcatttcgttaacagtggaagtgaatacagacaccaaccgatgagcagagacacctgtaattaaacaccaaaagaagaacctgttttgttcctatatattttccatttctatgctttaggtaatatactttctggttcggtgtcatagtacccgaaatagaatctttgcttgccaaaaaaagaaagaataacaagatagaacaacatcattaaattctgtcttagacaatacatttataatcgaccaacaaaaacagaccgatgagtgacgtggaataacaaatgaagtataaatgtcacgaaaaacacatctattgattttgtgtgataagccataggagcctacgaggaaccaagtgaatggttctgtacaccacatccattagcattaccataaatttgcctatttctatgatgccttccttaacatatttatcgacatgtgagcttgcaccataactaaatgttatggtaaactttaccttataaatgattgcaacatgtaccttttcatttcgataattaaaaacgagaaagcttccacctttttcctagatatattttttcgtttgattatgaatttagttgtacttgacaataaaataatgcgcttgtaagattatgcaattttcgttgcattacctccctctcttttcatttgtcagtttttttctactccctccgtttcatattttaagtcgtttgacttttttcctagtcagactttgttaagtttgacaaagtttatagtaaaatttagcaatatctaaaatattaaattggtttcatcaaatctaacattgaatatattttgataatatgttttttttgtgttgaaaatacttctatatttttctataaacttagtcaaacttggaagaaatttcagtaagaaaaaagtcaaacggcttataatatgaaacggagggagtaaatttttttttgcattaattaaaattcatgtattttcctccttttttggtttttgttgtgcgatatattacagtgtagagactctgaagctcatatggttggcattggttgatatgataacgtgtcacgttgtgagatgcagacccccgttatgggggttgattgaatgaagtggatagcactcttataactttcttgctcgaagtttcttgataccaaggcaagtagcttataagaagatttggactttggtggcataaatggatgaactaactggcggcttcaacgtggtgttgattacaggtgcttttattgctaactggagcagtttgtgtagtttctttctttctctttgagaattctctcagtttcttgggcaggctcttgagctaccaagtatgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggtaacattattcctatttgattcttgttagaaattctatatgttgaacatacttgcctcaatgggaaacttggttgcgtctcctctctgaaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcaggtactgtatctctaagcatgtttagtatcttctgtgataaaatctttctccagtgtgttaacttttgcattctttttcttgcatgcttgcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattaggtaatttgatccagtgatcaatacatgttccacaaacaattgaagaaagagtttgacatatttttctcatatttccagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgaggtaaggcccctactctaatggttagttccaatttattggttcatttctctgtgtttgataagattttgcctctgctgtgcatttcaatattatatctaaactgtgctgcttatagcttatcaaactactcttgctcataaagatgcaacttttatgcagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgagtcataataaatgataacatattattgttctgtggactgaaaatctgaaatcctcctttcttgagaaatacaacctcaattttatggtgatattgacaagaaggaagagcgcatataatttaaagttaaaacacggaaaataatagaagtatgtgagcagaacgcaggagcaaggggtggaactcataacaagttcaaacttcatgcatgttcataattccccaatgcaagtatggcaagaaaaatagacaacttttaaaagtatgctaatggaactgcatgcattgtgtaaacttgcttttgtcaagcatgttggaacatactatatatttgctgctgatgatgaaaaatttgcttgagtactcttctattgctaatggacgcttaccattccctgttttgtcttcaaatgcatcaggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactggttggtaaaatcagcaatggaattttctatcgtgaattttttataattattgtaacattaattactcccatgtctttttattctgttgagataccatgtgacaatgtctacaattgcatattttgcattctttggtagatcagttcccttcttaagatccccaaagggaaactggagtatcaataacttactccctcaaataaaaaaaaagtgatactttaggcataaacacattcttcaaaacaaaactttgaatagcagtttcttttgtaatatattaataaagcatggaaaaattataatgtgaagaatgtacattcagggacaaatatactcatcatttttgcatggttgatcttaatgtccatagagagattgatcagaccaaaagtttatttttgacaggagtgagtatggattataataggagtgaatggcaatatgctgtgaaagtgagcattttatatttgtttactactagcatttggtgacaaagcaaatgatgtatattgtaaatcaaagtaaacaagtagaacatgtcctgtggcgacaagacataacctacaaattaaagttagcctgattgtttattttactgggagcacattttttctagattctagagtacatattattaaaaaaggtgtgatatgtttgcaaattactttttaatggaagtttgtaaattactttgaaacttgcattcagtttttttgcttcttaaaaaacatagtgcatatgggctttatgagctcatactatgcatgtttttctgcattggagtgggacctttatctttttcttgcaactcggatgctaatgcttattataaacacaagttactaacatgtgcttgggtcattcagcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcataggtaagctcttatgatcctgtccactgttgtaccatgtgtactttccaaatgattgctaacatgctatttttcgtcagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcgggtatatcatacatctctctggctgttgtttcagtgctatcaatcaagatgacacatcatgcaatattatcataacagcacattatgtgttttgaatattttattgattgtcatagctttgttttatgttttagcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcaggtatgcagtgtttcctgcacctacttgctacctattctgtatttgatgtattcaatttcctttttatcagttttctcgaaatatatgctaataggcttgtcactaaacggctacattatccagactctgttctgaatttttttttcattaagtactgttagctgccagattacatcatcatgactaaagtagtttagctacaatattgcaaagctagatttgtctagcctctattgttttgaatagaaaaggcaggtatgcagtgtttcctgcacctacttgctacctattctgtatttgatgtattcaatttcctttttatcagttttctcgaaatatatgctaataggcttgtcactaaacggctacattatccagactctgttctgaatttttttttcattaagtactgttagctgccagattacatcatcatgactaaagtagtttagctacaatattgcaaagctagatttgtctagcctctattgttttgaatagaaaattaaatcttttggtcatttttttgtaaccttttaggagtctctgtatgcttgtcgtcaacatgctatgataagtttatatcagttcttttggagctatgatgtattcttgtatattttatgactgctaggatattatgttaaaactgttttaaaggtatcatcaaacaagttaatatagatattgctacctaccaaagttggaatttgattatgtcgtaaaatatatgcaccctacatgttttagcttcatattgacttatattttcatatgtgctccttcaactgcacaaagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggtatggtcattcctgcttataagataccagaataatgctatgaagaccgtccttggttgtcagcattcttatacacaatgataatctttgtttttcatgggtaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggtcaagatattgttattgttatatttacatacttgttatcatatttaaaagcttcatattgtatgtacctttgaaaaaaaaagttccatactgtagcttcttttgttctcatttcttggtttcaaagcatcagcatcattaccgatataaaatatacccatttcctcagtcatcttccccgactcttgtatctttcacaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagaggtaacaccttcattcatgaggtccttaggtgcaaatcccagtgaagaataatcagttctactttgtatggttcatctcgcttactgaccggttcatctcgggatcaattacgagcagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaaggcatgtcctcttctctgatctcactcacgttcatgccaaacttactcatcatcatccatgatccatcatcctgactgaatcttgatatgcagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctgacaaggccacaccatgactgtcactgcaagcagctgctgtaggatcagcgagtaggaaaggatgacttgccggccagtttcctgagatgtgatttaagactgatattagctgatgttcccaagcatgatacggggcttgctcccaagatgtgatttttactttacttttggttaatgtgctaccgctgacattagatgttcaagcatatgaaaactgcttgtgctgtaacttgtatgtctgtaacgtaaaaatctttccatatggtaaccggggaagcagaggtatatccttttctgctgttgtatctatttccagtttcctgctgaacttgcaaatgaatattcagcaatttccgttttgctc</dnaseqindica>|<br />
Link = [http://www.ncbi.nlm.nih.gov/nuccore/NM_001048755.1 RefSeq:Os01g0182600]|<br />
}}<br />
[[Category:Genes]]<br />
[[Category:Japonica mRNA]]<br />
[[Category:Oryza Sativa Japonica Group]]<br />
[[Category:Japonica Genes]]<br />
[[Category:Japonica Chromosome 1]]<br />
[[Category:Chromosome 1]]</div>Mikahttps://ngdc.cncb.ac.cn/ricewiki/index.php?title=Os01g0182600&diff=183830Os01g01826002014-06-11T03:21:19Z<p>Mika: /* Annotated Information */</p>
<hr />
<div>The rice gene of ''Gigantea(GI)'' is solo ortholog of ''GI'' in the Arabidopsis circadian system and related to photoperiodic control of flowering.<br />
<br />
==Annotated Information==<br />
===Function===<br />
*''OsGI'',a ''GI'' ortholog, regulates flowering time in response to photoperiodic conditions. Mutations in this gene delay flowering in inductive SDs (short-days),whereas flowering is marginally induced under LDs(long-days).In comparison, over expression of ''OsGI'' in transgenic rice causes late flowering in both SDs and LDs.In addition, ''OsGI'' has function to regulate ''Hd1'', a gene that also shows circadian rhythm<ref name="ref1" />. ''Os-GI'' affects gene expression of approximately 75% of genes among 27,201 genes and is required for strong amplitudes and proper phase-setting of global gene expression under natural field conditions<ref name="ref2" />. <br />
*The cDNA of ''OsGI'' firstly found in ''se5'' mutant that is completely photoperiod-insensitive, the expression patterns of rice and Arabidopsis ''GI'' gene are both photoperiodically regulated and are very similar under LD and SD conditions(FIG1)<ref name="ref3" />. ''GI'' is required for phase-dependent ''CO''transcription through the interaction with FKF1, which leads to cycling DNA binding with one finger (CDF) degradation and consequent derepression of CO. In addition, it has been known that GI protein interacts with ZEITLUPE (ZTL) F-box protein to control the diurnal rhythm of TOC1 protein in regulation of the Arabidopsis circadian clock<ref name="ref4" />. <br />
*''Os-GI'' also has function to maintain bioactive GA level through the regulation of the GA-deactivating enzyme genes in rice, ''Os-GI'' represses major ''OsGA2ox'' genes in order to maintain a proper bioactive GA level, ''osgi-1'' plants showed a semi-dwarf phenotype with elevated ''OsGA2ox'' gene expression <ref name="ref4" /> .[[File:Limeijia1.png|frame|'''Figure 2''' WT rice and osgi-1 plants grown under natural field conditions and osgi-1plant exhibits Semi-dwarf phenotype.]]<br />
<br />
===Expression===<br />
*''GI'' encodes a nuclear protein that defines the proper amplitude and period length of circadian rhythms. ''GI'' expression is regulated by the circadian clock, with a peak in transcript levels at 8 to 10 h after dawn. The ''GI'' protein acts between the circadian oscillator and ''CO'' to promote flowering by increasing ''CO'' and ''FT'' mRNA abundance<ref name="ref1" />. The expression of the ''OsGI'' messenger RNA was circadian-controlled and that its temporal expression pattern was very similar to that of ''GI'' under both SD and LD conditions. Decreased ''OsGI'' expression led to early flowering under LD conditions and late flowering under SD conditions relative to the wild type, and that this early flowering phenotype under LD conditions was the opposite of that of the Arabidopsis ''gi'' mutants, confirming the opposite role of the ''GI'' gene in flowering under LD conditions in rice and Arabidopsis<ref name="ref5" />.<br />
<br />
===Mutation===<br />
*The single ''osgi-1'' mutation extended the time to flowering only under SD conditions. Under LD conditions the single ''hd1'' mutation induced early flowering, but ''osgi-1'' plants exhibited no phenotypic change<ref name="ref4" /><br />
<br />
===Evolution===<br />
* There is some evolutionary conservation between rice circadian clock–related genes and those associated with Arabidopsis circadian clocks, where the circadian clock consists of interlocked subloops, it is also possible that ''Os-GI'' is a member of a subloop in rice circadian clocks<ref name="ref4" />.But there is a divergence in the photoperiodic pathways of these two species arises from the molecular mechanisms controlling the expression of ''FT/Hd3a'' by ''CO/Hd1(Se1)''<ref name="ref5" />(figure 3)<br />
<br />
==Labs working on this gene==<br />
*Photosynthesis and Photobiology Research Unit, National Institute of Agrobiological Sciences, 2-1-2 Kannondai, Tsukuba 305-8602, Japan<br />
*Laboratory of Plant Molecular Genetics, Nara Institute of Science and Technology, 8916-5 Takayama, Ikoma 630-0101, Japan<br />
*Plant Genomics Research Unit, National Institute of Agrobiological Sciences, 2-1-2 Kannondai, Tsukuba 305-8602, Japan<br />
<br />
==Knowledge Extension==<br />
*The rice orthologues of the two key regulators of flowering time in Arabidopsis, ''CO'' and ''FT'', were recently shown to be important in the photoperiodic control of flowering in rice. The rice ''Hd1(Se1)''gene, which is an orthologue of the Arabidopsis ''CO'' gene, was required for the suppression of flowering under LD conditions and for the promotion of flowering under SD conditions. Furthermore, the rice ''Hd3a'' gene, an orthologue of the Arabidopsis ''FT'' gene, was shown to be an activator of flowering in rice. It was previously demonstrated that ''CO'' controls the flowering time by integrating signals from the circadian clock and light to regulate ''FT'' expression in Arabidopsis<ref name="ref5" />.Unlike Arabidopsis ''FT'', expression of ''Hd3a'' mRNA is more or less opposite to that of ''Hd1'' under inductive short-day conditions<ref name="ref6" />.<br />
<br />
==References==<br />
<references><br />
<ref name="ref1">Song Lim Kim, Shinyoung Lee, Hyo Jung Kim, Hong Gil Nam, Gynheung An（2007）''OsMADS51'' is a short-day flowering promoter that functions upstream of ''Ehd1'', ''OsMADS14'', and ''Hd3a''.''Plant Physiology'' 145(4): 1484-1494.</ref><br />
<ref name="ref2">Hironori Itoh, Takeshi Izawa (2011) A study of phytohormone biosynthetic gene expression using a circadian clock-related mutant in rice. ''Plant Signaling & Behavior'' 6(12): 1932-1936.</ref> <br />
<ref name="ref3">Ryousuke Hayama, Takeshi Izawa, Ko Shimamoto (2002) Isolation of rice genes possibly involved in the photoperiodic control of flowering by a<br />
fluorescent differential display method. ''Plant and Cell Physiology'' 43(5): 494-504.</ref> <br />
<ref name="ref4">Takeshi Izawa, Motohiro Mihara, Yuji Suzuki, Meenu Gupta, Hironori Itoh, et al. (2011) ''Os-GIGANTEA'' confers robust diurnal rhythms on the global transcriptome of rice in the field. ''The Plant Cell'' 23(5): 1741-1755.</ref> <br />
<ref name="ref5">Ryosuke Hayama, Shuji Yokoi, Shojiro Tamaki, Masahiro Yano, Ko Shimamoto (2003) Adaptation of photoperiodic control pathways produces short-day flowering in rice. ''Nature'' 422(6933): 719-722.</ref> <br />
<ref name="ref6">Bong-Soo Shin, Jeong-Hyun Lee, Jeong-Hwan Lee, Hyun-Joo Jeong, Choong-Hyo Yun, Jeong-Kook Kim (2004) Circadian regulation of rice (''Oryza sativa L.'') ''CONSTANS-like'' gene transcripts. ''Molecules and Cells'' 17(1): 10-16.</ref><br />
<br />
==Structured Information==<br />
{{JaponicaGene|<br />
GeneName = Os01g0182600|<br />
Description = GIGANTEA protein|<br />
Version = NM_001048755.1 GI:115434923 GeneID:4325329|<br />
Length = 9083 bp|<br />
Definition = Oryza sativa Japonica Group Os01g0182600, complete gene.|<br />
Source = Oryza sativa Japonica Group<br />
<br />
ORGANISM Oryza sativa Japonica Group<br />
Eukaryota; Viridiplantae; Streptophyta; Embryophyta; Tracheophyta;<br />
Spermatophyta; Magnoliophyta; Liliopsida; Poales; Poaceae; BEP<br />
clade; Ehrhartoideae; Oryzeae; Oryza.<br />
|<br />
Chromosome = [[:category:Japonica Chromosome 1|Chromosome 1]]|<br />
AP = Chromosome 1:4328362..4337444|<br />
CDS = 4328725..4329030,4329123..4329215,4329333..4329551,4329752..4329820,4329925..4331421<br>,4332220..4332429,4332563..4332713,4332791..4333031,4333898..4333956<br>,4334379..4334653,4334813..4334955,4335034..4335088,4335181..4335258<br>,4335351..4335437|<br />
GCID = <gbrowseImage1><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage1>|<br />
GSID = <gbrowseImage2><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage2>|<br />
CDNA = <cdnaseq>atgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactgcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcatagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcggcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctga</cdnaseq>|<br />
AA = <aaseq>MSASNEKWIDGLQFSSLFWPPPQDSQQKQAQILAYVEYFGQFTA DSEQFPEDIAQLIQSCYPSKEKRLVDEVLATFVLHHPEHGHAVVHPILSRIIDGTLSY DRNGFPFMSFISLFSHTSEKEYSEQWALACGEILRVLTHYNRPIFKVDHQHSEAECSS TSDQASSCESMEKRANGSPRNEPDRKPLRPLSPWITDILLAAPLGIRSDYFRWCGGVM GKYAAGGELKPPTTAYSRGSGKHPQLMPSTPRWAVANGAGVILSVCDEEVARYETANL TAAAVPALLLPPPTTPLDEHLVAGLPPLEPYARLFHRYYAIATPSATQRLLFGLLEAP PSWAPDALDAAVQLVELLRAAEDYDSGMRLPKNWMHLHFLRAIGTAMSMRAGIAADTS AALLFRILSQPTLLFPPLRHAEGVELHHEPLGGYVSSYKRQLEVPASEATIDATAQGI ASMLCAHGPDVEWRICTIWEAAYGLLPLSSSAVDLPEIVVAAPLQPPTLSWSLYLPLL KVFEYLPRGSPSEACLMRIFVATVEAILRRTFPSETSEQSRKPRSQSKNLAVAELRTM IHSLFVESCASMDLASRLLFVVLTVCVSHQALPGGSKRPTGSDNHSSEEVTNDSRLTN GRNRCKKRQGPVATFDSYVLAAVCALSCELQLFPFISKNGNHSNLKDSIKIVIPGKTT GISNELHNSISSAILHTRRILGILEALFSLKPSSVGTSWSYSSNEIVAAAMVAAHVSE LFRRSRPCLNALSALKQCKWDAEISTRASSLYHLIDLHGKTVTSIVNKAEPLEAHLTL TPVKKDEPPIEEKNINSSDGGALEKKDASRSHRKNGFARPLLKCAEDVILNGDVASTS GKAIASLQVEASDLANFLTMDRNGGYRGSQTLLRSVLSEKQELCFSVVSLLWQKLIAS PEMQMSAESTSAHQGWRKVVDALCDIVSASPTKASAAIVLQAEKDLQPWIARDDEQGQ KMWRVNQRIVKLIAELMRNHDSPEALVILASASDLLLRATDGMLVDGEACTLPQLELL EVTARAVHLIVEWGDSGVSVADGLSNLLKCRLSTTIRCLSHPSAHVRALSMSVLRDIL NSGQINSSKLIQGEHRNGIQSPTYQCLAASIINWQADVERCIEWEAHSRRATGLTLAF LTAAAKELGCPLTC</aaseq>|<br />
DNA = <dnaseqindica>8415..8720#8230..8322#7894..8112#7625..7693#6024..7520#5016..5225#4732..4882#4414..4654#3489..3547#2792..3066#2490..2632#2357..2411#2187..2264#2008..2094#ccgagccgtttctctccctctctctctcctcctccttcccctgcgcctcctccaccgcccgtgtcgccgccgcgcgcgtctatcccctcgccgccgccgccgtctacgccgcgtgaggtccggattggttgaaaggctgatcgaggtaatgcttttttatttttcccgtgcggttttagtttttctgtgctgtgcttggtggttttttctttaaaactatttttaggctggttttgatgatggcctttctaaggttgtcgtgtatggggggcgaggtggttcgggtggatttgtggtgttatgttgaggggatttttgtttggggggagagagttttggtggtagatgcggttgggattttgaagacgtgaaagggaaattagttttgaaatttgacggcgaggttgagctgattagtattgtaataattcgtggcatcggatagatcaacgcgagatggatgcttattactgcgaatttcaggaggtttttgtggtttttttaggcgagtagcgttgattctagctggattttgctgcttcgtggaggcgtttagatgtggtgcatacggctgtggatgtaatttgtgcggggataattactgaaaattaattcgcagtaagctagctctgctttaatattttttcccctgaaacagtaacaagggatcccttcctcatttcgttaacagtggaagtgaatacagacaccaaccgatgagcagagacacctgtaattaaacaccaaaagaagaacctgttttgttcctatatattttccatttctatgctttaggtaatatactttctggttcggtgtcatagtacccgaaatagaatctttgcttgccaaaaaaagaaagaataacaagatagaacaacatcattaaattctgtcttagacaatacatttataatcgaccaacaaaaacagaccgatgagtgacgtggaataacaaatgaagtataaatgtcacgaaaaacacatctattgattttgtgtgataagccataggagcctacgaggaaccaagtgaatggttctgtacaccacatccattagcattaccataaatttgcctatttctatgatgccttccttaacatatttatcgacatgtgagcttgcaccataactaaatgttatggtaaactttaccttataaatgattgcaacatgtaccttttcatttcgataattaaaaacgagaaagcttccacctttttcctagatatattttttcgtttgattatgaatttagttgtacttgacaataaaataatgcgcttgtaagattatgcaattttcgttgcattacctccctctcttttcatttgtcagtttttttctactccctccgtttcatattttaagtcgtttgacttttttcctagtcagactttgttaagtttgacaaagtttatagtaaaatttagcaatatctaaaatattaaattggtttcatcaaatctaacattgaatatattttgataatatgttttttttgtgttgaaaatacttctatatttttctataaacttagtcaaacttggaagaaatttcagtaagaaaaaagtcaaacggcttataatatgaaacggagggagtaaatttttttttgcattaattaaaattcatgtattttcctccttttttggtttttgttgtgcgatatattacagtgtagagactctgaagctcatatggttggcattggttgatatgataacgtgtcacgttgtgagatgcagacccccgttatgggggttgattgaatgaagtggatagcactcttataactttcttgctcgaagtttcttgataccaaggcaagtagcttataagaagatttggactttggtggcataaatggatgaactaactggcggcttcaacgtggtgttgattacaggtgcttttattgctaactggagcagtttgtgtagtttctttctttctctttgagaattctctcagtttcttgggcaggctcttgagctaccaagtatgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggtaacattattcctatttgattcttgttagaaattctatatgttgaacatacttgcctcaatgggaaacttggttgcgtctcctctctgaaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcaggtactgtatctctaagcatgtttagtatcttctgtgataaaatctttctccagtgtgttaacttttgcattctttttcttgcatgcttgcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattaggtaatttgatccagtgatcaatacatgttccacaaacaattgaagaaagagtttgacatatttttctcatatttccagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgaggtaaggcccctactctaatggttagttccaatttattggttcatttctctgtgtttgataagattttgcctctgctgtgcatttcaatattatatctaaactgtgctgcttatagcttatcaaactactcttgctcataaagatgcaacttttatgcagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgagtcataataaatgataacatattattgttctgtggactgaaaatctgaaatcctcctttcttgagaaatacaacctcaattttatggtgatattgacaagaaggaagagcgcatataatttaaagttaaaacacggaaaataatagaagtatgtgagcagaacgcaggagcaaggggtggaactcataacaagttcaaacttcatgcatgttcataattccccaatgcaagtatggcaagaaaaatagacaacttttaaaagtatgctaatggaactgcatgcattgtgtaaacttgcttttgtcaagcatgttggaacatactatatatttgctgctgatgatgaaaaatttgcttgagtactcttctattgctaatggacgcttaccattccctgttttgtcttcaaatgcatcaggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactggttggtaaaatcagcaatggaattttctatcgtgaattttttataattattgtaacattaattactcccatgtctttttattctgttgagataccatgtgacaatgtctacaattgcatattttgcattctttggtagatcagttcccttcttaagatccccaaagggaaactggagtatcaataacttactccctcaaataaaaaaaaagtgatactttaggcataaacacattcttcaaaacaaaactttgaatagcagtttcttttgtaatatattaataaagcatggaaaaattataatgtgaagaatgtacattcagggacaaatatactcatcatttttgcatggttgatcttaatgtccatagagagattgatcagaccaaaagtttatttttgacaggagtgagtatggattataataggagtgaatggcaatatgctgtgaaagtgagcattttatatttgtttactactagcatttggtgacaaagcaaatgatgtatattgtaaatcaaagtaaacaagtagaacatgtcctgtggcgacaagacataacctacaaattaaagttagcctgattgtttattttactgggagcacattttttctagattctagagtacatattattaaaaaaggtgtgatatgtttgcaaattactttttaatggaagtttgtaaattactttgaaacttgcattcagtttttttgcttcttaaaaaacatagtgcatatgggctttatgagctcatactatgcatgtttttctgcattggagtgggacctttatctttttcttgcaactcggatgctaatgcttattataaacacaagttactaacatgtgcttgggtcattcagcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcataggtaagctcttatgatcctgtccactgttgtaccatgtgtactttccaaatgattgctaacatgctatttttcgtcagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtac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/>
Link = [http://www.ncbi.nlm.nih.gov/nuccore/NM_001048755.1 RefSeq:Os01g0182600]|<br />
}}<br />
[[Category:Genes]]<br />
[[Category:Japonica mRNA]]<br />
[[Category:Oryza Sativa Japonica Group]]<br />
[[Category:Japonica Genes]]<br />
[[Category:Japonica Chromosome 1]]<br />
[[Category:Chromosome 1]]</div>Mikahttps://ngdc.cncb.ac.cn/ricewiki/index.php?title=Os01g0182600&diff=183828Os01g01826002014-06-11T03:20:44Z<p>Mika: /* Function */</p>
<hr />
<div>The rice gene of ''Gigantea(GI)'' is solo ortholog of ''GI'' in the Arabidopsis circadian system and related to photoperiodic control of flowering.<br />
<br />
==Annotated Information==<br />
===Function===<br />
*''OsGI'',a ''GI'' ortholog, regulates flowering time in response to photoperiodic conditions. Mutations in this gene delay flowering in inductive SDs (short-days),whereas flowering is marginally induced under LDs(long-days).In comparison, over expression of ''OsGI'' in transgenic rice causes late flowering in both SDs and LDs.In addition, ''OsGI'' has function to regulate ''Hd1'', a gene that also shows circadian rhythm<ref name="ref1" />. ''Os-GI'' affects gene expression of approximately 75% of genes among 27,201 genes and is required for strong amplitudes and proper phase-setting of global gene expression under natural field conditions<ref name="ref2" />. <br />
*The cDNA of ''OsGI'' firstly found in ''se5'' mutant that is completely photoperiod-insensitive, the expression patterns of rice and Arabidopsis ''GI'' gene are both photoperiodically regulated and are very similar under LD and SD conditions(FIG1)<ref name="ref3" />. ''GI'' is required for phase-dependent ''CO''transcription through the interaction with FKF1, which leads to cycling DNA binding with one finger (CDF) degradation and consequent derepression of CO. In addition, it has been known that GI protein interacts with ZEITLUPE (ZTL) F-box protein to control the diurnal rhythm of TOC1 protein in regulation of the Arabidopsis circadian clock<ref name="ref4" />. <br />
*''Os-GI'' also has function to maintain bioactive GA level through the regulation of the GA-deactivating enzyme genes in rice, ''Os-GI'' represses major ''OsGA2ox'' genes in order to maintain a proper bioactive GA level, ''osgi-1'' plants showed a semi-dwarf phenotype with elevated ''OsGA2ox'' gene expression <ref name="ref4" /> .[[File:Limeijia1.png|frame|'''Figure 2''' WT rice and osgi-1plants grown under natural field conditions and osgi-1plant exhibits Semi-dwarf phenotype.]]<br />
<br />
===Expression===<br />
*''GI'' encodes a nuclear protein that defines the proper amplitude and period length of circadian rhythms. ''GI'' expression is regulated by the circadian clock, with a peak in transcript levels at 8 to 10 h after dawn. The ''GI'' protein acts between the circadian oscillator and ''CO'' to promote flowering by increasing ''CO'' and ''FT'' mRNA abundance<ref name="ref1" />. The expression of the ''OsGI'' messenger RNA was circadian-controlled and that its temporal expression pattern was very similar to that of ''GI'' under both SD and LD conditions. Decreased ''OsGI'' expression led to early flowering under LD conditions and late flowering under SD conditions relative to the wild type, and that this early flowering phenotype under LD conditions was the opposite of that of the Arabidopsis ''gi'' mutants, confirming the opposite role of the ''GI'' gene in flowering under LD conditions in rice and Arabidopsis<ref name="ref5" />.<br />
<br />
===Mutation===<br />
*The single ''osgi-1'' mutation extended the time to flowering only under SD conditions. Under LD conditions the single ''hd1'' mutation induced early flowering, but ''osgi-1'' plants exhibited no phenotypic change<ref name="ref4" /><br />
<br />
===Evolution===<br />
* There is some evolutionary conservation between rice circadian clock–related genes and those associated with Arabidopsis circadian clocks, where the circadian clock consists of interlocked subloops, it is also possible that ''Os-GI'' is a member of a subloop in rice circadian clocks<ref name="ref4" />.But there is a divergence in the photoperiodic pathways of these two species arises from the molecular mechanisms controlling the expression of ''FT/Hd3a'' by ''CO/Hd1(Se1)''<ref name="ref5" />(figure 3)<br />
<br />
==Labs working on this gene==<br />
*Photosynthesis and Photobiology Research Unit, National Institute of Agrobiological Sciences, 2-1-2 Kannondai, Tsukuba 305-8602, Japan<br />
*Laboratory of Plant Molecular Genetics, Nara Institute of Science and Technology, 8916-5 Takayama, Ikoma 630-0101, Japan<br />
*Plant Genomics Research Unit, National Institute of Agrobiological Sciences, 2-1-2 Kannondai, Tsukuba 305-8602, Japan<br />
<br />
==Knowledge Extension==<br />
*The rice orthologues of the two key regulators of flowering time in Arabidopsis, ''CO'' and ''FT'', were recently shown to be important in the photoperiodic control of flowering in rice. The rice ''Hd1(Se1)''gene, which is an orthologue of the Arabidopsis ''CO'' gene, was required for the suppression of flowering under LD conditions and for the promotion of flowering under SD conditions. Furthermore, the rice ''Hd3a'' gene, an orthologue of the Arabidopsis ''FT'' gene, was shown to be an activator of flowering in rice. It was previously demonstrated that ''CO'' controls the flowering time by integrating signals from the circadian clock and light to regulate ''FT'' expression in Arabidopsis<ref name="ref5" />.Unlike Arabidopsis ''FT'', expression of ''Hd3a'' mRNA is more or less opposite to that of ''Hd1'' under inductive short-day conditions<ref name="ref6" />.<br />
<br />
==References==<br />
<references><br />
<ref name="ref1">Song Lim Kim, Shinyoung Lee, Hyo Jung Kim, Hong Gil Nam, Gynheung An（2007）''OsMADS51'' is a short-day flowering promoter that functions upstream of ''Ehd1'', ''OsMADS14'', and ''Hd3a''.''Plant Physiology'' 145(4): 1484-1494.</ref><br />
<ref name="ref2">Hironori Itoh, Takeshi Izawa (2011) A study of phytohormone biosynthetic gene expression using a circadian clock-related mutant in rice. ''Plant Signaling & Behavior'' 6(12): 1932-1936.</ref> <br />
<ref name="ref3">Ryousuke Hayama, Takeshi Izawa, Ko Shimamoto (2002) Isolation of rice genes possibly involved in the photoperiodic control of flowering by a<br />
fluorescent differential display method. ''Plant and Cell Physiology'' 43(5): 494-504.</ref> <br />
<ref name="ref4">Takeshi Izawa, Motohiro Mihara, Yuji Suzuki, Meenu Gupta, Hironori Itoh, et al. (2011) ''Os-GIGANTEA'' confers robust diurnal rhythms on the global transcriptome of rice in the field. ''The Plant Cell'' 23(5): 1741-1755.</ref> <br />
<ref name="ref5">Ryosuke Hayama, Shuji Yokoi, Shojiro Tamaki, Masahiro Yano, Ko Shimamoto (2003) Adaptation of photoperiodic control pathways produces short-day flowering in rice. ''Nature'' 422(6933): 719-722.</ref> <br />
<ref name="ref6">Bong-Soo Shin, Jeong-Hyun Lee, Jeong-Hwan Lee, Hyun-Joo Jeong, Choong-Hyo Yun, Jeong-Kook Kim (2004) Circadian regulation of rice (''Oryza sativa L.'') ''CONSTANS-like'' gene transcripts. ''Molecules and Cells'' 17(1): 10-16.</ref><br />
<br />
==Structured Information==<br />
{{JaponicaGene|<br />
GeneName = Os01g0182600|<br />
Description = GIGANTEA protein|<br />
Version = NM_001048755.1 GI:115434923 GeneID:4325329|<br />
Length = 9083 bp|<br />
Definition = Oryza sativa Japonica Group Os01g0182600, complete gene.|<br />
Source = Oryza sativa Japonica Group<br />
<br />
ORGANISM Oryza sativa Japonica Group<br />
Eukaryota; Viridiplantae; Streptophyta; Embryophyta; Tracheophyta;<br />
Spermatophyta; Magnoliophyta; Liliopsida; Poales; Poaceae; BEP<br />
clade; Ehrhartoideae; Oryzeae; Oryza.<br />
|<br />
Chromosome = [[:category:Japonica Chromosome 1|Chromosome 1]]|<br />
AP = Chromosome 1:4328362..4337444|<br />
CDS = 4328725..4329030,4329123..4329215,4329333..4329551,4329752..4329820,4329925..4331421<br>,4332220..4332429,4332563..4332713,4332791..4333031,4333898..4333956<br>,4334379..4334653,4334813..4334955,4335034..4335088,4335181..4335258<br>,4335351..4335437|<br />
GCID = <gbrowseImage1><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage1>|<br />
GSID = <gbrowseImage2><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage2>|<br />
CDNA = <cdnaseq>atgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactgcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcatagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcggcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctga</cdnaseq>|<br />
AA = <aaseq>MSASNEKWIDGLQFSSLFWPPPQDSQQKQAQILAYVEYFGQFTA DSEQFPEDIAQLIQSCYPSKEKRLVDEVLATFVLHHPEHGHAVVHPILSRIIDGTLSY DRNGFPFMSFISLFSHTSEKEYSEQWALACGEILRVLTHYNRPIFKVDHQHSEAECSS TSDQASSCESMEKRANGSPRNEPDRKPLRPLSPWITDILLAAPLGIRSDYFRWCGGVM GKYAAGGELKPPTTAYSRGSGKHPQLMPSTPRWAVANGAGVILSVCDEEVARYETANL TAAAVPALLLPPPTTPLDEHLVAGLPPLEPYARLFHRYYAIATPSATQRLLFGLLEAP PSWAPDALDAAVQLVELLRAAEDYDSGMRLPKNWMHLHFLRAIGTAMSMRAGIAADTS AALLFRILSQPTLLFPPLRHAEGVELHHEPLGGYVSSYKRQLEVPASEATIDATAQGI ASMLCAHGPDVEWRICTIWEAAYGLLPLSSSAVDLPEIVVAAPLQPPTLSWSLYLPLL KVFEYLPRGSPSEACLMRIFVATVEAILRRTFPSETSEQSRKPRSQSKNLAVAELRTM IHSLFVESCASMDLASRLLFVVLTVCVSHQALPGGSKRPTGSDNHSSEEVTNDSRLTN GRNRCKKRQGPVATFDSYVLAAVCALSCELQLFPFISKNGNHSNLKDSIKIVIPGKTT GISNELHNSISSAILHTRRILGILEALFSLKPSSVGTSWSYSSNEIVAAAMVAAHVSE LFRRSRPCLNALSALKQCKWDAEISTRASSLYHLIDLHGKTVTSIVNKAEPLEAHLTL TPVKKDEPPIEEKNINSSDGGALEKKDASRSHRKNGFARPLLKCAEDVILNGDVASTS GKAIASLQVEASDLANFLTMDRNGGYRGSQTLLRSVLSEKQELCFSVVSLLWQKLIAS PEMQMSAESTSAHQGWRKVVDALCDIVSASPTKASAAIVLQAEKDLQPWIARDDEQGQ KMWRVNQRIVKLIAELMRNHDSPEALVILASASDLLLRATDGMLVDGEACTLPQLELL EVTARAVHLIVEWGDSGVSVADGLSNLLKCRLSTTIRCLSHPSAHVRALSMSVLRDIL NSGQINSSKLIQGEHRNGIQSPTYQCLAASIINWQADVERCIEWEAHSRRATGLTLAF LTAAAKELGCPLTC</aaseq>|<br />
DNA = <dnaseqindica>8415..8720#8230..8322#7894..8112#7625..7693#6024..7520#5016..5225#4732..4882#4414..4654#3489..3547#2792..3066#2490..2632#2357..2411#2187..2264#2008..2094#ccgagccgtttctctccctctctctctcctcctccttcccctgcgcctcctccaccgcccgtgtcgccgccgcgcgcgtctatcccctcgccgccgccgccgtctacgccgcgtgaggtccggattggttgaaaggctgatcgaggtaatgcttttttatttttcccgtgcggttttagtttttctgtgctgtgcttggtggttttttctttaaaactatttttaggctggttttgatgatggcctttctaaggttgtcgtgtatggggggcgaggtggttcgggtggatttgtggtgttatgttgaggggatttttgtttggggggagagagttttggtggtagatgcggttgggattttgaagacgtgaaagggaaattagttttgaaatttgacggcgaggttgagctgattagtattgtaataattcgtggcatcggatagatcaacgcgagatggatgcttattactgcgaatttcaggaggtttttgtggtttttttaggcgagtagcgttgattctagctggattttgctgcttcgtggaggcgtttagatgtggtgcatacggctgtggatgtaatttgtgcggggataattactgaaaattaattcgcagtaagctagctctgctttaatattttttcccctgaaacagtaacaagggatcccttcctcatttcgttaacagtggaagtgaatacagacaccaaccgatgagcagagacacctgtaattaaacaccaaaagaagaacctgttttgttcctatatattttccatttctatgctttaggtaatatactttctggttcggtgtcatagtacccgaaatagaatctttgcttgccaaaaaaagaaagaataacaagatagaacaacatcattaaattctgtcttagacaatacatttataatcgaccaacaaaaacagaccgatgagtgacgtggaataacaaatgaagtataaatgtcacgaaaaacacatctattgattttgtgtgataagccataggagcctacgaggaaccaagtgaatggttctgtacaccacatccattagcattaccataaatttgcctatttctatgatgccttccttaacatatttatcgacatgtgagcttgcaccataactaaatgttatggtaaactttaccttataaatgattgcaacatgtaccttttcatttcgataattaaaaacgagaaagcttccacctttttcctagatatattttttcgtttgattatgaatttagttgtacttgacaataaaataatgcgcttgtaagattatgcaattttcgttgcattacctccctctcttttcatttgtcagtttttttctactccctccgtttcatattttaagtcgtttgacttttttcctagtcagactttgttaagtttgacaaagtttatagtaaaatttagcaatatctaaaatattaaattggtttcatcaaatctaacattgaatatattttgataatatgttttttttgtgttgaaaatacttctatatttttctataaacttagtcaaacttggaagaaatttcagtaagaaaaaagtcaaacggcttataatatgaaacggagggagtaaatttttttttgcattaattaaaattcatgtattttcctccttttttggtttttgttgtgcgatatattacagtgtagagactctgaagctcatatggttggcattggttgatatgataacgtgtcacgttgtgagatgcagacccccgttatgggggttgattgaatgaagtggatagcactcttataactttcttgctcgaagtttcttgataccaaggcaagtagcttataagaagatttggactttggtggcataaatggatgaactaactggcggcttcaacgtggtgttgattacaggtgcttttattgctaactggagcagtttgtgtagtttctttctttctctttgagaattctctcagtttcttgggcaggctcttgagctaccaagtatgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggtaacattattcctatttgattcttgttagaaattctatatgttgaacatacttgcctcaatgggaaacttggttgcgtctcctctctgaaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcaggtactgtatctctaagcatgtttagtatcttctgtgataaaatctttctccagtgtgttaacttttgcattctttttcttgcatgcttgcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattaggtaatttgatccagtgatcaatacatgttccacaaacaattgaagaaagagtttgacatatttttctcatatttccagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgaggtaaggcccctactctaatggttagttccaatttattggttcatttctctgtgtttgataagattttgcctctgctgtgcatttcaatattatatctaaactgtgctgcttatagcttatcaaactactcttgctcataaagatgcaacttttatgcagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgagtcataataaatgataacatattattgttctgtggactgaaaatctgaaatcctcctttcttgagaaatacaacctcaattttatggtgatattgacaagaaggaagagcgcatataatttaaagttaaaacacggaaaataatagaagtatgtgagcagaacgcaggagcaaggggtggaactcataacaagttcaaacttcatgcatgttcataattccccaatgcaagtatggcaagaaaaatagacaacttttaaaagtatgctaatggaactgcatgcattgtgtaaacttgcttttgtcaagcatgttggaacatactatatatttgctgctgatgatgaaaaatttgcttgagtactcttctattgctaatggacgcttaccattccctgttttgtcttcaaatgcatcaggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactggttggtaaaatcagcaatggaattttctatcgtgaattttttataattattgtaacattaattactcccatgtctttttattctgttgagataccatgtgacaatgtctacaattgcatattttgcattctttggtagatcagttcccttcttaagatccccaaagggaaactggagtatcaataacttactccctcaaataaaaaaaaagtgatactttaggcataaacacattcttcaaaacaaaactttgaatagcagtttcttttgtaatatattaataaagcatggaaaaattataatgtgaagaatgtacattcagggacaaatatactcatcatttttgcatggttgatcttaatgtccatagagagattgatcagaccaaaagtttatttttgacaggagtgagtatggattataataggagtgaatggcaatatgctgtgaaagtgagcattttatatttgtttactactagcatttggtgacaaagcaaatgatgtatattgtaaatcaaagtaaacaagtagaacatgtcctgtggcgacaagacataacctacaaattaaagttagcctgattgtttattttactgggagcacattttttctagattctagagtacatattattaaaaaaggtgtgatatgtttgcaaattactttttaatggaagtttgtaaattactttgaaacttgcattcagtttttttgcttcttaaaaaacatagtgcatatgggctttatgagctcatactatgcatgtttttctgcattggagtgggacctttatctttttcttgcaactcggatgctaatgcttattataaacacaagttactaacatgtgcttgggtcattcagcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcataggtaagctcttatgatcctgtccactgttgtaccatgtgtactttccaaatgattgctaacatgctatttttcgtcagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcgggtatatcatacatctctctggctgttgtttcagtgctatcaatcaagatgacacatcatgcaatattatcataacagcacattatgtgttttgaatattttattgattgtcatagctttgttttatgttttagcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcaggtatgcagtgtttcctgcacctacttgctacctattctgtatttgatgtattcaatttcctttttatcagttttctcgaaatatatgctaataggcttgtcactaaacggctacattatccagactctgttctgaatttttttttcattaagtactgttagctgccagattacatcatcatgactaaagtagtttagctacaatattgcaaagctagatttgtctagcctctattgttttgaatagaaaaggcaggtatgcagtgtttcctgcacctacttgctacctattctgtatttgatgtattcaatttcctttttatcagttttctcgaaatatatgctaataggcttgtcactaaacggctacattatccagactctgttctgaatttttttttcattaagtactgttagctgccagattacatcatcatgactaaagtagtttagctacaatattgcaaagctagatttgtctagcctctattgttttgaatagaaaattaaatcttttggtcatttttttgtaaccttttaggagtctctgtatgcttgtcgtcaacatgctatgataagtttatatcagttcttttggagctatgatgtattcttgtatattttatgactgctaggatattatgttaaaactgttttaaaggtatcatcaaacaagttaatatagatattgctacctaccaaagttggaatttgattatgtcgtaaaatatatgcaccctacatgttttagcttcatattgacttatattttcatatgtgctccttcaactgcacaaagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggtatggtcattcctgcttataagataccagaataatgctatgaagaccgtccttggttgtcagcattcttatacacaatgataatctttgtttttcatgggtaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggtcaagatattgttattgttatatttacatacttgttatcatatttaaaagcttcatattgtatgtacctttgaaaaaaaaagttccatactgtagcttcttttgttctcatttcttggtttcaaagcatcagcatcattaccgatataaaatatacccatttcctcagtcatcttccccgactcttgtatctttcacaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagaggtaacaccttcattcatgaggtccttaggtgcaaatcccagtgaagaataatcagttctactttgtatggttcatctcgcttactgaccggttcatctcgggatcaattacgagcagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaaggcatgtcctcttctctgatctcactcacgttcatgccaaacttactcatcatcatccatgatccatcatcctgactgaatcttgatatgcagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctgacaaggccacaccatgactgtcactgcaagcagctgctgtaggatcagcgagtaggaaaggatgacttgccggccagtttcctgagatgtgatttaagactgatattagctgatgttcccaagcatgatacggggcttgctcccaagatgtgatttttactttacttttggttaatgtgctaccgctgacattagatgttcaagcatatgaaaactgcttgtgctgtaacttgtatgtctgtaacgtaaaaatctttccatatggtaaccggggaagcagaggtatatccttttctgctgttgtatctatttccagtttcctgctgaacttgcaaatgaatattcagcaatttccgttttgctc</dnaseqindica>|<br />
Link = [http://www.ncbi.nlm.nih.gov/nuccore/NM_001048755.1 RefSeq:Os01g0182600]|<br />
}}<br />
[[Category:Genes]]<br />
[[Category:Japonica mRNA]]<br />
[[Category:Oryza Sativa Japonica Group]]<br />
[[Category:Japonica Genes]]<br />
[[Category:Japonica Chromosome 1]]<br />
[[Category:Chromosome 1]]</div>Mikahttps://ngdc.cncb.ac.cn/ricewiki/index.php?title=File:Limeijia1.png&diff=183827File:Limeijia1.png2014-06-11T03:16:43Z<p>Mika: WT rice and osgi-1plants grown under natural field conditions and osgi-1plant exhibits Semi-dwarf phenotype</p>
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<div>WT rice and osgi-1plants grown under natural field conditions and osgi-1plant exhibits Semi-dwarf phenotype</div>Mikahttps://ngdc.cncb.ac.cn/ricewiki/index.php?title=Os01g0182600&diff=183823Os01g01826002014-06-11T03:11:37Z<p>Mika: /* Function */</p>
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<div>The rice gene of ''Gigantea(GI)'' is solo ortholog of ''GI'' in the Arabidopsis circadian system and related to photoperiodic control of flowering.<br />
<br />
==Annotated Information==<br />
===Function===<br />
*''OsGI'',a ''GI'' ortholog, regulates flowering time in response to photoperiodic conditions. Mutations in this gene delay flowering in inductive SDs (short-days),whereas flowering is marginally induced under LDs(long-days).In comparison, over expression of ''OsGI'' in transgenic rice causes late flowering in both SDs and LDs.In addition, ''OsGI'' has function to regulate ''Hd1'', a gene that also shows circadian rhythm<ref name="ref1" />. ''Os-GI'' affects gene expression of approximately 75% of genes among 27,201 genes and is required for strong amplitudes and proper phase-setting of global gene expression under natural field conditions<ref name="ref2" />. <br />
*The cDNA of ''OsGI'' firstly found in ''se5'' mutant that is completely photoperiod-insensitive, the expression patterns of rice and Arabidopsis ''GI'' gene are both photoperiodically regulated and are very similar under LD and SD conditions(FIG1)<ref name="ref3" />. ''GI'' is required for phase-dependent ''CO''transcription through the interaction with FKF1, which leads to cycling DNA binding with one finger (CDF) degradation and consequent derepression of CO. In addition, it has been known that GI protein interacts with ZEITLUPE (ZTL) F-box protein to control the diurnal rhythm of TOC1 protein in regulation of the Arabidopsis circadian clock<ref name="ref4" />. <br />
*''Os-GI'' also has function to maintain bioactive GA level through the regulation of the GA-deactivating enzyme genes in rice, ''Os-GI'' represses major ''OsGA2ox'' genes in order to maintain a proper bioactive GA level, ''osgi-1'' plants showed a semi-dwarf phenotype with elevated ''OsGA2ox'' gene expression <ref name="ref4" /> .[[File:Limeijia1.png|frame|'''Figure 1'''WT rice and osgi-1plants grown under natural field conditions and osgi-1plant exhibits Semi-dwarf phenotype.]]<br />
<br />
===Expression===<br />
*''GI'' encodes a nuclear protein that defines the proper amplitude and period length of circadian rhythms. ''GI'' expression is regulated by the circadian clock, with a peak in transcript levels at 8 to 10 h after dawn. The ''GI'' protein acts between the circadian oscillator and ''CO'' to promote flowering by increasing ''CO'' and ''FT'' mRNA abundance<ref name="ref1" />. The expression of the ''OsGI'' messenger RNA was circadian-controlled and that its temporal expression pattern was very similar to that of ''GI'' under both SD and LD conditions. Decreased ''OsGI'' expression led to early flowering under LD conditions and late flowering under SD conditions relative to the wild type, and that this early flowering phenotype under LD conditions was the opposite of that of the Arabidopsis ''gi'' mutants, confirming the opposite role of the ''GI'' gene in flowering under LD conditions in rice and Arabidopsis<ref name="ref5" />.<br />
<br />
===Mutation===<br />
*The single ''osgi-1'' mutation extended the time to flowering only under SD conditions. Under LD conditions the single ''hd1'' mutation induced early flowering, but ''osgi-1'' plants exhibited no phenotypic change<ref name="ref4" /><br />
<br />
===Evolution===<br />
* There is some evolutionary conservation between rice circadian clock–related genes and those associated with Arabidopsis circadian clocks, where the circadian clock consists of interlocked subloops, it is also possible that ''Os-GI'' is a member of a subloop in rice circadian clocks<ref name="ref4" />.But there is a divergence in the photoperiodic pathways of these two species arises from the molecular mechanisms controlling the expression of ''FT/Hd3a'' by ''CO/Hd1(Se1)''<ref name="ref5" />(figure 3)<br />
<br />
==Labs working on this gene==<br />
*Photosynthesis and Photobiology Research Unit, National Institute of Agrobiological Sciences, 2-1-2 Kannondai, Tsukuba 305-8602, Japan<br />
*Laboratory of Plant Molecular Genetics, Nara Institute of Science and Technology, 8916-5 Takayama, Ikoma 630-0101, Japan<br />
*Plant Genomics Research Unit, National Institute of Agrobiological Sciences, 2-1-2 Kannondai, Tsukuba 305-8602, Japan<br />
<br />
==Knowledge Extension==<br />
*The rice orthologues of the two key regulators of flowering time in Arabidopsis, ''CO'' and ''FT'', were recently shown to be important in the photoperiodic control of flowering in rice. The rice ''Hd1(Se1)''gene, which is an orthologue of the Arabidopsis ''CO'' gene, was required for the suppression of flowering under LD conditions and for the promotion of flowering under SD conditions. Furthermore, the rice ''Hd3a'' gene, an orthologue of the Arabidopsis ''FT'' gene, was shown to be an activator of flowering in rice. It was previously demonstrated that ''CO'' controls the flowering time by integrating signals from the circadian clock and light to regulate ''FT'' expression in Arabidopsis<ref name="ref5" />.Unlike Arabidopsis ''FT'', expression of ''Hd3a'' mRNA is more or less opposite to that of ''Hd1'' under inductive short-day conditions<ref name="ref6" />.<br />
<br />
==References==<br />
<references><br />
<ref name="ref1">Song Lim Kim, Shinyoung Lee, Hyo Jung Kim, Hong Gil Nam, Gynheung An（2007）''OsMADS51'' is a short-day flowering promoter that functions upstream of ''Ehd1'', ''OsMADS14'', and ''Hd3a''.''Plant Physiology'' 145(4): 1484-1494.</ref><br />
<ref name="ref2">Hironori Itoh, Takeshi Izawa (2011) A study of phytohormone biosynthetic gene expression using a circadian clock-related mutant in rice. ''Plant Signaling & Behavior'' 6(12): 1932-1936.</ref> <br />
<ref name="ref3">Ryousuke Hayama, Takeshi Izawa, Ko Shimamoto (2002) Isolation of rice genes possibly involved in the photoperiodic control of flowering by a<br />
fluorescent differential display method. ''Plant and Cell Physiology'' 43(5): 494-504.</ref> <br />
<ref name="ref4">Takeshi Izawa, Motohiro Mihara, Yuji Suzuki, Meenu Gupta, Hironori Itoh, et al. (2011) ''Os-GIGANTEA'' confers robust diurnal rhythms on the global transcriptome of rice in the field. ''The Plant Cell'' 23(5): 1741-1755.</ref> <br />
<ref name="ref5">Ryosuke Hayama, Shuji Yokoi, Shojiro Tamaki, Masahiro Yano, Ko Shimamoto (2003) Adaptation of photoperiodic control pathways produces short-day flowering in rice. ''Nature'' 422(6933): 719-722.</ref> <br />
<ref name="ref6">Bong-Soo Shin, Jeong-Hyun Lee, Jeong-Hwan Lee, Hyun-Joo Jeong, Choong-Hyo Yun, Jeong-Kook Kim (2004) Circadian regulation of rice (''Oryza sativa L.'') ''CONSTANS-like'' gene transcripts. ''Molecules and Cells'' 17(1): 10-16.</ref><br />
<br />
==Structured Information==<br />
{{JaponicaGene|<br />
GeneName = Os01g0182600|<br />
Description = GIGANTEA protein|<br />
Version = NM_001048755.1 GI:115434923 GeneID:4325329|<br />
Length = 9083 bp|<br />
Definition = Oryza sativa Japonica Group Os01g0182600, complete gene.|<br />
Source = Oryza sativa Japonica Group<br />
<br />
ORGANISM Oryza sativa Japonica Group<br />
Eukaryota; Viridiplantae; Streptophyta; Embryophyta; Tracheophyta;<br />
Spermatophyta; Magnoliophyta; Liliopsida; Poales; Poaceae; BEP<br />
clade; Ehrhartoideae; Oryzeae; Oryza.<br />
|<br />
Chromosome = [[:category:Japonica Chromosome 1|Chromosome 1]]|<br />
AP = Chromosome 1:4328362..4337444|<br />
CDS = 4328725..4329030,4329123..4329215,4329333..4329551,4329752..4329820,4329925..4331421<br>,4332220..4332429,4332563..4332713,4332791..4333031,4333898..4333956<br>,4334379..4334653,4334813..4334955,4335034..4335088,4335181..4335258<br>,4335351..4335437|<br />
GCID = <gbrowseImage1><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage1>|<br />
GSID = <gbrowseImage2><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage2>|<br />
CDNA = <cdnaseq>atgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactgcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcatagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcggcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctga</cdnaseq>|<br />
AA = <aaseq>MSASNEKWIDGLQFSSLFWPPPQDSQQKQAQILAYVEYFGQFTA DSEQFPEDIAQLIQSCYPSKEKRLVDEVLATFVLHHPEHGHAVVHPILSRIIDGTLSY DRNGFPFMSFISLFSHTSEKEYSEQWALACGEILRVLTHYNRPIFKVDHQHSEAECSS TSDQASSCESMEKRANGSPRNEPDRKPLRPLSPWITDILLAAPLGIRSDYFRWCGGVM GKYAAGGELKPPTTAYSRGSGKHPQLMPSTPRWAVANGAGVILSVCDEEVARYETANL TAAAVPALLLPPPTTPLDEHLVAGLPPLEPYARLFHRYYAIATPSATQRLLFGLLEAP PSWAPDALDAAVQLVELLRAAEDYDSGMRLPKNWMHLHFLRAIGTAMSMRAGIAADTS AALLFRILSQPTLLFPPLRHAEGVELHHEPLGGYVSSYKRQLEVPASEATIDATAQGI ASMLCAHGPDVEWRICTIWEAAYGLLPLSSSAVDLPEIVVAAPLQPPTLSWSLYLPLL KVFEYLPRGSPSEACLMRIFVATVEAILRRTFPSETSEQSRKPRSQSKNLAVAELRTM IHSLFVESCASMDLASRLLFVVLTVCVSHQALPGGSKRPTGSDNHSSEEVTNDSRLTN GRNRCKKRQGPVATFDSYVLAAVCALSCELQLFPFISKNGNHSNLKDSIKIVIPGKTT GISNELHNSISSAILHTRRILGILEALFSLKPSSVGTSWSYSSNEIVAAAMVAAHVSE LFRRSRPCLNALSALKQCKWDAEISTRASSLYHLIDLHGKTVTSIVNKAEPLEAHLTL TPVKKDEPPIEEKNINSSDGGALEKKDASRSHRKNGFARPLLKCAEDVILNGDVASTS GKAIASLQVEASDLANFLTMDRNGGYRGSQTLLRSVLSEKQELCFSVVSLLWQKLIAS PEMQMSAESTSAHQGWRKVVDALCDIVSASPTKASAAIVLQAEKDLQPWIARDDEQGQ KMWRVNQRIVKLIAELMRNHDSPEALVILASASDLLLRATDGMLVDGEACTLPQLELL EVTARAVHLIVEWGDSGVSVADGLSNLLKCRLSTTIRCLSHPSAHVRALSMSVLRDIL NSGQINSSKLIQGEHRNGIQSPTYQCLAASIINWQADVERCIEWEAHSRRATGLTLAF LTAAAKELGCPLTC</aaseq>|<br />
DNA = <dnaseqindica>8415..8720#8230..8322#7894..8112#7625..7693#6024..7520#5016..5225#4732..4882#4414..4654#3489..3547#2792..3066#2490..2632#2357..2411#2187..2264#2008..2094#ccgagccgtttctctccctctctctctcctcctccttcccctgcgcctcctccaccgcccgtgtcgccgccgcgcgcgtctatcccctcgccgccgccgccgtctacgccgcgtgaggtccggattggttgaaaggctgatcgaggtaatgcttttttatttttcccgtgcggttttagtttttctgtgctgtgcttggtggttttttctttaaaactatttttaggctggttttgatgatggcctttctaaggttgtcgtgtatggggggcgaggtggttcgggtggatttgtggtgttatgttgaggggatttttgtttggggggagagagttttggtggtagatgcggttgggattttgaagacgtgaaagggaaattagttttgaaatttgacggcgaggttgagctgattagtattgtaataattcgtggcatcggatagatcaacgcgagatggatgcttattactgcgaatttcaggaggtttttgtggtttttttaggcgagtagcgttgattctagctggattttgctgcttcgtggaggcgtttagatgtggtgcatacggctgtggatgtaatttgtgcggggataattactgaaaattaattcgcagtaagctagctctgctttaatattttttcccctgaaacagtaacaagggatcccttcctcatttcgttaacagtggaagtgaatacagacaccaaccgatgagcagagacacctgtaattaaacaccaaaagaagaacctgttttgttcctatatattttccatttctatgctttaggtaatatactttctggttcggtgtcatagtacccgaaatagaatctttgcttgccaaaaaaagaaagaataacaagatagaacaacatcattaaattctgtcttagacaatacatttataatcgaccaacaaaaacagaccgatgagtgacgtggaataacaaatgaagtataaatgtcacgaaaaacacatctattgattttgtgtgataagccataggagcctacgaggaaccaagtgaatggttctgtacaccacatccattagcattaccataaatttgcctatttctatgatgccttccttaacatatttatcgacatgtgagcttgcaccataactaaatgttatggtaaactttaccttataaatgattgcaacatgtaccttttcatttcgataattaaaaacgagaaagcttccacctttttcctagatatattttttcgtttgattatgaatttagttgtacttgacaataaaataatgcgcttgtaagattatgcaattttcgttgcattacctccctctcttttcatttgtcagtttttttctactccctccgtttcatattttaagtcgtttgacttttttcctagtcagactttgttaagtttgacaaagtttatagtaaaatttagcaatatctaaaatattaaattggtttcatcaaatctaacattgaatatattttgataatatgttttttttgtgttgaaaatacttctatatttttctataaacttagtcaaacttggaagaaatttcagtaagaaaaaagtcaaacggcttataatatgaaacggagggagtaaatttttttttgcattaattaaaattcatgtattttcctccttttttggtttttgttgtgcgatatattacagtgtagagactctgaagctcatatggttggcattggttgatatgataacgtgtcacgttgtgagatgcagacccccgttatgggggttgattgaatgaagtggatagcactcttataactttcttgctcgaagtttcttgataccaaggcaagtagcttataagaagatttggactttggtggcataaatggatgaactaactggcggcttcaacgtggtgttgattacaggtgcttttattgctaactggagcagtttgtgtagtttctttctttctctttgagaattctctcagtttcttgggcaggctcttgagctaccaagtatgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggtaacattattcctatttgattcttgttagaaattctatatgttgaacatacttgcctcaatgggaaacttggttgcgtctcctctctgaaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcaggtactgtatctctaagcatgtttagtatcttctgtgataaaatctttctccagtgtgttaacttttgcattctttttcttgcatgcttgcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattaggtaatttgatccagtgatcaatacatgttccacaaacaattgaagaaagagtttgacatatttttctcatatttccagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgaggtaaggcccctactctaatggttagttccaatttattggttcatttctctgtgtttgataagattttgcctctgctgtgcatttcaatattatatctaaactgtgctgcttatagcttatcaaactactcttgctcataaagatgcaacttttatgcagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgagtcataataaatgataacatattattgttctgtggactgaaaatctgaaatcctcctttcttgagaaatacaacctcaattttatggtgatattgacaagaaggaagagcgcatataatttaaagttaaaacacggaaaataatagaagtatgtgagcagaacgcaggagcaaggggtggaactcataacaagttcaaacttcatgcatgttcataattccccaatgcaagtatggcaagaaaaatagacaacttttaaaagtatgctaatggaactgcatgcattgtgtaaacttgcttttgtcaagcatgttggaacatactatatatttgctgctgatgatgaaaaatttgcttgagtactcttctattgctaatggacgcttaccattccctgttttgtcttcaaatgcatcaggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactggttggtaaaatcagcaatggaattttctatcgtgaattttttataattattgtaacattaattactcccatgtctttttattctgttgagataccatgtgacaatgtctacaattgcatattttgcattctttggtagatcagttcccttcttaagatccccaaagggaaactggagtatcaataacttactccctcaaataaaaaaaaagtgatactttaggcataaacacattcttcaaaacaaaactttgaatagcagtttcttttgtaatatattaataaagcatggaaaaattataatgtgaagaatgtacattcagggacaaatatactcatcatttttgcatggttgatcttaatgtccatagagagattgatcagaccaaaagtttatttttgacaggagtgagtatggattataataggagtgaatggcaatatgctgtgaaagtgagcattttatatttgtttactactagcatttggtgacaaagcaaatgatgtatattgtaaatcaaagtaaacaagtagaacatgtcctgtggcgacaagacataacctacaaattaaagttagcctgattgtttattttactgggagcacattttttctagattctagagtacatattattaaaaaaggtgtgatatgtttgcaaattactttttaatggaagtttgtaaattactttgaaacttgcattcagtttttttgcttcttaaaaaacatagtgcatatgggctttatgagctcatactatgcatgtttttctgcattggagtgggacctttatctttttcttgcaactcggatgctaatgcttattataaacacaagttactaacatgtgcttgggtcattcagcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcataggtaagctcttatgatcctgtccactgttgtaccatgtgtactttccaaatgattgctaacatgctatttttcgtcagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtac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/>
Link = [http://www.ncbi.nlm.nih.gov/nuccore/NM_001048755.1 RefSeq:Os01g0182600]|<br />
}}<br />
[[Category:Genes]]<br />
[[Category:Japonica mRNA]]<br />
[[Category:Oryza Sativa Japonica Group]]<br />
[[Category:Japonica Genes]]<br />
[[Category:Japonica Chromosome 1]]<br />
[[Category:Chromosome 1]]</div>Mikahttps://ngdc.cncb.ac.cn/ricewiki/index.php?title=Os01g0182600&diff=183819Os01g01826002014-06-11T03:05:05Z<p>Mika: /* Function */</p>
<hr />
<div>The rice gene of ''Gigantea(GI)'' is solo ortholog of ''GI'' in the Arabidopsis circadian system and related to photoperiodic control of flowering.<br />
<br />
==Annotated Information==<br />
===Function===<br />
*''OsGI'',a ''GI'' ortholog, regulates flowering time in response to photoperiodic conditions. Mutations in this gene delay flowering in inductive SDs (short-days),whereas flowering is marginally induced under LDs(long-days).In comparison, over expression of ''OsGI'' in transgenic rice causes late flowering in both SDs and LDs.In addition, ''OsGI'' has function to regulate ''Hd1'', a gene that also shows circadian rhythm<ref name="ref1" />. ''Os-GI'' affects gene expression of approximately 75% of genes among 27,201 genes and is required for strong amplitudes and proper phase-setting of global gene expression under natural field conditions<ref name="ref2" />. <br />
*The cDNA of ''OsGI'' firstly found in ''se5'' mutant that is completely photoperiod-insensitive, the expression patterns of rice and Arabidopsis ''GI'' gene are both photoperiodically regulated and are very similar under LD and SD conditions(FIG1)<ref name="ref3" />. ''GI'' is required for phase-dependent ''CO''transcription through the interaction with FKF1, which leads to cycling DNA binding with one finger (CDF) degradation and consequent derepression of CO. In addition, it has been known that GI protein interacts with ZEITLUPE (ZTL) F-box protein to control the diurnal rhythm of TOC1 protein in regulation of the Arabidopsis circadian clock<ref name="ref4" />. <br />
*''Os-GI'' also has function to maintain bioactive GA level through the regulation of the GA-deactivating enzyme genes in rice, ''Os-GI'' represses major ''OsGA2ox'' genes in order to maintain a proper bioactive GA level, ''osgi-1'' plants showed a semi-dwarf phenotype with elevated ''OsGA2ox'' gene expression <ref name="ref4" /> .<br />
<br />
===Expression===<br />
*''GI'' encodes a nuclear protein that defines the proper amplitude and period length of circadian rhythms. ''GI'' expression is regulated by the circadian clock, with a peak in transcript levels at 8 to 10 h after dawn. The ''GI'' protein acts between the circadian oscillator and ''CO'' to promote flowering by increasing ''CO'' and ''FT'' mRNA abundance<ref name="ref1" />. The expression of the ''OsGI'' messenger RNA was circadian-controlled and that its temporal expression pattern was very similar to that of ''GI'' under both SD and LD conditions. Decreased ''OsGI'' expression led to early flowering under LD conditions and late flowering under SD conditions relative to the wild type, and that this early flowering phenotype under LD conditions was the opposite of that of the Arabidopsis ''gi'' mutants, confirming the opposite role of the ''GI'' gene in flowering under LD conditions in rice and Arabidopsis<ref name="ref5" />.<br />
<br />
===Mutation===<br />
*The single ''osgi-1'' mutation extended the time to flowering only under SD conditions. Under LD conditions the single ''hd1'' mutation induced early flowering, but ''osgi-1'' plants exhibited no phenotypic change<ref name="ref4" /><br />
<br />
===Evolution===<br />
* There is some evolutionary conservation between rice circadian clock–related genes and those associated with Arabidopsis circadian clocks, where the circadian clock consists of interlocked subloops, it is also possible that ''Os-GI'' is a member of a subloop in rice circadian clocks<ref name="ref4" />.But there is a divergence in the photoperiodic pathways of these two species arises from the molecular mechanisms controlling the expression of ''FT/Hd3a'' by ''CO/Hd1(Se1)''<ref name="ref5" />(figure 3)<br />
<br />
==Labs working on this gene==<br />
*Photosynthesis and Photobiology Research Unit, National Institute of Agrobiological Sciences, 2-1-2 Kannondai, Tsukuba 305-8602, Japan<br />
*Laboratory of Plant Molecular Genetics, Nara Institute of Science and Technology, 8916-5 Takayama, Ikoma 630-0101, Japan<br />
*Plant Genomics Research Unit, National Institute of Agrobiological Sciences, 2-1-2 Kannondai, Tsukuba 305-8602, Japan<br />
<br />
==Knowledge Extension==<br />
*The rice orthologues of the two key regulators of flowering time in Arabidopsis, ''CO'' and ''FT'', were recently shown to be important in the photoperiodic control of flowering in rice. The rice ''Hd1(Se1)''gene, which is an orthologue of the Arabidopsis ''CO'' gene, was required for the suppression of flowering under LD conditions and for the promotion of flowering under SD conditions. Furthermore, the rice ''Hd3a'' gene, an orthologue of the Arabidopsis ''FT'' gene, was shown to be an activator of flowering in rice. It was previously demonstrated that ''CO'' controls the flowering time by integrating signals from the circadian clock and light to regulate ''FT'' expression in Arabidopsis<ref name="ref5" />.Unlike Arabidopsis ''FT'', expression of ''Hd3a'' mRNA is more or less opposite to that of ''Hd1'' under inductive short-day conditions<ref name="ref6" />.<br />
<br />
==References==<br />
<references><br />
<ref name="ref1">Song Lim Kim, Shinyoung Lee, Hyo Jung Kim, Hong Gil Nam, Gynheung An（2007）''OsMADS51'' is a short-day flowering promoter that functions upstream of ''Ehd1'', ''OsMADS14'', and ''Hd3a''.''Plant Physiology'' 145(4): 1484-1494.</ref><br />
<ref name="ref2">Hironori Itoh, Takeshi Izawa (2011) A study of phytohormone biosynthetic gene expression using a circadian clock-related mutant in rice. ''Plant Signaling & Behavior'' 6(12): 1932-1936.</ref> <br />
<ref name="ref3">Ryousuke Hayama, Takeshi Izawa, Ko Shimamoto (2002) Isolation of rice genes possibly involved in the photoperiodic control of flowering by a<br />
fluorescent differential display method. ''Plant and Cell Physiology'' 43(5): 494-504.</ref> <br />
<ref name="ref4">Takeshi Izawa, Motohiro Mihara, Yuji Suzuki, Meenu Gupta, Hironori Itoh, et al. (2011) ''Os-GIGANTEA'' confers robust diurnal rhythms on the global transcriptome of rice in the field. ''The Plant Cell'' 23(5): 1741-1755.</ref> <br />
<ref name="ref5">Ryosuke Hayama, Shuji Yokoi, Shojiro Tamaki, Masahiro Yano, Ko Shimamoto (2003) Adaptation of photoperiodic control pathways produces short-day flowering in rice. ''Nature'' 422(6933): 719-722.</ref> <br />
<ref name="ref6">Bong-Soo Shin, Jeong-Hyun Lee, Jeong-Hwan Lee, Hyun-Joo Jeong, Choong-Hyo Yun, Jeong-Kook Kim (2004) Circadian regulation of rice (''Oryza sativa L.'') ''CONSTANS-like'' gene transcripts. ''Molecules and Cells'' 17(1): 10-16.</ref><br />
<br />
==Structured Information==<br />
{{JaponicaGene|<br />
GeneName = Os01g0182600|<br />
Description = GIGANTEA protein|<br />
Version = NM_001048755.1 GI:115434923 GeneID:4325329|<br />
Length = 9083 bp|<br />
Definition = Oryza sativa Japonica Group Os01g0182600, complete gene.|<br />
Source = Oryza sativa Japonica Group<br />
<br />
ORGANISM Oryza sativa Japonica Group<br />
Eukaryota; Viridiplantae; Streptophyta; Embryophyta; Tracheophyta;<br />
Spermatophyta; Magnoliophyta; Liliopsida; Poales; Poaceae; BEP<br />
clade; Ehrhartoideae; Oryzeae; Oryza.<br />
|<br />
Chromosome = [[:category:Japonica Chromosome 1|Chromosome 1]]|<br />
AP = Chromosome 1:4328362..4337444|<br />
CDS = 4328725..4329030,4329123..4329215,4329333..4329551,4329752..4329820,4329925..4331421<br>,4332220..4332429,4332563..4332713,4332791..4333031,4333898..4333956<br>,4334379..4334653,4334813..4334955,4335034..4335088,4335181..4335258<br>,4335351..4335437|<br />
GCID = <gbrowseImage1><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage1>|<br />
GSID = <gbrowseImage2><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage2>|<br />
CDNA = <cdnaseq>atgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactgcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcatagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcggcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctga</cdnaseq>|<br />
AA = <aaseq>MSASNEKWIDGLQFSSLFWPPPQDSQQKQAQILAYVEYFGQFTA DSEQFPEDIAQLIQSCYPSKEKRLVDEVLATFVLHHPEHGHAVVHPILSRIIDGTLSY DRNGFPFMSFISLFSHTSEKEYSEQWALACGEILRVLTHYNRPIFKVDHQHSEAECSS TSDQASSCESMEKRANGSPRNEPDRKPLRPLSPWITDILLAAPLGIRSDYFRWCGGVM GKYAAGGELKPPTTAYSRGSGKHPQLMPSTPRWAVANGAGVILSVCDEEVARYETANL TAAAVPALLLPPPTTPLDEHLVAGLPPLEPYARLFHRYYAIATPSATQRLLFGLLEAP PSWAPDALDAAVQLVELLRAAEDYDSGMRLPKNWMHLHFLRAIGTAMSMRAGIAADTS AALLFRILSQPTLLFPPLRHAEGVELHHEPLGGYVSSYKRQLEVPASEATIDATAQGI ASMLCAHGPDVEWRICTIWEAAYGLLPLSSSAVDLPEIVVAAPLQPPTLSWSLYLPLL KVFEYLPRGSPSEACLMRIFVATVEAILRRTFPSETSEQSRKPRSQSKNLAVAELRTM IHSLFVESCASMDLASRLLFVVLTVCVSHQALPGGSKRPTGSDNHSSEEVTNDSRLTN GRNRCKKRQGPVATFDSYVLAAVCALSCELQLFPFISKNGNHSNLKDSIKIVIPGKTT GISNELHNSISSAILHTRRILGILEALFSLKPSSVGTSWSYSSNEIVAAAMVAAHVSE LFRRSRPCLNALSALKQCKWDAEISTRASSLYHLIDLHGKTVTSIVNKAEPLEAHLTL TPVKKDEPPIEEKNINSSDGGALEKKDASRSHRKNGFARPLLKCAEDVILNGDVASTS GKAIASLQVEASDLANFLTMDRNGGYRGSQTLLRSVLSEKQELCFSVVSLLWQKLIAS PEMQMSAESTSAHQGWRKVVDALCDIVSASPTKASAAIVLQAEKDLQPWIARDDEQGQ KMWRVNQRIVKLIAELMRNHDSPEALVILASASDLLLRATDGMLVDGEACTLPQLELL EVTARAVHLIVEWGDSGVSVADGLSNLLKCRLSTTIRCLSHPSAHVRALSMSVLRDIL NSGQINSSKLIQGEHRNGIQSPTYQCLAASIINWQADVERCIEWEAHSRRATGLTLAF LTAAAKELGCPLTC</aaseq>|<br />
DNA = <dnaseqindica>8415..8720#8230..8322#7894..8112#7625..7693#6024..7520#5016..5225#4732..4882#4414..4654#3489..3547#2792..3066#2490..2632#2357..2411#2187..2264#2008..2094#ccgagccgtttctctccctctctctctcctcctccttcccctgcgcctcctccaccgcccgtgtcgccgccgcgcgcgtctatcccctcgccgccgccgccgtctacgccgcgtgaggtccggattggttgaaaggctgatcgaggtaatgcttttttatttttcccgtgcggttttagtttttctgtgctgtgcttggtggttttttctttaaaactatttttaggctggttttgatgatggcctttctaaggttgtcgtgtatggggggcgaggtggttcgggtggatttgtggtgttatgttgaggggatttttgtttggggggagagagttttggtggtagatgcggttgggattttgaagacgtgaaagggaaattagttttgaaatttgacggcgaggttgagctgattagtattgtaataattcgtggcatcggatagatcaacgcgagatggatgcttattactgcgaatttcaggaggtttttgtggtttttttaggcgagtagcgttgattctagctggattttgctgcttcgtggaggcgtttagatgtggtgcatacggctgtggatgtaatttgtgcggggataattactgaaaattaattcgcagtaagctagctctgctttaatattttttcccctgaaacagtaacaagggatcccttcctcatttcgttaacagtggaagtgaatacagacaccaaccgatgagcagagacacctgtaattaaacaccaaaagaagaacctgttttgttcctatatattttccatttctatgctttaggtaatatactttctggttcggtgtcatagtacccgaaatagaatctttgcttgccaaaaaaagaaagaataacaagatagaacaacatcattaaattctgtcttagacaatacatttataatcgaccaacaaaaacagaccgatgagtgacgtggaataacaaatgaagtataaatgtcacgaaaaacacatctattgattttgtgtgataagccataggagcctacgaggaaccaagtgaatggttctgtacaccacatccattagcattaccataaatttgcctatttctatgatgccttccttaacatatttatcgacatgtgagcttgcaccataactaaatgttatggtaaactttaccttataaatgattgcaacatgtaccttttcatttcgataattaaaaacgagaaagcttccacctttttcctagatatattttttcgtttgattatgaatttagttgtacttgacaataaaataatgcgcttgtaagattatgcaattttcgttgcattacctccctctcttttcatttgtcagtttttttctactccctccgtttcatattttaagtcgtttgacttttttcctagtcagactttgttaagtttgacaaagtttatagtaaaatttagcaatatctaaaatattaaattggtttcatcaaatctaacattgaatatattttgataatatgttttttttgtgttgaaaatacttctatatttttctataaacttagtcaaacttggaagaaatttcagtaagaaaaaagtcaaacggcttataatatgaaacggagggagtaaatttttttttgcattaattaaaattcatgtattttcctccttttttggtttttgttgtgcgatatattacagtgtagagactctgaagctcatatggttggcattggttgatatgataacgtgtcacgttgtgagatgcagacccccgttatgggggttgattgaatgaagtggatagcactcttataactttcttgctcgaagtttcttgataccaaggcaagtagcttataagaagatttggactttggtggcataaatggatgaactaactggcggcttcaacgtggtgttgattacaggtgcttttattgctaactggagcagtttgtgtagtttctttctttctctttgagaattctctcagtttcttgggcaggctcttgagctaccaagtatgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggtaacattattcctatttgattcttgttagaaattctatatgttgaacatacttgcctcaatgggaaacttggttgcgtctcctctctgaaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcaggtactgtatctctaagcatgtttagtatcttctgtgataaaatctttctccagtgtgttaacttttgcattctttttcttgcatgcttgcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattaggtaatttgatccagtgatcaatacatgttccacaaacaattgaagaaagagtttgacatatttttctcatatttccagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgaggtaaggcccctactctaatggttagttccaatttattggttcatttctctgtgtttgataagattttgcctctgctgtgcatttcaatattatatctaaactgtgctgcttatagcttatcaaactactcttgctcataaagatgcaacttttatgcagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgagtcataataaatgataacatattattgttctgtggactgaaaatctgaaatcctcctttcttgagaaatacaacctcaattttatggtgatattgacaagaaggaagagcgcatataatttaaagttaaaacacggaaaataatagaagtatgtgagcagaacgcaggagcaaggggtggaactcataacaagttcaaacttcatgcatgttcataattccccaatgcaagtatggcaagaaaaatagacaacttttaaaagtatgctaatggaactgcatgcattgtgtaaacttgcttttgtcaagcatgttggaacatactatatatttgctgctgatgatgaaaaatttgcttgagtactcttctattgctaatggacgcttaccattccctgttttgtcttcaaatgcatcaggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactggttggtaaaatcagcaatggaattttctatcgtgaattttttataattattgtaacattaattactcccatgtctttttattctgttgagataccatgtgacaatgtctacaattgcatattttgcattctttggtagatcagttcccttcttaagatccccaaagggaaactggagtatcaataacttactccctcaaataaaaaaaaagtgatactttaggcataaacacattcttcaaaacaaaactttgaatagcagtttcttttgtaatatattaataaagcatggaaaaattataatgtgaagaatgtacattcagggacaaatatactcatcatttttgcatggttgatcttaatgtccatagagagattgatcagaccaaaagtttatttttgacaggagtgagtatggattataataggagtgaatggcaatatgctgtgaaagtgagcattttatatttgtttactactagcatttggtgacaaagcaaatgatgtatattgtaaatcaaagtaaacaagtagaacatgtcctgtggcgacaagacataacctacaaattaaagttagcctgattgtttattttactgggagcacattttttctagattctagagtacatattattaaaaaaggtgtgatatgtttgcaaattactttttaatggaagtttgtaaattactttgaaacttgcattcagtttttttgcttcttaaaaaacatagtgcatatgggctttatgagctcatactatgcatgtttttctgcattggagtgggacctttatctttttcttgcaactcggatgctaatgcttattataaacacaagttactaacatgtgcttgggtcattcagcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcataggtaagctcttatgatcctgtccactgttgtaccatgtgtactttccaaatgattgctaacatgctatttttcgtcagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcgggtatatcatacatctctctggctgttgtttcagtgctatcaatcaagatgacacatcatgcaatattatcataacagcacattatgtgttttgaatattttattgattgtcatagctttgttttatgttttagcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcaggtatgcagtgtttcctgcacctacttgctacctattctgtatttgatgtattcaatttcctttttatcagttttctcgaaatatatgctaataggcttgtcactaaacggctacattatccagactctgttctgaatttttttttcattaagtactgttagctgccagattacatcatcatgactaaagtagtttagctacaatattgcaaagctagatttgtctagcctctattgttttgaatagaaaaggcaggtatgcagtgtttcctgcacctacttgctacctattctgtatttgatgtattcaatttcctttttatcagttttctcgaaatatatgctaataggcttgtcactaaacggctacattatccagactctgttctgaatttttttttcattaagtactgttagctgccagattacatcatcatgactaaagtagtttagctacaatattgcaaagctagatttgtctagcctctattgttttgaatagaaaattaaatcttttggtcatttttttgtaaccttttaggagtctctgtatgcttgtcgtcaacatgctatgataagtttatatcagttcttttggagctatgatgtattcttgtatattttatgactgctaggatattatgttaaaactgttttaaaggtatcatcaaacaagttaatatagatattgctacctaccaaagttggaatttgattatgtcgtaaaatatatgcaccctacatgttttagcttcatattgacttatattttcatatgtgctccttcaactgcacaaagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggtatggtcattcctgcttataagataccagaataatgctatgaagaccgtccttggttgtcagcattcttatacacaatgataatctttgtttttcatgggtaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggtcaagatattgttattgttatatttacatacttgttatcatatttaaaagcttcatattgtatgtacctttgaaaaaaaaagttccatactgtagcttcttttgttctcatttcttggtttcaaagcatcagcatcattaccgatataaaatatacccatttcctcagtcatcttccccgactcttgtatctttcacaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagaggtaacaccttcattcatgaggtccttaggtgcaaatcccagtgaagaataatcagttctactttgtatggttcatctcgcttactgaccggttcatctcgggatcaattacgagcagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaaggcatgtcctcttctctgatctcactcacgttcatgccaaacttactcatcatcatccatgatccatcatcctgactgaatcttgatatgcagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctgacaaggccacaccatgactgtcactgcaagcagctgctgtaggatcagcgagtaggaaaggatgacttgccggccagtttcctgagatgtgatttaagactgatattagctgatgttcccaagcatgatacggggcttgctcccaagatgtgatttttactttacttttggttaatgtgctaccgctgacattagatgttcaagcatatgaaaactgcttgtgctgtaacttgtatgtctgtaacgtaaaaatctttccatatggtaaccggggaagcagaggtatatccttttctgctgttgtatctatttccagtttcctgctgaacttgcaaatgaatattcagcaatttccgttttgctc</dnaseqindica>|<br />
Link = [http://www.ncbi.nlm.nih.gov/nuccore/NM_001048755.1 RefSeq:Os01g0182600]|<br />
}}<br />
[[Category:Genes]]<br />
[[Category:Japonica mRNA]]<br />
[[Category:Oryza Sativa Japonica Group]]<br />
[[Category:Japonica Genes]]<br />
[[Category:Japonica Chromosome 1]]<br />
[[Category:Chromosome 1]]</div>Mikahttps://ngdc.cncb.ac.cn/ricewiki/index.php?title=Os01g0182600&diff=183808Os01g01826002014-06-11T01:42:42Z<p>Mika: /* Function */</p>
<hr />
<div>The rice gene of ''Gigantea(GI)'' is solo ortholog of ''GI'' in the Arabidopsis circadian system and related to photoperiodic control of flowering.<br />
<br />
==Annotated Information==<br />
===Function===<br />
*''OsGI'',a ''GI'' ortholog, regulates flowering time in response to photoperiodic conditions. Mutations in this gene delay flowering in inductive SDs (short-days),whereas flowering is marginally induced under LDs(long-days).In comparison, over expression of ''OsGI'' in transgenic rice causes late flowering in both SDs and LDs.In addition, ''OsGI'' has function to regulate ''Hd1'', a gene that also shows circadian rhythm<ref name="ref1" />. ''Os-GI'' affects gene expression of approximately 75% of genes among 27,201 genes and is required for strong amplitudes and proper phase-setting of global gene expression under natural field conditions<ref name="ref2" />. <br />
*The cDNA of ''OsGI'' firstly found in ''se5'' mutant that is completely photoperiod-insensitive, the expression patterns of rice and Arabidopsis ''GI'' gene are both photoperiodically regulated and are very similar under LD and SD conditions(FIG1)<ref name="ref3" />. ''GI'' is required for phase-dependent ''CO''transcription through the interaction with FKF1, which leads to cycling DNA binding with one finger (CDF) degradation and consequent derepression of CO. In addition, it has been known that GI protein interacts with ZEITLUPE (ZTL) F-box protein to control the diurnal rhythm of TOC1 protein in regulation of the Arabidopsis circadian clock<ref name="ref4" />. <br />
*''Os-GI'' also has function to maintain bioactive GA level through the regulation of the GA-deactivating enzyme genes in rice, ''Os-GI'' represses major ''OsGA2ox'' genes in order to maintain a proper bioactive GA level, ''osgi-1'' plants showed a semi-dwarf phenotype with elevated ''OsGA2ox'' gene expression <ref name="ref4" /> [[File:limeijia1.png|right|thumb|150px|''WT rice VS. osgi-1 plants (from reference <ref name="ref4" />).'']].<br />
<br />
===Expression===<br />
*''GI'' encodes a nuclear protein that defines the proper amplitude and period length of circadian rhythms. ''GI'' expression is regulated by the circadian clock, with a peak in transcript levels at 8 to 10 h after dawn. The ''GI'' protein acts between the circadian oscillator and ''CO'' to promote flowering by increasing ''CO'' and ''FT'' mRNA abundance<ref name="ref1" />. The expression of the ''OsGI'' messenger RNA was circadian-controlled and that its temporal expression pattern was very similar to that of ''GI'' under both SD and LD conditions. Decreased ''OsGI'' expression led to early flowering under LD conditions and late flowering under SD conditions relative to the wild type, and that this early flowering phenotype under LD conditions was the opposite of that of the Arabidopsis ''gi'' mutants, confirming the opposite role of the ''GI'' gene in flowering under LD conditions in rice and Arabidopsis<ref name="ref5" />.<br />
<br />
===Mutation===<br />
*The single ''osgi-1'' mutation extended the time to flowering only under SD conditions. Under LD conditions the single ''hd1'' mutation induced early flowering, but ''osgi-1'' plants exhibited no phenotypic change<ref name="ref4" /><br />
<br />
===Evolution===<br />
* There is some evolutionary conservation between rice circadian clock–related genes and those associated with Arabidopsis circadian clocks, where the circadian clock consists of interlocked subloops, it is also possible that ''Os-GI'' is a member of a subloop in rice circadian clocks<ref name="ref4" />.But there is a divergence in the photoperiodic pathways of these two species arises from the molecular mechanisms controlling the expression of ''FT/Hd3a'' by ''CO/Hd1(Se1)''<ref name="ref5" />(figure 3)<br />
<br />
==Labs working on this gene==<br />
*Photosynthesis and Photobiology Research Unit, National Institute of Agrobiological Sciences, 2-1-2 Kannondai, Tsukuba 305-8602, Japan<br />
*Laboratory of Plant Molecular Genetics, Nara Institute of Science and Technology, 8916-5 Takayama, Ikoma 630-0101, Japan<br />
*Plant Genomics Research Unit, National Institute of Agrobiological Sciences, 2-1-2 Kannondai, Tsukuba 305-8602, Japan<br />
<br />
==Knowledge Extension==<br />
*The rice orthologues of the two key regulators of flowering time in Arabidopsis, ''CO'' and ''FT'', were recently shown to be important in the photoperiodic control of flowering in rice. The rice ''Hd1(Se1)''gene, which is an orthologue of the Arabidopsis ''CO'' gene, was required for the suppression of flowering under LD conditions and for the promotion of flowering under SD conditions. Furthermore, the rice ''Hd3a'' gene, an orthologue of the Arabidopsis ''FT'' gene, was shown to be an activator of flowering in rice. It was previously demonstrated that ''CO'' controls the flowering time by integrating signals from the circadian clock and light to regulate ''FT'' expression in Arabidopsis<ref name="ref5" />.Unlike Arabidopsis ''FT'', expression of ''Hd3a'' mRNA is more or less opposite to that of ''Hd1'' under inductive short-day conditions<ref name="ref6" />.<br />
<br />
==References==<br />
<references><br />
<ref name="ref1">Song Lim Kim, Shinyoung Lee, Hyo Jung Kim, Hong Gil Nam, Gynheung An（2007）''OsMADS51'' is a short-day flowering promoter that functions upstream of ''Ehd1'', ''OsMADS14'', and ''Hd3a''.''Plant Physiology'' 145(4): 1484-1494.</ref><br />
<ref name="ref2">Hironori Itoh, Takeshi Izawa (2011) A study of phytohormone biosynthetic gene expression using a circadian clock-related mutant in rice. ''Plant Signaling & Behavior'' 6(12): 1932-1936.</ref> <br />
<ref name="ref3">Ryousuke Hayama, Takeshi Izawa, Ko Shimamoto (2002) Isolation of rice genes possibly involved in the photoperiodic control of flowering by a<br />
fluorescent differential display method. ''Plant and Cell Physiology'' 43(5): 494-504.</ref> <br />
<ref name="ref4">Takeshi Izawa, Motohiro Mihara, Yuji Suzuki, Meenu Gupta, Hironori Itoh, et al. (2011) ''Os-GIGANTEA'' confers robust diurnal rhythms on the global transcriptome of rice in the field. ''The Plant Cell'' 23(5): 1741-1755.</ref> <br />
<ref name="ref5">Ryosuke Hayama, Shuji Yokoi, Shojiro Tamaki, Masahiro Yano, Ko Shimamoto (2003) Adaptation of photoperiodic control pathways produces short-day flowering in rice. ''Nature'' 422(6933): 719-722.</ref> <br />
<ref name="ref6">Bong-Soo Shin, Jeong-Hyun Lee, Jeong-Hwan Lee, Hyun-Joo Jeong, Choong-Hyo Yun, Jeong-Kook Kim (2004) Circadian regulation of rice (''Oryza sativa L.'') ''CONSTANS-like'' gene transcripts. ''Molecules and Cells'' 17(1): 10-16.</ref><br />
<br />
==Structured Information==<br />
{{JaponicaGene|<br />
GeneName = Os01g0182600|<br />
Description = GIGANTEA protein|<br />
Version = NM_001048755.1 GI:115434923 GeneID:4325329|<br />
Length = 9083 bp|<br />
Definition = Oryza sativa Japonica Group Os01g0182600, complete gene.|<br />
Source = Oryza sativa Japonica Group<br />
<br />
ORGANISM Oryza sativa Japonica Group<br />
Eukaryota; Viridiplantae; Streptophyta; Embryophyta; Tracheophyta;<br />
Spermatophyta; Magnoliophyta; Liliopsida; Poales; Poaceae; BEP<br />
clade; Ehrhartoideae; Oryzeae; Oryza.<br />
|<br />
Chromosome = [[:category:Japonica Chromosome 1|Chromosome 1]]|<br />
AP = Chromosome 1:4328362..4337444|<br />
CDS = 4328725..4329030,4329123..4329215,4329333..4329551,4329752..4329820,4329925..4331421<br>,4332220..4332429,4332563..4332713,4332791..4333031,4333898..4333956<br>,4334379..4334653,4334813..4334955,4335034..4335088,4335181..4335258<br>,4335351..4335437|<br />
GCID = <gbrowseImage1><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage1>|<br />
GSID = <gbrowseImage2><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage2>|<br />
CDNA = <cdnaseq>atgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactgcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcatagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcggcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctga</cdnaseq>|<br />
AA = <aaseq>MSASNEKWIDGLQFSSLFWPPPQDSQQKQAQILAYVEYFGQFTA DSEQFPEDIAQLIQSCYPSKEKRLVDEVLATFVLHHPEHGHAVVHPILSRIIDGTLSY DRNGFPFMSFISLFSHTSEKEYSEQWALACGEILRVLTHYNRPIFKVDHQHSEAECSS TSDQASSCESMEKRANGSPRNEPDRKPLRPLSPWITDILLAAPLGIRSDYFRWCGGVM GKYAAGGELKPPTTAYSRGSGKHPQLMPSTPRWAVANGAGVILSVCDEEVARYETANL TAAAVPALLLPPPTTPLDEHLVAGLPPLEPYARLFHRYYAIATPSATQRLLFGLLEAP PSWAPDALDAAVQLVELLRAAEDYDSGMRLPKNWMHLHFLRAIGTAMSMRAGIAADTS AALLFRILSQPTLLFPPLRHAEGVELHHEPLGGYVSSYKRQLEVPASEATIDATAQGI ASMLCAHGPDVEWRICTIWEAAYGLLPLSSSAVDLPEIVVAAPLQPPTLSWSLYLPLL KVFEYLPRGSPSEACLMRIFVATVEAILRRTFPSETSEQSRKPRSQSKNLAVAELRTM IHSLFVESCASMDLASRLLFVVLTVCVSHQALPGGSKRPTGSDNHSSEEVTNDSRLTN GRNRCKKRQGPVATFDSYVLAAVCALSCELQLFPFISKNGNHSNLKDSIKIVIPGKTT GISNELHNSISSAILHTRRILGILEALFSLKPSSVGTSWSYSSNEIVAAAMVAAHVSE LFRRSRPCLNALSALKQCKWDAEISTRASSLYHLIDLHGKTVTSIVNKAEPLEAHLTL TPVKKDEPPIEEKNINSSDGGALEKKDASRSHRKNGFARPLLKCAEDVILNGDVASTS GKAIASLQVEASDLANFLTMDRNGGYRGSQTLLRSVLSEKQELCFSVVSLLWQKLIAS PEMQMSAESTSAHQGWRKVVDALCDIVSASPTKASAAIVLQAEKDLQPWIARDDEQGQ KMWRVNQRIVKLIAELMRNHDSPEALVILASASDLLLRATDGMLVDGEACTLPQLELL EVTARAVHLIVEWGDSGVSVADGLSNLLKCRLSTTIRCLSHPSAHVRALSMSVLRDIL NSGQINSSKLIQGEHRNGIQSPTYQCLAASIINWQADVERCIEWEAHSRRATGLTLAF LTAAAKELGCPLTC</aaseq>|<br />
DNA = <dnaseqindica>8415..8720#8230..8322#7894..8112#7625..7693#6024..7520#5016..5225#4732..4882#4414..4654#3489..3547#2792..3066#2490..2632#2357..2411#2187..2264#2008..2094#ccgagccgtttctctccctctctctctcctcctccttcccctgcgcctcctccaccgcccgtgtcgccgccgcgcgcgtctatcccctcgccgccgccgccgtctacgccgcgtgaggtccggattggttgaaaggctgatcgaggtaatgcttttttatttttcccgtgcggttttagtttttctgtgctgtgcttggtggttttttctttaaaactatttttaggctggttttgatgatggcctttctaaggttgtcgtgtatggggggcgaggtggttcgggtggatttgtggtgttatgttgaggggatttttgtttggggggagagagttttggtggtagatgcggttgggattttgaagacgtgaaagggaaattagttttgaaatttgacggcgaggttgagctgattagtattgtaataattcgtggcatcggatagatcaacgcgagatggatgcttattactgcgaatttcaggaggtttttgtggtttttttaggcgagtagcgttgattctagctggattttgctgcttcgtggaggcgtttagatgtggtgcatacggctgtggatgtaatttgtgcggggataattactgaaaattaattcgcagtaagctagctctgctttaatattttttcccctgaaacagtaacaagggatcccttcctcatttcgttaacagtggaagtgaatacagacaccaaccgatgagcagagacacctgtaattaaacaccaaaagaagaacctgttttgttcctatatattttccatttctatgctttaggtaatatactttctggttcggtgtcatagtacccgaaatagaatctttgcttgccaaaaaaagaaagaataacaagatagaacaacatcattaaattctgtcttagacaatacatttataatcgaccaacaaaaacagaccgatgagtgacgtggaataacaaatgaagtataaatgtcacgaaaaacacatctattgattttgtgtgataagccataggagcctacgaggaaccaagtgaatggttctgtacaccacatccattagcattaccataaatttgcctatttctatgatgccttccttaacatatttatcgacatgtgagcttgcaccataactaaatgttatggtaaactttaccttataaatgattgcaacatgtaccttttcatttcgataattaaaaacgagaaagcttccacctttttcctagatatattttttcgtttgattatgaatttagttgtacttgacaataaaataatgcgcttgtaagattatgcaattttcgttgcattacctccctctcttttcatttgtcagtttttttctactccctccgtttcatattttaagtcgtttgacttttttcctagtcagactttgttaagtttgacaaagtttatagtaaaatttagcaatatctaaaatattaaattggtttcatcaaatctaacattgaatatattttgataatatgttttttttgtgttgaaaatacttctatatttttctataaacttagtcaaacttggaagaaatttcagtaagaaaaaagtcaaacggcttataatatgaaacggagggagtaaatttttttttgcattaattaaaattcatgtattttcctccttttttggtttttgttgtgcgatatattacagtgtagagactctgaagctcatatggttggcattggttgatatgataacgtgtcacgttgtgagatgcagacccccgttatgggggttgattgaatgaagtggatagcactcttataactttcttgctcgaagtttcttgataccaaggcaagtagcttataagaagatttggactttggtggcataaatggatgaactaactggcggcttcaacgtggtgttgattacaggtgcttttattgctaactggagcagtttgtgtagtttctttctttctctttgagaattctctcagtttcttgggcaggctcttgagctaccaagtatgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggtaacattattcctatttgattcttgttagaaattctatatgttgaacatacttgcctcaatgggaaacttggttgcgtctcctctctgaaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcaggtactgtatctctaagcatgtttagtatcttctgtgataaaatctttctccagtgtgttaacttttgcattctttttcttgcatgcttgcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattaggtaatttgatccagtgatcaatacatgttccacaaacaattgaagaaagagtttgacatatttttctcatatttccagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgaggtaaggcccctactctaatggttagttccaatttattggttcatttctctgtgtttgataagattttgcctctgctgtgcatttcaatattatatctaaactgtgctgcttatagcttatcaaactactcttgctcataaagatgcaacttttatgcagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgagtcataataaatgataacatattattgttctgtggactgaaaatctgaaatcctcctttcttgagaaatacaacctcaattttatggtgatattgacaagaaggaagagcgcatataatttaaagttaaaacacggaaaataatagaagtatgtgagcagaacgcaggagcaaggggtggaactcataacaagttcaaacttcatgcatgttcataattccccaatgcaagtatggcaagaaaaatagacaacttttaaaagtatgctaatggaactgcatgcattgtgtaaacttgcttttgtcaagcatgttggaacatactatatatttgctgctgatgatgaaaaatttgcttgagtactcttctattgctaatggacgcttaccattccctgttttgtcttcaaatgcatcaggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactggttggtaaaatcagcaatggaattttctatcgtgaattttttataattattgtaacattaattactcccatgtctttttattctgttgagataccatgtgacaatgtctacaattgcatattttgcattctttggtagatcagttcccttcttaagatccccaaagggaaactggagtatcaataacttactccctcaaataaaaaaaaagtgatactttaggcataaacacattcttcaaaacaaaactttgaatagcagtttcttttgtaatatattaataaagcatggaaaaattataatgtgaagaatgtacattcagggacaaatatactcatcatttttgcatggttgatcttaatgtccatagagagattgatcagaccaaaagtttatttttgacaggagtgagtatggattataataggagtgaatggcaatatgctgtgaaagtgagcattttatatttgtttactactagcatttggtgacaaagcaaatgatgtatattgtaaatcaaagtaaacaagtagaacatgtcctgtggcgacaagacataacctacaaattaaagttagcctgattgtttattttactgggagcacattttttctagattctagagtacatattattaaaaaaggtgtgatatgtttgcaaattactttttaatggaagtttgtaaattactttgaaacttgcattcagtttttttgcttcttaaaaaacatagtgcatatgggctttatgagctcatactatgcatgtttttctgcattggagtgggacctttatctttttcttgcaactcggatgctaatgcttattataaacacaagttactaacatgtgcttgggtcattcagcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcataggtaagctcttatgatcctgtccactgttgtaccatgtgtactttccaaatgattgctaacatgctatttttcgtcagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtac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/>
Link = [http://www.ncbi.nlm.nih.gov/nuccore/NM_001048755.1 RefSeq:Os01g0182600]|<br />
}}<br />
[[Category:Genes]]<br />
[[Category:Japonica mRNA]]<br />
[[Category:Oryza Sativa Japonica Group]]<br />
[[Category:Japonica Genes]]<br />
[[Category:Japonica Chromosome 1]]<br />
[[Category:Chromosome 1]]</div>Mikahttps://ngdc.cncb.ac.cn/ricewiki/index.php?title=Os01g0182600&diff=183806Os01g01826002014-06-11T01:42:11Z<p>Mika: </p>
<hr />
<div>The rice gene of ''Gigantea(GI)'' is solo ortholog of ''GI'' in the Arabidopsis circadian system and related to photoperiodic control of flowering.<br />
<br />
==Annotated Information==<br />
===Function===<br />
*''OsGI'',a ''GI'' ortholog, regulates flowering time in response to photoperiodic conditions. Mutations in this gene delay flowering in inductive SDs (short-days),whereas flowering is marginally induced under LDs(long-days).In comparison, over expression of ''OsGI'' in transgenic rice causes late flowering in both SDs and LDs.In addition, ''OsGI'' has function to regulate ''Hd1'', a gene that also shows circadian rhythm<ref name="ref1" />. ''Os-GI'' affects gene expression of approximately 75% of genes among 27,201 genes and is required for strong amplitudes and proper phase-setting of global gene expression under natural field conditions<ref name="ref2" />. <br />
*The cDNA of ''OsGI'' firstly found in ''se5'' mutant that is completely photoperiod-insensitive, the expression patterns of rice and Arabidopsis ''GI'' gene are both photoperiodically regulated and are very similar under LD and SD conditions(FIG1)<ref name="ref3" />. ''GI'' is required for phase-dependent ''CO''transcription through the interaction with FKF1, which leads to cycling DNA binding with one finger (CDF) degradation and consequent derepression of CO. In addition, it has been known that GI protein interacts with ZEITLUPE (ZTL) F-box protein to control the diurnal rhythm of TOC1 protein in regulation of the Arabidopsis circadian clock<ref name="ref4" />. <br />
*''Os-GI'' also has function to maintain bioactive GA level through the regulation of the GA-deactivating enzyme genes in rice, ''Os-GI'' represses major ''OsGA2ox'' genes in order to maintain a proper bioactive GA level, ''osgi-1'' plants showed a semi-dwarf phenotype with elevated ''OsGA2ox'' gene expression<br />
<ref name="ref4" /> [[File:limeijia1.png|right|thumb|150px|''WT rice VS. osgi-1 plants (from reference <ref name="ref4" />).'']].<br />
<br />
===Expression===<br />
*''GI'' encodes a nuclear protein that defines the proper amplitude and period length of circadian rhythms. ''GI'' expression is regulated by the circadian clock, with a peak in transcript levels at 8 to 10 h after dawn. The ''GI'' protein acts between the circadian oscillator and ''CO'' to promote flowering by increasing ''CO'' and ''FT'' mRNA abundance<ref name="ref1" />. The expression of the ''OsGI'' messenger RNA was circadian-controlled and that its temporal expression pattern was very similar to that of ''GI'' under both SD and LD conditions. Decreased ''OsGI'' expression led to early flowering under LD conditions and late flowering under SD conditions relative to the wild type, and that this early flowering phenotype under LD conditions was the opposite of that of the Arabidopsis ''gi'' mutants, confirming the opposite role of the ''GI'' gene in flowering under LD conditions in rice and Arabidopsis<ref name="ref5" />.<br />
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===Mutation===<br />
*The single ''osgi-1'' mutation extended the time to flowering only under SD conditions. Under LD conditions the single ''hd1'' mutation induced early flowering, but ''osgi-1'' plants exhibited no phenotypic change<ref name="ref4" /><br />
<br />
===Evolution===<br />
* There is some evolutionary conservation between rice circadian clock–related genes and those associated with Arabidopsis circadian clocks, where the circadian clock consists of interlocked subloops, it is also possible that ''Os-GI'' is a member of a subloop in rice circadian clocks<ref name="ref4" />.But there is a divergence in the photoperiodic pathways of these two species arises from the molecular mechanisms controlling the expression of ''FT/Hd3a'' by ''CO/Hd1(Se1)''<ref name="ref5" />(figure 3)<br />
<br />
==Labs working on this gene==<br />
*Photosynthesis and Photobiology Research Unit, National Institute of Agrobiological Sciences, 2-1-2 Kannondai, Tsukuba 305-8602, Japan<br />
*Laboratory of Plant Molecular Genetics, Nara Institute of Science and Technology, 8916-5 Takayama, Ikoma 630-0101, Japan<br />
*Plant Genomics Research Unit, National Institute of Agrobiological Sciences, 2-1-2 Kannondai, Tsukuba 305-8602, Japan<br />
<br />
==Knowledge Extension==<br />
*The rice orthologues of the two key regulators of flowering time in Arabidopsis, ''CO'' and ''FT'', were recently shown to be important in the photoperiodic control of flowering in rice. The rice ''Hd1(Se1)''gene, which is an orthologue of the Arabidopsis ''CO'' gene, was required for the suppression of flowering under LD conditions and for the promotion of flowering under SD conditions. Furthermore, the rice ''Hd3a'' gene, an orthologue of the Arabidopsis ''FT'' gene, was shown to be an activator of flowering in rice. It was previously demonstrated that ''CO'' controls the flowering time by integrating signals from the circadian clock and light to regulate ''FT'' expression in Arabidopsis<ref name="ref5" />.Unlike Arabidopsis ''FT'', expression of ''Hd3a'' mRNA is more or less opposite to that of ''Hd1'' under inductive short-day conditions<ref name="ref6" />.<br />
<br />
==References==<br />
<references><br />
<ref name="ref1">Song Lim Kim, Shinyoung Lee, Hyo Jung Kim, Hong Gil Nam, Gynheung An（2007）''OsMADS51'' is a short-day flowering promoter that functions upstream of ''Ehd1'', ''OsMADS14'', and ''Hd3a''.''Plant Physiology'' 145(4): 1484-1494.</ref><br />
<ref name="ref2">Hironori Itoh, Takeshi Izawa (2011) A study of phytohormone biosynthetic gene expression using a circadian clock-related mutant in rice. ''Plant Signaling & Behavior'' 6(12): 1932-1936.</ref> <br />
<ref name="ref3">Ryousuke Hayama, Takeshi Izawa, Ko Shimamoto (2002) Isolation of rice genes possibly involved in the photoperiodic control of flowering by a<br />
fluorescent differential display method. ''Plant and Cell Physiology'' 43(5): 494-504.</ref> <br />
<ref name="ref4">Takeshi Izawa, Motohiro Mihara, Yuji Suzuki, Meenu Gupta, Hironori Itoh, et al. (2011) ''Os-GIGANTEA'' confers robust diurnal rhythms on the global transcriptome of rice in the field. ''The Plant Cell'' 23(5): 1741-1755.</ref> <br />
<ref name="ref5">Ryosuke Hayama, Shuji Yokoi, Shojiro Tamaki, Masahiro Yano, Ko Shimamoto (2003) Adaptation of photoperiodic control pathways produces short-day flowering in rice. ''Nature'' 422(6933): 719-722.</ref> <br />
<ref name="ref6">Bong-Soo Shin, Jeong-Hyun Lee, Jeong-Hwan Lee, Hyun-Joo Jeong, Choong-Hyo Yun, Jeong-Kook Kim (2004) Circadian regulation of rice (''Oryza sativa L.'') ''CONSTANS-like'' gene transcripts. ''Molecules and Cells'' 17(1): 10-16.</ref><br />
<br />
==Structured Information==<br />
{{JaponicaGene|<br />
GeneName = Os01g0182600|<br />
Description = GIGANTEA protein|<br />
Version = NM_001048755.1 GI:115434923 GeneID:4325329|<br />
Length = 9083 bp|<br />
Definition = Oryza sativa Japonica Group Os01g0182600, complete gene.|<br />
Source = Oryza sativa Japonica Group<br />
<br />
ORGANISM Oryza sativa Japonica Group<br />
Eukaryota; Viridiplantae; Streptophyta; Embryophyta; Tracheophyta;<br />
Spermatophyta; Magnoliophyta; Liliopsida; Poales; Poaceae; BEP<br />
clade; Ehrhartoideae; Oryzeae; Oryza.<br />
|<br />
Chromosome = [[:category:Japonica Chromosome 1|Chromosome 1]]|<br />
AP = Chromosome 1:4328362..4337444|<br />
CDS = 4328725..4329030,4329123..4329215,4329333..4329551,4329752..4329820,4329925..4331421<br>,4332220..4332429,4332563..4332713,4332791..4333031,4333898..4333956<br>,4334379..4334653,4334813..4334955,4335034..4335088,4335181..4335258<br>,4335351..4335437|<br />
GCID = <gbrowseImage1><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage1>|<br />
GSID = <gbrowseImage2><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage2>|<br />
CDNA = <cdnaseq>atgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactgcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcatagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcggcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctga</cdnaseq>|<br />
AA = <aaseq>MSASNEKWIDGLQFSSLFWPPPQDSQQKQAQILAYVEYFGQFTA DSEQFPEDIAQLIQSCYPSKEKRLVDEVLATFVLHHPEHGHAVVHPILSRIIDGTLSY DRNGFPFMSFISLFSHTSEKEYSEQWALACGEILRVLTHYNRPIFKVDHQHSEAECSS TSDQASSCESMEKRANGSPRNEPDRKPLRPLSPWITDILLAAPLGIRSDYFRWCGGVM GKYAAGGELKPPTTAYSRGSGKHPQLMPSTPRWAVANGAGVILSVCDEEVARYETANL TAAAVPALLLPPPTTPLDEHLVAGLPPLEPYARLFHRYYAIATPSATQRLLFGLLEAP PSWAPDALDAAVQLVELLRAAEDYDSGMRLPKNWMHLHFLRAIGTAMSMRAGIAADTS AALLFRILSQPTLLFPPLRHAEGVELHHEPLGGYVSSYKRQLEVPASEATIDATAQGI ASMLCAHGPDVEWRICTIWEAAYGLLPLSSSAVDLPEIVVAAPLQPPTLSWSLYLPLL KVFEYLPRGSPSEACLMRIFVATVEAILRRTFPSETSEQSRKPRSQSKNLAVAELRTM IHSLFVESCASMDLASRLLFVVLTVCVSHQALPGGSKRPTGSDNHSSEEVTNDSRLTN GRNRCKKRQGPVATFDSYVLAAVCALSCELQLFPFISKNGNHSNLKDSIKIVIPGKTT GISNELHNSISSAILHTRRILGILEALFSLKPSSVGTSWSYSSNEIVAAAMVAAHVSE LFRRSRPCLNALSALKQCKWDAEISTRASSLYHLIDLHGKTVTSIVNKAEPLEAHLTL TPVKKDEPPIEEKNINSSDGGALEKKDASRSHRKNGFARPLLKCAEDVILNGDVASTS GKAIASLQVEASDLANFLTMDRNGGYRGSQTLLRSVLSEKQELCFSVVSLLWQKLIAS PEMQMSAESTSAHQGWRKVVDALCDIVSASPTKASAAIVLQAEKDLQPWIARDDEQGQ KMWRVNQRIVKLIAELMRNHDSPEALVILASASDLLLRATDGMLVDGEACTLPQLELL EVTARAVHLIVEWGDSGVSVADGLSNLLKCRLSTTIRCLSHPSAHVRALSMSVLRDIL NSGQINSSKLIQGEHRNGIQSPTYQCLAASIINWQADVERCIEWEAHSRRATGLTLAF LTAAAKELGCPLTC</aaseq>|<br />
DNA = <dnaseqindica>8415..8720#8230..8322#7894..8112#7625..7693#6024..7520#5016..5225#4732..4882#4414..4654#3489..3547#2792..3066#2490..2632#2357..2411#2187..2264#2008..2094#ccgagccgtttctctccctctctctctcctcctccttcccctgcgcctcctccaccgcccgtgtcgccgccgcgcgcgtctatcccctcgccgccgccgccgtctacgccgcgtgaggtccggattggttgaaaggctgatcgaggtaatgcttttttatttttcccgtgcggttttagtttttctgtgctgtgcttggtggttttttctttaaaactatttttaggctggttttgatgatggcctttctaaggttgtcgtgtatggggggcgaggtggttcgggtggatttgtggtgttatgttgaggggatttttgtttggggggagagagttttggtggtagatgcggttgggattttgaagacgtgaaagggaaattagttttgaaatttgacggcgaggttgagctgattagtattgtaataattcgtggcatcggatagatcaacgcgagatggatgcttattactgcgaatttcaggaggtttttgtggtttttttaggcgagtagcgttgattctagctggattttgctgcttcgtggaggcgtttagatgtggtgcatacggctgtggatgtaatttgtgcggggataattactgaaaattaattcgcagtaagctagctctgctttaatattttttcccctgaaacagtaacaagggatcccttcctcatttcgttaacagtggaagtgaatacagacaccaaccgatgagcagagacacctgtaattaaacaccaaaagaagaacctgttttgttcctatatattttccatttctatgctttaggtaatatactttctggttcggtgtcatagtacccgaaatagaatctttgcttgccaaaaaaagaaagaataacaagatagaacaacatcattaaattctgtcttagacaatacatttataatcgaccaacaaaaacagaccgatgagtgacgtggaataacaaatgaagtataaatgtcacgaaaaacacatctattgattttgtgtgataagccataggagcctacgaggaaccaagtgaatggttctgtacaccacatccattagcattaccataaatttgcctatttctatgatgccttccttaacatatttatcgacatgtgagcttgcaccataactaaatgttatggtaaactttaccttataaatgattgcaacatgtaccttttcatttcgataattaaaaacgagaaagcttccacctttttcctagatatattttttcgtttgattatgaatttagttgtacttgacaataaaataatgcgcttgtaagattatgcaattttcgttgcattacctccctctcttttcatttgtcagtttttttctactccctccgtttcatattttaagtcgtttgacttttttcctagtcagactttgttaagtttgacaaagtttatagtaaaatttagcaatatctaaaatattaaattggtttcatcaaatctaacattgaatatattttgataatatgttttttttgtgttgaaaatacttctatatttttctataaacttagtcaaacttggaagaaatttcagtaagaaaaaagtcaaacggcttataatatgaaacggagggagtaaatttttttttgcattaattaaaattcatgtattttcctccttttttggtttttgttgtgcgatatattacagtgtagagactctgaagctcatatggttggcattggttgatatgataacgtgtcacgttgtgagatgcagacccccgttatgggggttgattgaatgaagtggatagcactcttataactttcttgctcgaagtttcttgataccaaggcaagtagcttataagaagatttggactttggtggcataaatggatgaactaactggcggcttcaacgtggtgttgattacaggtgcttttattgctaactggagcagtttgtgtagtttctttctttctctttgagaattctctcagtttcttgggcaggctcttgagctaccaagtatgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggtaacattattcctatttgattcttgttagaaattctatatgttgaacatacttgcctcaatgggaaacttggttgcgtctcctctctgaaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcaggtactgtatctctaagcatgtttagtatcttctgtgataaaatctttctccagtgtgttaacttttgcattctttttcttgcatgcttgcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattaggtaatttgatccagtgatcaatacatgttccacaaacaattgaagaaagagtttgacatatttttctcatatttccagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgaggtaaggcccctactctaatggttagttccaatttattggttcatttctctgtgtttgataagattttgcctctgctgtgcatttcaatattatatctaaactgtgctgcttatagcttatcaaactactcttgctcataaagatgcaacttttatgcagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgagtcataataaatgataacatattattgttctgtggactgaaaatctgaaatcctcctttcttgagaaatacaacctcaattttatggtgatattgacaagaaggaagagcgcatataatttaaagttaaaacacggaaaataatagaagtatgtgagcagaacgcaggagcaaggggtggaactcataacaagttcaaacttcatgcatgttcataattccccaatgcaagtatggcaagaaaaatagacaacttttaaaagtatgctaatggaactgcatgcattgtgtaaacttgcttttgtcaagcatgttggaacatactatatatttgctgctgatgatgaaaaatttgcttgagtactcttctattgctaatggacgcttaccattccctgttttgtcttcaaatgcatcaggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactggttggtaaaatcagcaatggaattttctatcgtgaattttttataattattgtaacattaattactcccatgtctttttattctgttgagataccatgtgacaatgtctacaattgcatattttgcattctttggtagatcagttcccttcttaagatccccaaagggaaactggagtatcaataacttactccctcaaataaaaaaaaagtgatactttaggcataaacacattcttcaaaacaaaactttgaatagcagtttcttttgtaatatattaataaagcatggaaaaattataatgtgaagaatgtacattcagggacaaatatactcatcatttttgcatggttgatcttaatgtccatagagagattgatcagaccaaaagtttatttttgacaggagtgagtatggattataataggagtgaatggcaatatgctgtgaaagtgagcattttatatttgtttactactagcatttggtgacaaagcaaatgatgtatattgtaaatcaaagtaaacaagtagaacatgtcctgtggcgacaagacataacctacaaattaaagttagcctgattgtttattttactgggagcacattttttctagattctagagtacatattattaaaaaaggtgtgatatgtttgcaaattactttttaatggaagtttgtaaattactttgaaacttgcattcagtttttttgcttcttaaaaaacatagtgcatatgggctttatgagctcatactatgcatgtttttctgcattggagtgggacctttatctttttcttgcaactcggatgctaatgcttattataaacacaagttactaacatgtgcttgggtcattcagcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcataggtaagctcttatgatcctgtccactgttgtaccatgtgtactttccaaatgattgctaacatgctatttttcgtcagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcgggtatatcatacatctctctggctgttgtttcagtgctatcaatcaagatgacacatcatgcaatattatcataacagcacattatgtgttttgaatattttattgattgtcatagctttgttttatgttttagcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcaggtatgcagtgtttcctgcacctacttgctacctattctgtatttgatgtattcaatttcctttttatcagttttctcgaaatatatgctaataggcttgtcactaaacggctacattatccagactctgttctgaatttttttttcattaagtactgttagctgccagattacatcatcatgactaaagtagtttagctacaatattgcaaagctagatttgtctagcctctattgttttgaatagaaaaggcaggtatgcagtgtttcctgcacctacttgctacctattctgtatttgatgtattcaatttcctttttatcagttttctcgaaatatatgctaataggcttgtcactaaacggctacattatccagactctgttctgaatttttttttcattaagtactgttagctgccagattacatcatcatgactaaagtagtttagctacaatattgcaaagctagatttgtctagcctctattgttttgaatagaaaattaaatcttttggtcatttttttgtaaccttttaggagtctctgtatgcttgtcgtcaacatgctatgataagtttatatcagttcttttggagctatgatgtattcttgtatattttatgactgctaggatattatgttaaaactgttttaaaggtatcatcaaacaagttaatatagatattgctacctaccaaagttggaatttgattatgtcgtaaaatatatgcaccctacatgttttagcttcatattgacttatattttcatatgtgctccttcaactgcacaaagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggtatggtcattcctgcttataagataccagaataatgctatgaagaccgtccttggttgtcagcattcttatacacaatgataatctttgtttttcatgggtaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggtcaagatattgttattgttatatttacatacttgttatcatatttaaaagcttcatattgtatgtacctttgaaaaaaaaagttccatactgtagcttcttttgttctcatttcttggtttcaaagcatcagcatcattaccgatataaaatatacccatttcctcagtcatcttccccgactcttgtatctttcacaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagaggtaacaccttcattcatgaggtccttaggtgcaaatcccagtgaagaataatcagttctactttgtatggttcatctcgcttactgaccggttcatctcgggatcaattacgagcagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaaggcatgtcctcttctctgatctcactcacgttcatgccaaacttactcatcatcatccatgatccatcatcctgactgaatcttgatatgcagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctgacaaggccacaccatgactgtcactgcaagcagctgctgtaggatcagcgagtaggaaaggatgacttgccggccagtttcctgagatgtgatttaagactgatattagctgatgttcccaagcatgatacggggcttgctcccaagatgtgatttttactttacttttggttaatgtgctaccgctgacattagatgttcaagcatatgaaaactgcttgtgctgtaacttgtatgtctgtaacgtaaaaatctttccatatggtaaccggggaagcagaggtatatccttttctgctgttgtatctatttccagtttcctgctgaacttgcaaatgaatattcagcaatttccgttttgctc</dnaseqindica>|<br />
Link = [http://www.ncbi.nlm.nih.gov/nuccore/NM_001048755.1 RefSeq:Os01g0182600]|<br />
}}<br />
[[Category:Genes]]<br />
[[Category:Japonica mRNA]]<br />
[[Category:Oryza Sativa Japonica Group]]<br />
[[Category:Japonica Genes]]<br />
[[Category:Japonica Chromosome 1]]<br />
[[Category:Chromosome 1]]</div>Mikahttps://ngdc.cncb.ac.cn/ricewiki/index.php?title=File:Mika1.png&diff=183802File:Mika1.png2014-06-11T01:40:32Z<p>Mika: WT rice and osgi-1plants grown under natural field conditions and osgi-1 plant exhibits semi-dwarf phenotype</p>
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<div>WT rice and osgi-1plants grown under natural field conditions and osgi-1 plant exhibits semi-dwarf phenotype</div>Mikahttps://ngdc.cncb.ac.cn/ricewiki/index.php?title=Os01g0182600&diff=183801Os01g01826002014-06-11T01:38:48Z<p>Mika: </p>
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<div>The rice gene of ''Gigantea(GI)'' is solo ortholog of ''GI'' in the Arabidopsis circadian system and related to photoperiodic control of flowering.<br />
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==Annotated Information==<br />
===Function===<br />
*''OsGI'',a ''GI'' ortholog, regulates flowering time in response to photoperiodic conditions. Mutations in this gene delay flowering in inductive SDs (short-days),whereas flowering is marginally induced under LDs(long-days).In comparison, over expression of ''OsGI'' in transgenic rice causes late flowering in both SDs and LDs.In addition, ''OsGI'' has function to regulate ''Hd1'', a gene that also shows circadian rhythm<ref name="ref1" />. ''Os-GI'' affects gene expression of approximately 75% of genes among 27,201 genes and is required for strong amplitudes and proper phase-setting of global gene expression under natural field conditions<ref name="ref2" />. <br />
*The cDNA of ''OsGI'' firstly found in ''se5'' mutant that is completely photoperiod-insensitive, the expression patterns of rice and Arabidopsis ''GI'' gene are both photoperiodically regulated and are very similar under LD and SD conditions(FIG1)<ref name="ref3" />. ''GI'' is required for phase-dependent ''CO''transcription through the interaction with FKF1, which leads to cycling DNA binding with one finger (CDF) degradation and consequent derepression of CO. In addition, it has been known that GI protein interacts with ZEITLUPE (ZTL) F-box protein to control the diurnal rhythm of TOC1 protein in regulation of the Arabidopsis circadian clock<ref name="ref4" />. <br />
*''Os-GI'' also has function to maintain bioactive GA level through the regulation of the GA-deactivating enzyme genes in rice, ''Os-GI'' represses major ''OsGA2ox'' genes in order to maintain a proper bioactive GA level, ''osgi-1'' plants showed a semi-dwarf phenotype with elevated ''OsGA2ox'' gene expression [[File:limeijia1.png|right|thumb|150px|''WT rice VS. osgi-1 plants (from reference <ref name="ref4" />).'']].<br />
<br />
===Expression===<br />
*''GI'' encodes a nuclear protein that defines the proper amplitude and period length of circadian rhythms. ''GI'' expression is regulated by the circadian clock, with a peak in transcript levels at 8 to 10 h after dawn. The ''GI'' protein acts between the circadian oscillator and ''CO'' to promote flowering by increasing ''CO'' and ''FT'' mRNA abundance<ref name="ref1" />. The expression of the ''OsGI'' messenger RNA was circadian-controlled and that its temporal expression pattern was very similar to that of ''GI'' under both SD and LD conditions. Decreased ''OsGI'' expression led to early flowering under LD conditions and late flowering under SD conditions relative to the wild type, and that this early flowering phenotype under LD conditions was the opposite of that of the Arabidopsis ''gi'' mutants, confirming the opposite role of the ''GI'' gene in flowering under LD conditions in rice and Arabidopsis<ref name="ref5" />.<br />
<br />
===Mutation===<br />
*The single ''osgi-1'' mutation extended the time to flowering only under SD conditions. Under LD conditions the single ''hd1'' mutation induced early flowering, but ''osgi-1'' plants exhibited no phenotypic change<ref name="ref4" /><br />
<br />
===Evolution===<br />
* There is some evolutionary conservation between rice circadian clock–related genes and those associated with Arabidopsis circadian clocks, where the circadian clock consists of interlocked subloops, it is also possible that ''Os-GI'' is a member of a subloop in rice circadian clocks<ref name="ref4" />.But there is a divergence in the photoperiodic pathways of these two species arises from the molecular mechanisms controlling the expression of ''FT/Hd3a'' by ''CO/Hd1(Se1)''<ref name="ref5" />(figure 3)<br />
<br />
==Labs working on this gene==<br />
*Photosynthesis and Photobiology Research Unit, National Institute of Agrobiological Sciences, 2-1-2 Kannondai, Tsukuba 305-8602, Japan<br />
*Laboratory of Plant Molecular Genetics, Nara Institute of Science and Technology, 8916-5 Takayama, Ikoma 630-0101, Japan<br />
*Plant Genomics Research Unit, National Institute of Agrobiological Sciences, 2-1-2 Kannondai, Tsukuba 305-8602, Japan<br />
<br />
==Knowledge Extension==<br />
*The rice orthologues of the two key regulators of flowering time in Arabidopsis, ''CO'' and ''FT'', were recently shown to be important in the photoperiodic control of flowering in rice. The rice ''Hd1(Se1)''gene, which is an orthologue of the Arabidopsis ''CO'' gene, was required for the suppression of flowering under LD conditions and for the promotion of flowering under SD conditions. Furthermore, the rice ''Hd3a'' gene, an orthologue of the Arabidopsis ''FT'' gene, was shown to be an activator of flowering in rice. It was previously demonstrated that ''CO'' controls the flowering time by integrating signals from the circadian clock and light to regulate ''FT'' expression in Arabidopsis<ref name="ref5" />.Unlike Arabidopsis ''FT'', expression of ''Hd3a'' mRNA is more or less opposite to that of ''Hd1'' under inductive short-day conditions<ref name="ref6" />.<br />
<br />
==References==<br />
<references><br />
<ref name="ref1">Song Lim Kim, Shinyoung Lee, Hyo Jung Kim, Hong Gil Nam, Gynheung An（2007）''OsMADS51'' is a short-day flowering promoter that functions upstream of ''Ehd1'', ''OsMADS14'', and ''Hd3a''.''Plant Physiology'' 145(4): 1484-1494.</ref><br />
<ref name="ref2">Hironori Itoh, Takeshi Izawa (2011) A study of phytohormone biosynthetic gene expression using a circadian clock-related mutant in rice. ''Plant Signaling & Behavior'' 6(12): 1932-1936.</ref> <br />
<ref name="ref3">Ryousuke Hayama, Takeshi Izawa, Ko Shimamoto (2002) Isolation of rice genes possibly involved in the photoperiodic control of flowering by a<br />
fluorescent differential display method. ''Plant and Cell Physiology'' 43(5): 494-504.</ref> <br />
<ref name="ref4">Takeshi Izawa, Motohiro Mihara, Yuji Suzuki, Meenu Gupta, Hironori Itoh, et al. (2011) ''Os-GIGANTEA'' confers robust diurnal rhythms on the global transcriptome of rice in the field. ''The Plant Cell'' 23(5): 1741-1755.</ref> <br />
<ref name="ref5">Ryosuke Hayama, Shuji Yokoi, Shojiro Tamaki, Masahiro Yano, Ko Shimamoto (2003) Adaptation of photoperiodic control pathways produces short-day flowering in rice. ''Nature'' 422(6933): 719-722.</ref> <br />
<ref name="ref6">Bong-Soo Shin, Jeong-Hyun Lee, Jeong-Hwan Lee, Hyun-Joo Jeong, Choong-Hyo Yun, Jeong-Kook Kim (2004) Circadian regulation of rice (''Oryza sativa L.'') ''CONSTANS-like'' gene transcripts. ''Molecules and Cells'' 17(1): 10-16.</ref><br />
<br />
==Structured Information==<br />
{{JaponicaGene|<br />
GeneName = Os01g0182600|<br />
Description = GIGANTEA protein|<br />
Version = NM_001048755.1 GI:115434923 GeneID:4325329|<br />
Length = 9083 bp|<br />
Definition = Oryza sativa Japonica Group Os01g0182600, complete gene.|<br />
Source = Oryza sativa Japonica Group<br />
<br />
ORGANISM Oryza sativa Japonica Group<br />
Eukaryota; Viridiplantae; Streptophyta; Embryophyta; Tracheophyta;<br />
Spermatophyta; Magnoliophyta; Liliopsida; Poales; Poaceae; BEP<br />
clade; Ehrhartoideae; Oryzeae; Oryza.<br />
|<br />
Chromosome = [[:category:Japonica Chromosome 1|Chromosome 1]]|<br />
AP = Chromosome 1:4328362..4337444|<br />
CDS = 4328725..4329030,4329123..4329215,4329333..4329551,4329752..4329820,4329925..4331421<br>,4332220..4332429,4332563..4332713,4332791..4333031,4333898..4333956<br>,4334379..4334653,4334813..4334955,4335034..4335088,4335181..4335258<br>,4335351..4335437|<br />
GCID = <gbrowseImage1><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage1>|<br />
GSID = <gbrowseImage2><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage2>|<br />
CDNA = <cdnaseq>atgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactgcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcatagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcggcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctga</cdnaseq>|<br />
AA = <aaseq>MSASNEKWIDGLQFSSLFWPPPQDSQQKQAQILAYVEYFGQFTA DSEQFPEDIAQLIQSCYPSKEKRLVDEVLATFVLHHPEHGHAVVHPILSRIIDGTLSY DRNGFPFMSFISLFSHTSEKEYSEQWALACGEILRVLTHYNRPIFKVDHQHSEAECSS TSDQASSCESMEKRANGSPRNEPDRKPLRPLSPWITDILLAAPLGIRSDYFRWCGGVM GKYAAGGELKPPTTAYSRGSGKHPQLMPSTPRWAVANGAGVILSVCDEEVARYETANL TAAAVPALLLPPPTTPLDEHLVAGLPPLEPYARLFHRYYAIATPSATQRLLFGLLEAP PSWAPDALDAAVQLVELLRAAEDYDSGMRLPKNWMHLHFLRAIGTAMSMRAGIAADTS AALLFRILSQPTLLFPPLRHAEGVELHHEPLGGYVSSYKRQLEVPASEATIDATAQGI ASMLCAHGPDVEWRICTIWEAAYGLLPLSSSAVDLPEIVVAAPLQPPTLSWSLYLPLL KVFEYLPRGSPSEACLMRIFVATVEAILRRTFPSETSEQSRKPRSQSKNLAVAELRTM IHSLFVESCASMDLASRLLFVVLTVCVSHQALPGGSKRPTGSDNHSSEEVTNDSRLTN GRNRCKKRQGPVATFDSYVLAAVCALSCELQLFPFISKNGNHSNLKDSIKIVIPGKTT GISNELHNSISSAILHTRRILGILEALFSLKPSSVGTSWSYSSNEIVAAAMVAAHVSE LFRRSRPCLNALSALKQCKWDAEISTRASSLYHLIDLHGKTVTSIVNKAEPLEAHLTL TPVKKDEPPIEEKNINSSDGGALEKKDASRSHRKNGFARPLLKCAEDVILNGDVASTS GKAIASLQVEASDLANFLTMDRNGGYRGSQTLLRSVLSEKQELCFSVVSLLWQKLIAS PEMQMSAESTSAHQGWRKVVDALCDIVSASPTKASAAIVLQAEKDLQPWIARDDEQGQ KMWRVNQRIVKLIAELMRNHDSPEALVILASASDLLLRATDGMLVDGEACTLPQLELL EVTARAVHLIVEWGDSGVSVADGLSNLLKCRLSTTIRCLSHPSAHVRALSMSVLRDIL NSGQINSSKLIQGEHRNGIQSPTYQCLAASIINWQADVERCIEWEAHSRRATGLTLAF LTAAAKELGCPLTC</aaseq>|<br />
DNA = <dnaseqindica>8415..8720#8230..8322#7894..8112#7625..7693#6024..7520#5016..5225#4732..4882#4414..4654#3489..3547#2792..3066#2490..2632#2357..2411#2187..2264#2008..2094#ccgagccgtttctctccctctctctctcctcctccttcccctgcgcctcctccaccgcccgtgtcgccgccgcgcgcgtctatcccctcgccgccgccgccgtctacgccgcgtgaggtccggattggttgaaaggctgatcgaggtaatgcttttttatttttcccgtgcggttttagtttttctgtgctgtgcttggtggttttttctttaaaactatttttaggctggttttgatgatggcctttctaaggttgtcgtgtatggggggcgaggtggttcgggtggatttgtggtgttatgttgaggggatttttgtttggggggagagagttttggtggtagatgcggttgggattttgaagacgtgaaagggaaattagttttgaaatttgacggcgaggttgagctgattagtattgtaataattcgtggcatcggatagatcaacgcgagatggatgcttattactgcgaatttcaggaggtttttgtggtttttttaggcgagtagcgttgattctagctggattttgctgcttcgtggaggcgtttagatgtggtgcatacggctgtggatgtaatttgtgcggggataattactgaaaattaattcgcagtaagctagctctgctttaatattttttcccctgaaacagtaacaagggatcccttcctcatttcgttaacagtggaagtgaatacagacaccaaccgatgagcagagacacctgtaattaaacaccaaaagaagaacctgttttgttcctatatattttccatttctatgctttaggtaatatactttctggttcggtgtcatagtacccgaaatagaatctttgcttgccaaaaaaagaaagaataacaagatagaacaacatcattaaattctgtcttagacaatacatttataatcgaccaacaaaaacagaccgatgagtgacgtggaataacaaatgaagtataaatgtcacgaaaaacacatctattgattttgtgtgataagccataggagcctacgaggaaccaagtgaatggttctgtacaccacatccattagcattaccataaatttgcctatttctatgatgccttccttaacatatttatcgacatgtgagcttgcaccataactaaatgttatggtaaactttaccttataaatgattgcaacatgtaccttttcatttcgataattaaaaacgagaaagcttccacctttttcctagatatattttttcgtttgattatgaatttagttgtacttgacaataaaataatgcgcttgtaagattatgcaattttcgttgcattacctccctctcttttcatttgtcagtttttttctactccctccgtttcatattttaagtcgtttgacttttttcctagtcagactttgttaagtttgacaaagtttatagtaaaatttagcaatatctaaaatattaaattggtttcatcaaatctaacattgaatatattttgataatatgttttttttgtgttgaaaatacttctatatttttctataaacttagtcaaacttggaagaaatttcagtaagaaaaaagtcaaacggcttataatatgaaacggagggagtaaatttttttttgcattaattaaaattcatgtattttcctccttttttggtttttgttgtgcgatatattacagtgtagagactctgaagctcatatggttggcattggttgatatgataacgtgtcacgttgtgagatgcagacccccgttatgggggttgattgaatgaagtggatagcactcttataactttcttgctcgaagtttcttgataccaaggcaagtagcttataagaagatttggactttggtggcataaatggatgaactaactggcggcttcaacgtggtgttgattacaggtgcttttattgctaactggagcagtttgtgtagtttctttctttctctttgagaattctctcagtttcttgggcaggctcttgagctaccaagtatgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggtaacattattcctatttgattcttgttagaaattctatatgttgaacatacttgcctcaatgggaaacttggttgcgtctcctctctgaaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcaggtactgtatctctaagcatgtttagtatcttctgtgataaaatctttctccagtgtgttaacttttgcattctttttcttgcatgcttgcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattaggtaatttgatccagtgatcaatacatgttccacaaacaattgaagaaagagtttgacatatttttctcatatttccagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgaggtaaggcccctactctaatggttagttccaatttattggttcatttctctgtgtttgataagattttgcctctgctgtgcatttcaatattatatctaaactgtgctgcttatagcttatcaaactactcttgctcataaagatgcaacttttatgcagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgagtcataataaatgataacatattattgttctgtggactgaaaatctgaaatcctcctttcttgagaaatacaacctcaattttatggtgatattgacaagaaggaagagcgcatataatttaaagttaaaacacggaaaataatagaagtatgtgagcagaacgcaggagcaaggggtggaactcataacaagttcaaacttcatgcatgttcataattccccaatgcaagtatggcaagaaaaatagacaacttttaaaagtatgctaatggaactgcatgcattgtgtaaacttgcttttgtcaagcatgttggaacatactatatatttgctgctgatgatgaaaaatttgcttgagtactcttctattgctaatggacgcttaccattccctgttttgtcttcaaatgcatcaggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactggttggtaaaatcagcaatggaattttctatcgtgaattttttataattattgtaacattaattactcccatgtctttttattctgttgagataccatgtgacaatgtctacaattgcatattttgcattctttggtagatcagttcccttcttaagatccccaaagggaaactggagtatcaataacttactccctcaaataaaaaaaaagtgatactttaggcataaacacattcttcaaaacaaaactttgaatagcagtttcttttgtaatatattaataaagcatggaaaaattataatgtgaagaatgtacattcagggacaaatatactcatcatttttgcatggttgatcttaatgtccatagagagattgatcagaccaaaagtttatttttgacaggagtgagtatggattataataggagtgaatggcaatatgctgtgaaagtgagcattttatatttgtttactactagcatttggtgacaaagcaaatgatgtatattgtaaatcaaagtaaacaagtagaacatgtcctgtggcgacaagacataacctacaaattaaagttagcctgattgtttattttactgggagcacattttttctagattctagagtacatattattaaaaaaggtgtgatatgtttgcaaattactttttaatggaagtttgtaaattactttgaaacttgcattcagtttttttgcttcttaaaaaacatagtgcatatgggctttatgagctcatactatgcatgtttttctgcattggagtgggacctttatctttttcttgcaactcggatgctaatgcttattataaacacaagttactaacatgtgcttgggtcattcagcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcataggtaagctcttatgatcctgtccactgttgtaccatgtgtactttccaaatgattgctaacatgctatttttcgtcagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtac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/>
Link = [http://www.ncbi.nlm.nih.gov/nuccore/NM_001048755.1 RefSeq:Os01g0182600]|<br />
}}<br />
[[Category:Genes]]<br />
[[Category:Japonica mRNA]]<br />
[[Category:Oryza Sativa Japonica Group]]<br />
[[Category:Japonica Genes]]<br />
[[Category:Japonica Chromosome 1]]<br />
[[Category:Chromosome 1]]</div>Mikahttps://ngdc.cncb.ac.cn/ricewiki/index.php?title=File:%E6%9D%8E%E7%BE%8E%E4%BD%B31.png&diff=183800File:李美佳1.png2014-06-11T01:28:11Z<p>Mika: WT and osgi-1 plants grown under natural field conditions and osgi-1 plant shows Semi-dwarf phenotype</p>
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<div>WT and osgi-1 plants grown under natural field conditions and osgi-1 plant shows Semi-dwarf phenotype</div>Mikahttps://ngdc.cncb.ac.cn/ricewiki/index.php?title=Os01g0182600&diff=183799Os01g01826002014-06-11T01:20:29Z<p>Mika: /* Labs working on this gene */</p>
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<div>The rice gene of ''Gigantea(GI)'' is solo ortholog of ''GI'' in the Arabidopsis circadian system and related to photoperiodic control of flowering.<br />
<br />
==Annotated Information==<br />
===Function===<br />
*''OsGI'',a ''GI'' ortholog, regulates flowering time in response to photoperiodic conditions. Mutations in this gene delay flowering in inductive SDs (short-days),whereas flowering is marginally induced under LDs(long-days).In comparison, over expression of ''OsGI'' in transgenic rice causes late flowering in both SDs and LDs.In addition, ''OsGI'' has function to regulate ''Hd1'', a gene that also shows circadian rhythm<ref name="ref1" />. ''Os-GI'' affects gene expression of approximately 75% of genes among 27,201 genes and is required for strong amplitudes and proper phase-setting of global gene expression under natural field conditions<ref name="ref2" />. <br />
*The cDNA of ''OsGI'' firstly found in ''se5'' mutant that is completely photoperiod-insensitive, the expression patterns of rice and Arabidopsis ''GI'' gene are both photoperiodically regulated and are very similar under LD and SD conditions(FIG1)<ref name="ref3" />. ''GI'' is required for phase-dependent ''CO''transcription through the interaction with FKF1, which leads to cycling DNA binding with one finger (CDF) degradation and consequent derepression of CO. In addition, it has been known that GI protein interacts with ZEITLUPE (ZTL) F-box protein to control the diurnal rhythm of TOC1 protein in regulation of the Arabidopsis circadian clock<ref name="ref4" />. <br />
*''Os-GI'' also has function to maintain bioactive GA level through the regulation of the GA-deactivating enzyme genes in rice, ''Os-GI'' represses major ''OsGA2ox'' genes in order to maintain a proper bioactive GA level, ''osgi-1'' plants showed a semi-dwarf phenotype with elevated ''OsGA2ox'' gene expression(FIG2) <ref name="ref4" />.<br />
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===Expression===<br />
*''GI'' encodes a nuclear protein that defines the proper amplitude and period length of circadian rhythms. ''GI'' expression is regulated by the circadian clock, with a peak in transcript levels at 8 to 10 h after dawn. The ''GI'' protein acts between the circadian oscillator and ''CO'' to promote flowering by increasing ''CO'' and ''FT'' mRNA abundance<ref name="ref1" />. The expression of the ''OsGI'' messenger RNA was circadian-controlled and that its temporal expression pattern was very similar to that of ''GI'' under both SD and LD conditions. Decreased ''OsGI'' expression led to early flowering under LD conditions and late flowering under SD conditions relative to the wild type, and that this early flowering phenotype under LD conditions was the opposite of that of the Arabidopsis ''gi'' mutants, confirming the opposite role of the ''GI'' gene in flowering under LD conditions in rice and Arabidopsis<ref name="ref5" />.<br />
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===Mutation===<br />
*The single ''osgi-1'' mutation extended the time to flowering only under SD conditions. Under LD conditions the single ''hd1'' mutation induced early flowering, but ''osgi-1'' plants exhibited no phenotypic change<ref name="ref4" /><br />
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===Evolution===<br />
* There is some evolutionary conservation between rice circadian clock–related genes and those associated with Arabidopsis circadian clocks, where the circadian clock consists of interlocked subloops, it is also possible that ''Os-GI'' is a member of a subloop in rice circadian clocks<ref name="ref4" />.But there is a divergence in the photoperiodic pathways of these two species arises from the molecular mechanisms controlling the expression of ''FT/Hd3a'' by ''CO/Hd1(Se1)''<ref name="ref5" />(figure 3)<br />
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==Labs working on this gene==<br />
*Photosynthesis and Photobiology Research Unit, National Institute of Agrobiological Sciences, 2-1-2 Kannondai, Tsukuba 305-8602, Japan<br />
*Laboratory of Plant Molecular Genetics, Nara Institute of Science and Technology, 8916-5 Takayama, Ikoma 630-0101, Japan<br />
*Plant Genomics Research Unit, National Institute of Agrobiological Sciences, 2-1-2 Kannondai, Tsukuba 305-8602, Japan<br />
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==Knowledge Extension==<br />
*The rice orthologues of the two key regulators of flowering time in Arabidopsis, ''CO'' and ''FT'', were recently shown to be important in the photoperiodic control of flowering in rice. The rice ''Hd1(Se1)''gene, which is an orthologue of the Arabidopsis ''CO'' gene, was required for the suppression of flowering under LD conditions and for the promotion of flowering under SD conditions. Furthermore, the rice ''Hd3a'' gene, an orthologue of the Arabidopsis ''FT'' gene, was shown to be an activator of flowering in rice. It was previously demonstrated that ''CO'' controls the flowering time by integrating signals from the circadian clock and light to regulate ''FT'' expression in Arabidopsis<ref name="ref5" />.Unlike Arabidopsis ''FT'', expression of ''Hd3a'' mRNA is more or less opposite to that of ''Hd1'' under inductive short-day conditions<ref name="ref6" />.<br />
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==References==<br />
<references><br />
<ref name="ref1">Song Lim Kim, Shinyoung Lee, Hyo Jung Kim, Hong Gil Nam, Gynheung An（2007）''OsMADS51'' is a short-day flowering promoter that functions upstream of ''Ehd1'', ''OsMADS14'', and ''Hd3a''.''Plant Physiology'' 145(4): 1484-1494.</ref><br />
<ref name="ref2">Hironori Itoh, Takeshi Izawa (2011) A study of phytohormone biosynthetic gene expression using a circadian clock-related mutant in rice. ''Plant Signaling & Behavior'' 6(12): 1932-1936.</ref> <br />
<ref name="ref3">Ryousuke Hayama, Takeshi Izawa, Ko Shimamoto (2002) Isolation of rice genes possibly involved in the photoperiodic control of flowering by a<br />
fluorescent differential display method. ''Plant and Cell Physiology'' 43(5): 494-504.</ref> <br />
<ref name="ref4">Takeshi Izawa, Motohiro Mihara, Yuji Suzuki, Meenu Gupta, Hironori Itoh, et al. (2011) ''Os-GIGANTEA'' confers robust diurnal rhythms on the global transcriptome of rice in the field. ''The Plant Cell'' 23(5): 1741-1755.</ref> <br />
<ref name="ref5">Ryosuke Hayama, Shuji Yokoi, Shojiro Tamaki, Masahiro Yano, Ko Shimamoto (2003) Adaptation of photoperiodic control pathways produces short-day flowering in rice. ''Nature'' 422(6933): 719-722.</ref> <br />
<ref name="ref6">Bong-Soo Shin, Jeong-Hyun Lee, Jeong-Hwan Lee, Hyun-Joo Jeong, Choong-Hyo Yun, Jeong-Kook Kim (2004) Circadian regulation of rice (''Oryza sativa L.'') ''CONSTANS-like'' gene transcripts. ''Molecules and Cells'' 17(1): 10-16.</ref><br />
<br />
==Structured Information==<br />
{{JaponicaGene|<br />
GeneName = Os01g0182600|<br />
Description = GIGANTEA protein|<br />
Version = NM_001048755.1 GI:115434923 GeneID:4325329|<br />
Length = 9083 bp|<br />
Definition = Oryza sativa Japonica Group Os01g0182600, complete gene.|<br />
Source = Oryza sativa Japonica Group<br />
<br />
ORGANISM Oryza sativa Japonica Group<br />
Eukaryota; Viridiplantae; Streptophyta; Embryophyta; Tracheophyta;<br />
Spermatophyta; Magnoliophyta; Liliopsida; Poales; Poaceae; BEP<br />
clade; Ehrhartoideae; Oryzeae; Oryza.<br />
|<br />
Chromosome = [[:category:Japonica Chromosome 1|Chromosome 1]]|<br />
AP = Chromosome 1:4328362..4337444|<br />
CDS = 4328725..4329030,4329123..4329215,4329333..4329551,4329752..4329820,4329925..4331421<br>,4332220..4332429,4332563..4332713,4332791..4333031,4333898..4333956<br>,4334379..4334653,4334813..4334955,4335034..4335088,4335181..4335258<br>,4335351..4335437|<br />
GCID = <gbrowseImage1><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage1>|<br />
GSID = <gbrowseImage2><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage2>|<br />
CDNA = <cdnaseq>atgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactgcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcatagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcggcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctga</cdnaseq>|<br />
AA = <aaseq>MSASNEKWIDGLQFSSLFWPPPQDSQQKQAQILAYVEYFGQFTA DSEQFPEDIAQLIQSCYPSKEKRLVDEVLATFVLHHPEHGHAVVHPILSRIIDGTLSY DRNGFPFMSFISLFSHTSEKEYSEQWALACGEILRVLTHYNRPIFKVDHQHSEAECSS TSDQASSCESMEKRANGSPRNEPDRKPLRPLSPWITDILLAAPLGIRSDYFRWCGGVM GKYAAGGELKPPTTAYSRGSGKHPQLMPSTPRWAVANGAGVILSVCDEEVARYETANL TAAAVPALLLPPPTTPLDEHLVAGLPPLEPYARLFHRYYAIATPSATQRLLFGLLEAP PSWAPDALDAAVQLVELLRAAEDYDSGMRLPKNWMHLHFLRAIGTAMSMRAGIAADTS AALLFRILSQPTLLFPPLRHAEGVELHHEPLGGYVSSYKRQLEVPASEATIDATAQGI ASMLCAHGPDVEWRICTIWEAAYGLLPLSSSAVDLPEIVVAAPLQPPTLSWSLYLPLL KVFEYLPRGSPSEACLMRIFVATVEAILRRTFPSETSEQSRKPRSQSKNLAVAELRTM IHSLFVESCASMDLASRLLFVVLTVCVSHQALPGGSKRPTGSDNHSSEEVTNDSRLTN GRNRCKKRQGPVATFDSYVLAAVCALSCELQLFPFISKNGNHSNLKDSIKIVIPGKTT GISNELHNSISSAILHTRRILGILEALFSLKPSSVGTSWSYSSNEIVAAAMVAAHVSE LFRRSRPCLNALSALKQCKWDAEISTRASSLYHLIDLHGKTVTSIVNKAEPLEAHLTL TPVKKDEPPIEEKNINSSDGGALEKKDASRSHRKNGFARPLLKCAEDVILNGDVASTS GKAIASLQVEASDLANFLTMDRNGGYRGSQTLLRSVLSEKQELCFSVVSLLWQKLIAS PEMQMSAESTSAHQGWRKVVDALCDIVSASPTKASAAIVLQAEKDLQPWIARDDEQGQ KMWRVNQRIVKLIAELMRNHDSPEALVILASASDLLLRATDGMLVDGEACTLPQLELL EVTARAVHLIVEWGDSGVSVADGLSNLLKCRLSTTIRCLSHPSAHVRALSMSVLRDIL NSGQINSSKLIQGEHRNGIQSPTYQCLAASIINWQADVERCIEWEAHSRRATGLTLAF LTAAAKELGCPLTC</aaseq>|<br />
DNA = <dnaseqindica>8415..8720#8230..8322#7894..8112#7625..7693#6024..7520#5016..5225#4732..4882#4414..4654#3489..3547#2792..3066#2490..2632#2357..2411#2187..2264#2008..2094#ccgagccgtttctctccctctctctctcctcctccttcccctgcgcctcctccaccgcccgtgtcgccgccgcgcgcgtctatcccctcgccgccgccgccgtctacgccgcgtgaggtccggattggttgaaaggctgatcgaggtaatgcttttttatttttcccgtgcggttttagtttttctgtgctgtgcttggtggttttttctttaaaactatttttaggctggttttgatgatggcctttctaaggttgtcgtgtatggggggcgaggtggttcgggtggatttgtggtgttatgttgaggggatttttgtttggggggagagagttttggtggtagatgcggttgggattttgaagacgtgaaagggaaattagttttgaaatttgacggcgaggttgagctgattagtattgtaataattcgtggcatcggatagatcaacgcgagatggatgcttattactgcgaatttcaggaggtttttgtggtttttttaggcgagtagcgttgattctagctggattttgctgcttcgtggaggcgtttagatgtggtgcatacggctgtggatgtaatttgtgcggggataattactgaaaattaattcgcagtaagctagctctgctttaatattttttcccctgaaacagtaacaagggatcccttcctcatttcgttaacagtggaagtgaatacagacaccaaccgatgagcagagacacctgtaattaaacaccaaaagaagaacctgttttgttcctatatattttccatttctatgctttaggtaatatactttctggttcggtgtcatagtacccgaaatagaatctttgcttgccaaaaaaagaaagaataacaagatagaacaacatcattaaattctgtcttagacaatacatttataatcgaccaacaaaaacagaccgatgagtgacgtggaataacaaatgaagtataaatgtcacgaaaaacacatctattgattttgtgtgataagccataggagcctacgaggaaccaagtgaatggttctgtacaccacatccattagcattaccataaatttgcctatttctatgatgccttccttaacatatttatcgacatgtgagcttgcaccataactaaatgttatggtaaactttaccttataaatgattgcaacatgtaccttttcatttcgataattaaaaacgagaaagcttccacctttttcctagatatattttttcgtttgattatgaatttagttgtacttgacaataaaataatgcgcttgtaagattatgcaattttcgttgcattacctccctctcttttcatttgtcagtttttttctactccctccgtttcatattttaagtcgtttgacttttttcctagtcagactttgttaagtttgacaaagtttatagtaaaatttagcaatatctaaaatattaaattggtttcatcaaatctaacattgaatatattttgataatatgttttttttgtgttgaaaatacttctatatttttctataaacttagtcaaacttggaagaaatttcagtaagaaaaaagtcaaacggcttataatatgaaacggagggagtaaatttttttttgcattaattaaaattcatgtattttcctccttttttggtttttgttgtgcgatatattacagtgtagagactctgaagctcatatggttggcattggttgatatgataacgtgtcacgttgtgagatgcagacccccgttatgggggttgattgaatgaagtggatagcactcttataactttcttgctcgaagtttcttgataccaaggcaagtagcttataagaagatttggactttggtggcataaatggatgaactaactggcggcttcaacgtggtgttgattacaggtgcttttattgctaactggagcagtttgtgtagtttctttctttctctttgagaattctctcagtttcttgggcaggctcttgagctaccaagtatgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggtaacattattcctatttgattcttgttagaaattctatatgttgaacatacttgcctcaatgggaaacttggttgcgtctcctctctgaaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcaggtactgtatctctaagcatgtttagtatcttctgtgataaaatctttctccagtgtgttaacttttgcattctttttcttgcatgcttgcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattaggtaatttgatccagtgatcaatacatgttccacaaacaattgaagaaagagtttgacatatttttctcatatttccagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgaggtaaggcccctactctaatggttagttccaatttattggttcatttctctgtgtttgataagattttgcctctgctgtgcatttcaatattatatctaaactgtgctgcttatagcttatcaaactactcttgctcataaagatgcaacttttatgcagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgagtcataataaatgataacatattattgttctgtggactgaaaatctgaaatcctcctttcttgagaaatacaacctcaattttatggtgatattgacaagaaggaagagcgcatataatttaaagttaaaacacggaaaataatagaagtatgtgagcagaacgcaggagcaaggggtggaactcataacaagttcaaacttcatgcatgttcataattccccaatgcaagtatggcaagaaaaatagacaacttttaaaagtatgctaatggaactgcatgcattgtgtaaacttgcttttgtcaagcatgttggaacatactatatatttgctgctgatgatgaaaaatttgcttgagtactcttctattgctaatggacgcttaccattccctgttttgtcttcaaatgcatcaggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactggttggtaaaatcagcaatggaattttctatcgtgaattttttataattattgtaacattaattactcccatgtctttttattctgttgagataccatgtgacaatgtctacaattgcatattttgcattctttggtagatcagttcccttcttaagatccccaaagggaaactggagtatcaataacttactccctcaaataaaaaaaaagtgatactttaggcataaacacattcttcaaaacaaaactttgaatagcagtttcttttgtaatatattaataaagcatggaaaaattataatgtgaagaatgtacattcagggacaaatatactcatcatttttgcatggttgatcttaatgtccatagagagattgatcagaccaaaagtttatttttgacaggagtgagtatggattataataggagtgaatggcaatatgctgtgaaagtgagcattttatatttgtttactactagcatttggtgacaaagcaaatgatgtatattgtaaatcaaagtaaacaagtagaacatgtcctgtggcgacaagacataacctacaaattaaagttagcctgattgtttattttactgggagcacattttttctagattctagagtacatattattaaaaaaggtgtgatatgtttgcaaattactttttaatggaagtttgtaaattactttgaaacttgcattcagtttttttgcttcttaaaaaacatagtgcatatgggctttatgagctcatactatgcatgtttttctgcattggagtgggacctttatctttttcttgcaactcggatgctaatgcttattataaacacaagttactaacatgtgcttgggtcattcagcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcataggtaagctcttatgatcctgtccactgttgtaccatgtgtactttccaaatgattgctaacatgctatttttcgtcagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcgggtatatcatacatctctctggctgttgtttcagtgctatcaatcaagatgacacatcatgcaatattatcataacagcacattatgtgttttgaatattttattgattgtcatagctttgttttatgttttagcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcaggtatgcagtgtttcctgcacctacttgctacctattctgtatttgatgtattcaatttcctttttatcagttttctcgaaatatatgctaataggcttgtcactaaacggctacattatccagactctgttctgaatttttttttcattaagtactgttagctgccagattacatcatcatgactaaagtagtttagctacaatattgcaaagctagatttgtctagcctctattgttttgaatagaaaaggcaggtatgcagtgtttcctgcacctacttgctacctattctgtatttgatgtattcaatttcctttttatcagttttctcgaaatatatgctaataggcttgtcactaaacggctacattatccagactctgttctgaatttttttttcattaagtactgttagctgccagattacatcatcatgactaaagtagtttagctacaatattgcaaagctagatttgtctagcctctattgttttgaatagaaaattaaatcttttggtcatttttttgtaaccttttaggagtctctgtatgcttgtcgtcaacatgctatgataagtttatatcagttcttttggagctatgatgtattcttgtatattttatgactgctaggatattatgttaaaactgttttaaaggtatcatcaaacaagttaatatagatattgctacctaccaaagttggaatttgattatgtcgtaaaatatatgcaccctacatgttttagcttcatattgacttatattttcatatgtgctccttcaactgcacaaagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggtatggtcattcctgcttataagataccagaataatgctatgaagaccgtccttggttgtcagcattcttatacacaatgataatctttgtttttcatgggtaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggtcaagatattgttattgttatatttacatacttgttatcatatttaaaagcttcatattgtatgtacctttgaaaaaaaaagttccatactgtagcttcttttgttctcatttcttggtttcaaagcatcagcatcattaccgatataaaatatacccatttcctcagtcatcttccccgactcttgtatctttcacaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagaggtaacaccttcattcatgaggtccttaggtgcaaatcccagtgaagaataatcagttctactttgtatggttcatctcgcttactgaccggttcatctcgggatcaattacgagcagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaaggcatgtcctcttctctgatctcactcacgttcatgccaaacttactcatcatcatccatgatccatcatcctgactgaatcttgatatgcagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctgacaaggccacaccatgactgtcactgcaagcagctgctgtaggatcagcgagtaggaaaggatgacttgccggccagtttcctgagatgtgatttaagactgatattagctgatgttcccaagcatgatacggggcttgctcccaagatgtgatttttactttacttttggttaatgtgctaccgctgacattagatgttcaagcatatgaaaactgcttgtgctgtaacttgtatgtctgtaacgtaaaaatctttccatatggtaaccggggaagcagaggtatatccttttctgctgttgtatctatttccagtttcctgctgaacttgcaaatgaatattcagcaatttccgttttgctc</dnaseqindica>|<br />
Link = [http://www.ncbi.nlm.nih.gov/nuccore/NM_001048755.1 RefSeq:Os01g0182600]|<br />
}}<br />
[[Category:Genes]]<br />
[[Category:Japonica mRNA]]<br />
[[Category:Oryza Sativa Japonica Group]]<br />
[[Category:Japonica Genes]]<br />
[[Category:Japonica Chromosome 1]]<br />
[[Category:Chromosome 1]]</div>Mikahttps://ngdc.cncb.ac.cn/ricewiki/index.php?title=Os01g0182600&diff=183794Os01g01826002014-06-11T00:54:56Z<p>Mika: /* References */</p>
<hr />
<div>The rice gene of ''Gigantea(GI)'' is solo ortholog of ''GI'' in the Arabidopsis circadian system and related to photoperiodic control of flowering.<br />
<br />
==Annotated Information==<br />
===Function===<br />
*''OsGI'',a ''GI'' ortholog, regulates flowering time in response to photoperiodic conditions. Mutations in this gene delay flowering in inductive SDs (short-days),whereas flowering is marginally induced under LDs(long-days).In comparison, over expression of ''OsGI'' in transgenic rice causes late flowering in both SDs and LDs.In addition, ''OsGI'' has function to regulate ''Hd1'', a gene that also shows circadian rhythm<ref name="ref1" />. ''Os-GI'' affects gene expression of approximately 75% of genes among 27,201 genes and is required for strong amplitudes and proper phase-setting of global gene expression under natural field conditions<ref name="ref2" />. <br />
*The cDNA of ''OsGI'' firstly found in ''se5'' mutant that is completely photoperiod-insensitive, the expression patterns of rice and Arabidopsis ''GI'' gene are both photoperiodically regulated and are very similar under LD and SD conditions(FIG1)<ref name="ref3" />. ''GI'' is required for phase-dependent ''CO''transcription through the interaction with FKF1, which leads to cycling DNA binding with one finger (CDF) degradation and consequent derepression of CO. In addition, it has been known that GI protein interacts with ZEITLUPE (ZTL) F-box protein to control the diurnal rhythm of TOC1 protein in regulation of the Arabidopsis circadian clock<ref name="ref4" />. <br />
*''Os-GI'' also has function to maintain bioactive GA level through the regulation of the GA-deactivating enzyme genes in rice, ''Os-GI'' represses major ''OsGA2ox'' genes in order to maintain a proper bioactive GA level, ''osgi-1'' plants showed a semi-dwarf phenotype with elevated ''OsGA2ox'' gene expression(FIG2) <ref name="ref4" />.<br />
<br />
===Expression===<br />
*''GI'' encodes a nuclear protein that defines the proper amplitude and period length of circadian rhythms. ''GI'' expression is regulated by the circadian clock, with a peak in transcript levels at 8 to 10 h after dawn. The ''GI'' protein acts between the circadian oscillator and ''CO'' to promote flowering by increasing ''CO'' and ''FT'' mRNA abundance<ref name="ref1" />. The expression of the ''OsGI'' messenger RNA was circadian-controlled and that its temporal expression pattern was very similar to that of ''GI'' under both SD and LD conditions. Decreased ''OsGI'' expression led to early flowering under LD conditions and late flowering under SD conditions relative to the wild type, and that this early flowering phenotype under LD conditions was the opposite of that of the Arabidopsis ''gi'' mutants, confirming the opposite role of the ''GI'' gene in flowering under LD conditions in rice and Arabidopsis<ref name="ref5" />.<br />
<br />
===Mutation===<br />
*The single ''osgi-1'' mutation extended the time to flowering only under SD conditions. Under LD conditions the single ''hd1'' mutation induced early flowering, but ''osgi-1'' plants exhibited no phenotypic change<ref name="ref4" /><br />
<br />
===Evolution===<br />
* There is some evolutionary conservation between rice circadian clock–related genes and those associated with Arabidopsis circadian clocks, where the circadian clock consists of interlocked subloops, it is also possible that ''Os-GI'' is a member of a subloop in rice circadian clocks<ref name="ref4" />.But there is a divergence in the photoperiodic pathways of these two species arises from the molecular mechanisms controlling the expression of ''FT/Hd3a'' by ''CO/Hd1(Se1)''<ref name="ref5" />(figure 3)<br />
<br />
==Labs working on this gene==<br />
Please input related labs here.<br />
<br />
==Knowledge Extension==<br />
*The rice orthologues of the two key regulators of flowering time in Arabidopsis, ''CO'' and ''FT'', were recently shown to be important in the photoperiodic control of flowering in rice. The rice ''Hd1(Se1)''gene, which is an orthologue of the Arabidopsis ''CO'' gene, was required for the suppression of flowering under LD conditions and for the promotion of flowering under SD conditions. Furthermore, the rice ''Hd3a'' gene, an orthologue of the Arabidopsis ''FT'' gene, was shown to be an activator of flowering in rice. It was previously demonstrated that ''CO'' controls the flowering time by integrating signals from the circadian clock and light to regulate ''FT'' expression in Arabidopsis<ref name="ref5" />.Unlike Arabidopsis ''FT'', expression of ''Hd3a'' mRNA is more or less opposite to that of ''Hd1'' under inductive short-day conditions<ref name="ref6" />.<br />
<br />
==References==<br />
<references><br />
<ref name="ref1">Song Lim Kim, Shinyoung Lee, Hyo Jung Kim, Hong Gil Nam, Gynheung An（2007）''OsMADS51'' is a short-day flowering promoter that functions upstream of ''Ehd1'', ''OsMADS14'', and ''Hd3a''.''Plant Physiology'' 145(4): 1484-1494.</ref><br />
<ref name="ref2">Hironori Itoh, Takeshi Izawa (2011) A study of phytohormone biosynthetic gene expression using a circadian clock-related mutant in rice. ''Plant Signaling & Behavior'' 6(12): 1932-1936.</ref> <br />
<ref name="ref3">Ryousuke Hayama, Takeshi Izawa, Ko Shimamoto (2002) Isolation of rice genes possibly involved in the photoperiodic control of flowering by a<br />
fluorescent differential display method. ''Plant and Cell Physiology'' 43(5): 494-504.</ref> <br />
<ref name="ref4">Takeshi Izawa, Motohiro Mihara, Yuji Suzuki, Meenu Gupta, Hironori Itoh, et al. (2011) ''Os-GIGANTEA'' confers robust diurnal rhythms on the global transcriptome of rice in the field. ''The Plant Cell'' 23(5): 1741-1755.</ref> <br />
<ref name="ref5">Ryosuke Hayama, Shuji Yokoi, Shojiro Tamaki, Masahiro Yano, Ko Shimamoto (2003) Adaptation of photoperiodic control pathways produces short-day flowering in rice. ''Nature'' 422(6933): 719-722.</ref> <br />
<ref name="ref6">Bong-Soo Shin, Jeong-Hyun Lee, Jeong-Hwan Lee, Hyun-Joo Jeong, Choong-Hyo Yun, Jeong-Kook Kim (2004) Circadian regulation of rice (''Oryza sativa L.'') ''CONSTANS-like'' gene transcripts. ''Molecules and Cells'' 17(1): 10-16.</ref><br />
<br />
==Structured Information==<br />
{{JaponicaGene|<br />
GeneName = Os01g0182600|<br />
Description = GIGANTEA protein|<br />
Version = NM_001048755.1 GI:115434923 GeneID:4325329|<br />
Length = 9083 bp|<br />
Definition = Oryza sativa Japonica Group Os01g0182600, complete gene.|<br />
Source = Oryza sativa Japonica Group<br />
<br />
ORGANISM Oryza sativa Japonica Group<br />
Eukaryota; Viridiplantae; Streptophyta; Embryophyta; Tracheophyta;<br />
Spermatophyta; Magnoliophyta; Liliopsida; Poales; Poaceae; BEP<br />
clade; Ehrhartoideae; Oryzeae; Oryza.<br />
|<br />
Chromosome = [[:category:Japonica Chromosome 1|Chromosome 1]]|<br />
AP = Chromosome 1:4328362..4337444|<br />
CDS = 4328725..4329030,4329123..4329215,4329333..4329551,4329752..4329820,4329925..4331421<br>,4332220..4332429,4332563..4332713,4332791..4333031,4333898..4333956<br>,4334379..4334653,4334813..4334955,4335034..4335088,4335181..4335258<br>,4335351..4335437|<br />
GCID = <gbrowseImage1><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage1>|<br />
GSID = <gbrowseImage2><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage2>|<br />
CDNA = <cdnaseq>atgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactgcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcatagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcggcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctga</cdnaseq>|<br />
AA = <aaseq>MSASNEKWIDGLQFSSLFWPPPQDSQQKQAQILAYVEYFGQFTA DSEQFPEDIAQLIQSCYPSKEKRLVDEVLATFVLHHPEHGHAVVHPILSRIIDGTLSY DRNGFPFMSFISLFSHTSEKEYSEQWALACGEILRVLTHYNRPIFKVDHQHSEAECSS TSDQASSCESMEKRANGSPRNEPDRKPLRPLSPWITDILLAAPLGIRSDYFRWCGGVM GKYAAGGELKPPTTAYSRGSGKHPQLMPSTPRWAVANGAGVILSVCDEEVARYETANL TAAAVPALLLPPPTTPLDEHLVAGLPPLEPYARLFHRYYAIATPSATQRLLFGLLEAP PSWAPDALDAAVQLVELLRAAEDYDSGMRLPKNWMHLHFLRAIGTAMSMRAGIAADTS AALLFRILSQPTLLFPPLRHAEGVELHHEPLGGYVSSYKRQLEVPASEATIDATAQGI ASMLCAHGPDVEWRICTIWEAAYGLLPLSSSAVDLPEIVVAAPLQPPTLSWSLYLPLL KVFEYLPRGSPSEACLMRIFVATVEAILRRTFPSETSEQSRKPRSQSKNLAVAELRTM IHSLFVESCASMDLASRLLFVVLTVCVSHQALPGGSKRPTGSDNHSSEEVTNDSRLTN GRNRCKKRQGPVATFDSYVLAAVCALSCELQLFPFISKNGNHSNLKDSIKIVIPGKTT GISNELHNSISSAILHTRRILGILEALFSLKPSSVGTSWSYSSNEIVAAAMVAAHVSE LFRRSRPCLNALSALKQCKWDAEISTRASSLYHLIDLHGKTVTSIVNKAEPLEAHLTL TPVKKDEPPIEEKNINSSDGGALEKKDASRSHRKNGFARPLLKCAEDVILNGDVASTS GKAIASLQVEASDLANFLTMDRNGGYRGSQTLLRSVLSEKQELCFSVVSLLWQKLIAS PEMQMSAESTSAHQGWRKVVDALCDIVSASPTKASAAIVLQAEKDLQPWIARDDEQGQ KMWRVNQRIVKLIAELMRNHDSPEALVILASASDLLLRATDGMLVDGEACTLPQLELL EVTARAVHLIVEWGDSGVSVADGLSNLLKCRLSTTIRCLSHPSAHVRALSMSVLRDIL NSGQINSSKLIQGEHRNGIQSPTYQCLAASIINWQADVERCIEWEAHSRRATGLTLAF LTAAAKELGCPLTC</aaseq>|<br />
DNA = <dnaseqindica>8415..8720#8230..8322#7894..8112#7625..7693#6024..7520#5016..5225#4732..4882#4414..4654#3489..3547#2792..3066#2490..2632#2357..2411#2187..2264#2008..2094#ccgagccgtttctctccctctctctctcctcctccttcccctgcgcctcctccaccgcccgtgtcgccgccgcgcgcgtctatcccctcgccgccgccgccgtctacgccgcgtgaggtccggattggttgaaaggctgatcgaggtaatgcttttttatttttcccgtgcggttttagtttttctgtgctgtgcttggtggttttttctttaaaactatttttaggctggttttgatgatggcctttctaaggttgtcgtgtatggggggcgaggtggttcgggtggatttgtggtgttatgttgaggggatttttgtttggggggagagagttttggtggtagatgcggttgggattttgaagacgtgaaagggaaattagttttgaaatttgacggcgaggttgagctgattagtattgtaataattcgtggcatcggatagatcaacgcgagatggatgcttattactgcgaatttcaggaggtttttgtggtttttttaggcgagtagcgttgattctagctggattttgctgcttcgtggaggcgtttagatgtggtgcatacggctgtggatgtaatttgtgcggggataattactgaaaattaattcgcagtaagctagctctgctttaatattttttcccctgaaacagtaacaagggatcccttcctcatttcgttaacagtggaagtgaatacagacaccaaccgatgagcagagacacctgtaattaaacaccaaaagaagaacctgttttgttcctatatattttccatttctatgctttaggtaatatactttctggttcggtgtcatagtacccgaaatagaatctttgcttgccaaaaaaagaaagaataacaagatagaacaacatcattaaattctgtcttagacaatacatttataatcgaccaacaaaaacagaccgatgagtgacgtggaataacaaatgaagtataaatgtcacgaaaaacacatctattgattttgtgtgataagccataggagcctacgaggaaccaagtgaatggttctgtacaccacatccattagcattaccataaatttgcctatttctatgatgccttccttaacatatttatcgacatgtgagcttgcaccataactaaatgttatggtaaactttaccttataaatgattgcaacatgtaccttttcatttcgataattaaaaacgagaaagcttccacctttttcctagatatattttttcgtttgattatgaatttagttgtacttgacaataaaataatgcgcttgtaagattatgcaattttcgttgcattacctccctctcttttcatttgtcagtttttttctactccctccgtttcatattttaagtcgtttgacttttttcctagtcagactttgttaagtttgacaaagtttatagtaaaatttagcaatatctaaaatattaaattggtttcatcaaatctaacattgaatatattttgataatatgttttttttgtgttgaaaatacttctatatttttctataaacttagtcaaacttggaagaaatttcagtaagaaaaaagtcaaacggcttataatatgaaacggagggagtaaatttttttttgcattaattaaaattcatgtattttcctccttttttggtttttgttgtgcgatatattacagtgtagagactctgaagctcatatggttggcattggttgatatgataacgtgtcacgttgtgagatgcagacccccgttatgggggttgattgaatgaagtggatagcactcttataactttcttgctcgaagtttcttgataccaaggcaagtagcttataagaagatttggactttggtggcataaatggatgaactaactggcggcttcaacgtggtgttgattacaggtgcttttattgctaactggagcagtttgtgtagtttctttctttctctttgagaattctctcagtttcttgggcaggctcttgagctaccaagtatgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggtaacattattcctatttgattcttgttagaaattctatatgttgaacatacttgcctcaatgggaaacttggttgcgtctcctctctgaaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcaggtactgtatctctaagcatgtttagtatcttctgtgataaaatctttctccagtgtgttaacttttgcattctttttcttgcatgcttgcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattaggtaatttgatccagtgatcaatacatgttccacaaacaattgaagaaagagtttgacatatttttctcatatttccagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgaggtaaggcccctactctaatggttagttccaatttattggttcatttctctgtgtttgataagattttgcctctgctgtgcatttcaatattatatctaaactgtgctgcttatagcttatcaaactactcttgctcataaagatgcaacttttatgcagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgagtcataataaatgataacatattattgttctgtggactgaaaatctgaaatcctcctttcttgagaaatacaacctcaattttatggtgatattgacaagaaggaagagcgcatataatttaaagttaaaacacggaaaataatagaagtatgtgagcagaacgcaggagcaaggggtggaactcataacaagttcaaacttcatgcatgttcataattccccaatgcaagtatggcaagaaaaatagacaacttttaaaagtatgctaatggaactgcatgcattgtgtaaacttgcttttgtcaagcatgttggaacatactatatatttgctgctgatgatgaaaaatttgcttgagtactcttctattgctaatggacgcttaccattccctgttttgtcttcaaatgcatcaggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactggttggtaaaatcagcaatggaattttctatcgtgaattttttataattattgtaacattaattactcccatgtctttttattctgttgagataccatgtgacaatgtctacaattgcatattttgcattctttggtagatcagttcccttcttaagatccccaaagggaaactggagtatcaataacttactccctcaaataaaaaaaaagtgatactttaggcataaacacattcttcaaaacaaaactttgaatagcagtttcttttgtaatatattaataaagcatggaaaaattataatgtgaagaatgtacattcagggacaaatatactcatcatttttgcatggttgatcttaatgtccatagagagattgatcagaccaaaagtttatttttgacaggagtgagtatggattataataggagtgaatggcaatatgctgtgaaagtgagcattttatatttgtttactactagcatttggtgacaaagcaaatgatgtatattgtaaatcaaagtaaacaagtagaacatgtcctgtggcgacaagacataacctacaaattaaagttagcctgattgtttattttactgggagcacattttttctagattctagagtacatattattaaaaaaggtgtgatatgtttgcaaattactttttaatggaagtttgtaaattactttgaaacttgcattcagtttttttgcttcttaaaaaacatagtgcatatgggctttatgagctcatactatgcatgtttttctgcattggagtgggacctttatctttttcttgcaactcggatgctaatgcttattataaacacaagttactaacatgtgcttgggtcattcagcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcataggtaagctcttatgatcctgtccactgttgtaccatgtgtactttccaaatgattgctaacatgctatttttcgtcagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtac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/>
Link = [http://www.ncbi.nlm.nih.gov/nuccore/NM_001048755.1 RefSeq:Os01g0182600]|<br />
}}<br />
[[Category:Genes]]<br />
[[Category:Japonica mRNA]]<br />
[[Category:Oryza Sativa Japonica Group]]<br />
[[Category:Japonica Genes]]<br />
[[Category:Japonica Chromosome 1]]<br />
[[Category:Chromosome 1]]</div>Mikahttps://ngdc.cncb.ac.cn/ricewiki/index.php?title=Os01g0182600&diff=183712Os01g01826002014-06-10T15:06:25Z<p>Mika: </p>
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<div>The rice gene of ''Gigantea(GI)'' is solo ortholog of ''GI'' in the Arabidopsis circadian system and related to photoperiodic control of flowering.<br />
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==Annotated Information==<br />
===Function===<br />
*''OsGI'',a ''GI'' ortholog, regulates flowering time in response to photoperiodic conditions. Mutations in this gene delay flowering in inductive SDs (short-days),whereas flowering is marginally induced under LDs(long-days).In comparison, over expression of ''OsGI'' in transgenic rice causes late flowering in both SDs and LDs.In addition, ''OsGI'' has function to regulate ''Hd1'', a gene that also shows circadian rhythm<ref name="ref1" />. ''Os-GI'' affects gene expression of approximately 75% of genes among 27,201 genes and is required for strong amplitudes and proper phase-setting of global gene expression under natural field conditions<ref name="ref2" />. <br />
*The cDNA of ''OsGI'' firstly found in ''se5'' mutant that is completely photoperiod-insensitive, the expression patterns of rice and Arabidopsis ''GI'' gene are both photoperiodically regulated and are very similar under LD and SD conditions(FIG1)<ref name="ref3" />. ''GI'' is required for phase-dependent ''CO''transcription through the interaction with FKF1, which leads to cycling DNA binding with one finger (CDF) degradation and consequent derepression of CO. In addition, it has been known that GI protein interacts with ZEITLUPE (ZTL) F-box protein to control the diurnal rhythm of TOC1 protein in regulation of the Arabidopsis circadian clock<ref name="ref4" />. <br />
*''Os-GI'' also has function to maintain bioactive GA level through the regulation of the GA-deactivating enzyme genes in rice, ''Os-GI'' represses major ''OsGA2ox'' genes in order to maintain a proper bioactive GA level, ''osgi-1'' plants showed a semi-dwarf phenotype with elevated ''OsGA2ox'' gene expression(FIG2) <ref name="ref4" />.<br />
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===Expression===<br />
*''GI'' encodes a nuclear protein that defines the proper amplitude and period length of circadian rhythms. ''GI'' expression is regulated by the circadian clock, with a peak in transcript levels at 8 to 10 h after dawn. The ''GI'' protein acts between the circadian oscillator and ''CO'' to promote flowering by increasing ''CO'' and ''FT'' mRNA abundance<ref name="ref1" />. The expression of the ''OsGI'' messenger RNA was circadian-controlled and that its temporal expression pattern was very similar to that of ''GI'' under both SD and LD conditions. Decreased ''OsGI'' expression led to early flowering under LD conditions and late flowering under SD conditions relative to the wild type, and that this early flowering phenotype under LD conditions was the opposite of that of the Arabidopsis ''gi'' mutants, confirming the opposite role of the ''GI'' gene in flowering under LD conditions in rice and Arabidopsis<ref name="ref5" />.<br />
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===Mutation===<br />
*The single ''osgi-1'' mutation extended the time to flowering only under SD conditions. Under LD conditions the single ''hd1'' mutation induced early flowering, but ''osgi-1'' plants exhibited no phenotypic change<ref name="ref4" /><br />
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===Evolution===<br />
* There is some evolutionary conservation between rice circadian clock–related genes and those associated with Arabidopsis circadian clocks, where the circadian clock consists of interlocked subloops, it is also possible that ''Os-GI'' is a member of a subloop in rice circadian clocks<ref name="ref4" />.But there is a divergence in the photoperiodic pathways of these two species arises from the molecular mechanisms controlling the expression of ''FT/Hd3a'' by ''CO/Hd1(Se1)''<ref name="ref5" />(figure 3)<br />
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==Labs working on this gene==<br />
Please input related labs here.<br />
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==Knowledge Extension==<br />
*The rice orthologues of the two key regulators of flowering time in Arabidopsis, ''CO'' and ''FT'', were recently shown to be important in the photoperiodic control of flowering in rice. The rice ''Hd1(Se1)''gene, which is an orthologue of the Arabidopsis ''CO'' gene, was required for the suppression of flowering under LD conditions and for the promotion of flowering under SD conditions. Furthermore, the rice ''Hd3a'' gene, an orthologue of the Arabidopsis ''FT'' gene, was shown to be an activator of flowering in rice. It was previously demonstrated that ''CO'' controls the flowering time by integrating signals from the circadian clock and light to regulate ''FT'' expression in Arabidopsis<ref name="ref5" />.Unlike Arabidopsis ''FT'', expression of ''Hd3a'' mRNA is more or less opposite to that of ''Hd1'' under inductive short-day conditions<ref name="ref6" />.<br />
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==References==<br />
Please input cited references here.<br />
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==Structured Information==<br />
{{JaponicaGene|<br />
GeneName = Os01g0182600|<br />
Description = GIGANTEA protein|<br />
Version = NM_001048755.1 GI:115434923 GeneID:4325329|<br />
Length = 9083 bp|<br />
Definition = Oryza sativa Japonica Group Os01g0182600, complete gene.|<br />
Source = Oryza sativa Japonica Group<br />
<br />
ORGANISM Oryza sativa Japonica Group<br />
Eukaryota; Viridiplantae; Streptophyta; Embryophyta; Tracheophyta;<br />
Spermatophyta; Magnoliophyta; Liliopsida; Poales; Poaceae; BEP<br />
clade; Ehrhartoideae; Oryzeae; Oryza.<br />
|<br />
Chromosome = [[:category:Japonica Chromosome 1|Chromosome 1]]|<br />
AP = Chromosome 1:4328362..4337444|<br />
CDS = 4328725..4329030,4329123..4329215,4329333..4329551,4329752..4329820,4329925..4331421<br>,4332220..4332429,4332563..4332713,4332791..4333031,4333898..4333956<br>,4334379..4334653,4334813..4334955,4335034..4335088,4335181..4335258<br>,4335351..4335437|<br />
GCID = <gbrowseImage1><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage1>|<br />
GSID = <gbrowseImage2><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage2>|<br />
CDNA = <cdnaseq>atgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactgcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcatagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcggcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctga</cdnaseq>|<br />
AA = <aaseq>MSASNEKWIDGLQFSSLFWPPPQDSQQKQAQILAYVEYFGQFTA DSEQFPEDIAQLIQSCYPSKEKRLVDEVLATFVLHHPEHGHAVVHPILSRIIDGTLSY DRNGFPFMSFISLFSHTSEKEYSEQWALACGEILRVLTHYNRPIFKVDHQHSEAECSS TSDQASSCESMEKRANGSPRNEPDRKPLRPLSPWITDILLAAPLGIRSDYFRWCGGVM GKYAAGGELKPPTTAYSRGSGKHPQLMPSTPRWAVANGAGVILSVCDEEVARYETANL TAAAVPALLLPPPTTPLDEHLVAGLPPLEPYARLFHRYYAIATPSATQRLLFGLLEAP PSWAPDALDAAVQLVELLRAAEDYDSGMRLPKNWMHLHFLRAIGTAMSMRAGIAADTS AALLFRILSQPTLLFPPLRHAEGVELHHEPLGGYVSSYKRQLEVPASEATIDATAQGI ASMLCAHGPDVEWRICTIWEAAYGLLPLSSSAVDLPEIVVAAPLQPPTLSWSLYLPLL KVFEYLPRGSPSEACLMRIFVATVEAILRRTFPSETSEQSRKPRSQSKNLAVAELRTM IHSLFVESCASMDLASRLLFVVLTVCVSHQALPGGSKRPTGSDNHSSEEVTNDSRLTN GRNRCKKRQGPVATFDSYVLAAVCALSCELQLFPFISKNGNHSNLKDSIKIVIPGKTT GISNELHNSISSAILHTRRILGILEALFSLKPSSVGTSWSYSSNEIVAAAMVAAHVSE LFRRSRPCLNALSALKQCKWDAEISTRASSLYHLIDLHGKTVTSIVNKAEPLEAHLTL TPVKKDEPPIEEKNINSSDGGALEKKDASRSHRKNGFARPLLKCAEDVILNGDVASTS GKAIASLQVEASDLANFLTMDRNGGYRGSQTLLRSVLSEKQELCFSVVSLLWQKLIAS PEMQMSAESTSAHQGWRKVVDALCDIVSASPTKASAAIVLQAEKDLQPWIARDDEQGQ KMWRVNQRIVKLIAELMRNHDSPEALVILASASDLLLRATDGMLVDGEACTLPQLELL EVTARAVHLIVEWGDSGVSVADGLSNLLKCRLSTTIRCLSHPSAHVRALSMSVLRDIL NSGQINSSKLIQGEHRNGIQSPTYQCLAASIINWQADVERCIEWEAHSRRATGLTLAF LTAAAKELGCPLTC</aaseq>|<br />
DNA = <dnaseqindica>8415..8720#8230..8322#7894..8112#7625..7693#6024..7520#5016..5225#4732..4882#4414..4654#3489..3547#2792..3066#2490..2632#2357..2411#2187..2264#2008..2094#ccgagccgtttctctccctctctctctcctcctccttcccctgcgcctcctccaccgcccgtgtcgccgccgcgcgcgtctatcccctcgccgccgccgccgtctacgccgcgtgaggtccggattggttgaaaggctgatcgaggtaatgcttttttatttttcccgtgcggttttagtttttctgtgctgtgcttggtggttttttctttaaaactatttttaggctggttttgatgatggcctttctaaggttgtcgtgtatggggggcgaggtggttcgggtggatttgtggtgttatgttgaggggatttttgtttggggggagagagttttggtggtagatgcggttgggattttgaagacgtgaaagggaaattagttttgaaatttgacggcgaggttgagctgattagtattgtaataattcgtggcatcggatagatcaacgcgagatggatgcttattactgcgaatttcaggaggtttttgtggtttttttaggcgagtagcgttgattctagctggattttgctgcttcgtggaggcgtttagatgtggtgcatacggctgtggatgtaatttgtgcggggataattactgaaaattaattcgcagtaagctagctctgctttaatattttttcccctgaaacagtaacaagggatcccttcctcatttcgttaacagtggaagtgaatacagacaccaaccgatgagcagagacacctgtaattaaacaccaaaagaagaacctgttttgttcctatatattttccatttctatgctttaggtaatatactttctggttcggtgtcatagtacccgaaatagaatctttgcttgccaaaaaaagaaagaataacaagatagaacaacatcattaaattctgtcttagacaatacatttataatcgaccaacaaaaacagaccgatgagtgacgtggaataacaaatgaagtataaatgtcacgaaaaacacatctattgattttgtgtgataagccataggagcctacgaggaaccaagtgaatggttctgtacaccacatccattagcattaccataaatttgcctatttctatgatgccttccttaacatatttatcgacatgtgagcttgcaccataactaaatgttatggtaaactttaccttataaatgattgcaacatgtaccttttcatttcgataattaaaaacgagaaagcttccacctttttcctagatatattttttcgtttgattatgaatttagttgtacttgacaataaaataatgcgcttgtaagattatgcaattttcgttgcattacctccctctcttttcatttgtcagtttttttctactccctccgtttcatattttaagtcgtttgacttttttcctagtcagactttgttaagtttgacaaagtttatagtaaaatttagcaatatctaaaatattaaattggtttcatcaaatctaacattgaatatattttgataatatgttttttttgtgttgaaaatacttctatatttttctataaacttagtcaaacttggaagaaatttcagtaagaaaaaagtcaaacggcttataatatgaaacggagggagtaaatttttttttgcattaattaaaattcatgtattttcctccttttttggtttttgttgtgcgatatattacagtgtagagactctgaagctcatatggttggcattggttgatatgataacgtgtcacgttgtgagatgcagacccccgttatgggggttgattgaatgaagtggatagcactcttataactttcttgctcgaagtttcttgataccaaggcaagtagcttataagaagatttggactttggtggcataaatggatgaactaactggcggcttcaacgtggtgttgattacaggtgcttttattgctaactggagcagtttgtgtagtttctttctttctctttgagaattctctcagtttcttgggcaggctcttgagctaccaagtatgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggtaacattattcctatttgattcttgttagaaattctatatgttgaacatacttgcctcaatgggaaacttggttgcgtctcctctctgaaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcaggtactgtatctctaagcatgtttagtatcttctgtgataaaatctttctccagtgtgttaacttttgcattctttttcttgcatgcttgcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattaggtaatttgatccagtgatcaatacatgttccacaaacaattgaagaaagagtttgacatatttttctcatatttccagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgaggtaaggcccctactctaatggttagttccaatttattggttcatttctctgtgtttgataagattttgcctctgctgtgcatttcaatattatatctaaactgtgctgcttatagcttatcaaactactcttgctcataaagatgcaacttttatgcagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgagtcataataaatgataacatattattgttctgtggactgaaaatctgaaatcctcctttcttgagaaatacaacctcaattttatggtgatattgacaagaaggaagagcgcatataatttaaagttaaaacacggaaaataatagaagtatgtgagcagaacgcaggagcaaggggtggaactcataacaagttcaaacttcatgcatgttcataattccccaatgcaagtatggcaagaaaaatagacaacttttaaaagtatgctaatggaactgcatgcattgtgtaaacttgcttttgtcaagcatgttggaacatactatatatttgctgctgatgatgaaaaatttgcttgagtactcttctattgctaatggacgcttaccattccctgttttgtcttcaaatgcatcaggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactggttggtaaaatcagcaatggaattttctatcgtgaattttttataattattgtaacattaattactcccatgtctttttattctgttgagataccatgtgacaatgtctacaattgcatattttgcattctttggtagatcagttcccttcttaagatccccaaagggaaactggagtatcaataacttactccctcaaataaaaaaaaagtgatactttaggcataaacacattcttcaaaacaaaactttgaatagcagtttcttttgtaatatattaataaagcatggaaaaattataatgtgaagaatgtacattcagggacaaatatactcatcatttttgcatggttgatcttaatgtccatagagagattgatcagaccaaaagtttatttttgacaggagtgagtatggattataataggagtgaatggcaatatgctgtgaaagtgagcattttatatttgtttactactagcatttggtgacaaagcaaatgatgtatattgtaaatcaaagtaaacaagtagaacatgtcctgtggcgacaagacataacctacaaattaaagttagcctgattgtttattttactgggagcacattttttctagattctagagtacatattattaaaaaaggtgtgatatgtttgcaaattactttttaatggaagtttgtaaattactttgaaacttgcattcagtttttttgcttcttaaaaaacatagtgcatatgggctttatgagctcatactatgcatgtttttctgcattggagtgggacctttatctttttcttgcaactcggatgctaatgcttattataaacacaagttactaacatgtgcttgggtcattcagcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcataggtaagctcttatgatcctgtccactgttgtaccatgtgtactttccaaatgattgctaacatgctatttttcgtcagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcgggtatatcatacatctctctggctgttgtttcagtgctatcaatcaagatgacacatcatgcaatattatcataacagcacattatgtgttttgaatattttattgattgtcatagctttgttttatgttttagcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcaggtatgcagtgtttcctgcacctacttgctacctattctgtatttgatgtattcaatttcctttttatcagttttctcgaaatatatgctaataggcttgtcactaaacggctacattatccagactctgttctgaatttttttttcattaagtactgttagctgccagattacatcatcatgactaaagtagtttagctacaatattgcaaagctagatttgtctagcctctattgttttgaatagaaaaggcaggtatgcagtgtttcctgcacctacttgctacctattctgtatttgatgtattcaatttcctttttatcagttttctcgaaatatatgctaataggcttgtcactaaacggctacattatccagactctgttctgaatttttttttcattaagtactgttagctgccagattacatcatcatgactaaagtagtttagctacaatattgcaaagctagatttgtctagcctctattgttttgaatagaaaattaaatcttttggtcatttttttgtaaccttttaggagtctctgtatgcttgtcgtcaacatgctatgataagtttatatcagttcttttggagctatgatgtattcttgtatattttatgactgctaggatattatgttaaaactgttttaaaggtatcatcaaacaagttaatatagatattgctacctaccaaagttggaatttgattatgtcgtaaaatatatgcaccctacatgttttagcttcatattgacttatattttcatatgtgctccttcaactgcacaaagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggtatggtcattcctgcttataagataccagaataatgctatgaagaccgtccttggttgtcagcattcttatacacaatgataatctttgtttttcatgggtaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggtcaagatattgttattgttatatttacatacttgttatcatatttaaaagcttcatattgtatgtacctttgaaaaaaaaagttccatactgtagcttcttttgttctcatttcttggtttcaaagcatcagcatcattaccgatataaaatatacccatttcctcagtcatcttccccgactcttgtatctttcacaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagaggtaacaccttcattcatgaggtccttaggtgcaaatcccagtgaagaataatcagttctactttgtatggttcatctcgcttactgaccggttcatctcgggatcaattacgagcagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaaggcatgtcctcttctctgatctcactcacgttcatgccaaacttactcatcatcatccatgatccatcatcctgactgaatcttgatatgcagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctgacaaggccacaccatgactgtcactgcaagcagctgctgtaggatcagcgagtaggaaaggatgacttgccggccagtttcctgagatgtgatttaagactgatattagctgatgttcccaagcatgatacggggcttgctcccaagatgtgatttttactttacttttggttaatgtgctaccgctgacattagatgttcaagcatatgaaaactgcttgtgctgtaacttgtatgtctgtaacgtaaaaatctttccatatggtaaccggggaagcagaggtatatccttttctgctgttgtatctatttccagtttcctgctgaacttgcaaatgaatattcagcaatttccgttttgctc</dnaseqindica>|<br />
Link = [http://www.ncbi.nlm.nih.gov/nuccore/NM_001048755.1 RefSeq:Os01g0182600]|<br />
}}<br />
[[Category:Genes]]<br />
[[Category:Japonica mRNA]]<br />
[[Category:Oryza Sativa Japonica Group]]<br />
[[Category:Japonica Genes]]<br />
[[Category:Japonica Chromosome 1]]<br />
[[Category:Chromosome 1]]</div>Mikahttps://ngdc.cncb.ac.cn/ricewiki/index.php?title=Os01g0182600&diff=183711Os01g01826002014-06-10T15:06:02Z<p>Mika: </p>
<hr />
<div>The rice gene of Gigantea(GI) is solo ortholog of GI in the Arabidopsis circadian system and related to photoperiodic control of flowering.<br />
<br />
==Annotated Information==<br />
===Function===<br />
*''OsGI'',a ''GI'' ortholog, regulates flowering time in response to photoperiodic conditions. Mutations in this gene delay flowering in inductive SDs (short-days),whereas flowering is marginally induced under LDs(long-days).In comparison, over expression of ''OsGI'' in transgenic rice causes late flowering in both SDs and LDs.In addition, ''OsGI'' has function to regulate ''Hd1'', a gene that also shows circadian rhythm<ref name="ref1" />. ''Os-GI'' affects gene expression of approximately 75% of genes among 27,201 genes and is required for strong amplitudes and proper phase-setting of global gene expression under natural field conditions<ref name="ref2" />. <br />
*The cDNA of ''OsGI'' firstly found in ''se5'' mutant that is completely photoperiod-insensitive, the expression patterns of rice and Arabidopsis ''GI'' gene are both photoperiodically regulated and are very similar under LD and SD conditions(FIG1)<ref name="ref3" />. ''GI'' is required for phase-dependent ''CO''transcription through the interaction with FKF1, which leads to cycling DNA binding with one finger (CDF) degradation and consequent derepression of CO. In addition, it has been known that GI protein interacts with ZEITLUPE (ZTL) F-box protein to control the diurnal rhythm of TOC1 protein in regulation of the Arabidopsis circadian clock<ref name="ref4" />. <br />
*''Os-GI'' also has function to maintain bioactive GA level through the regulation of the GA-deactivating enzyme genes in rice, ''Os-GI'' represses major ''OsGA2ox'' genes in order to maintain a proper bioactive GA level, ''osgi-1'' plants showed a semi-dwarf phenotype with elevated ''OsGA2ox'' gene expression(FIG2) <ref name="ref4" />.<br />
<br />
===Expression===<br />
*''GI'' encodes a nuclear protein that defines the proper amplitude and period length of circadian rhythms. ''GI'' expression is regulated by the circadian clock, with a peak in transcript levels at 8 to 10 h after dawn. The ''GI'' protein acts between the circadian oscillator and ''CO'' to promote flowering by increasing ''CO'' and ''FT'' mRNA abundance<ref name="ref1" />. The expression of the ''OsGI'' messenger RNA was circadian-controlled and that its temporal expression pattern was very similar to that of ''GI'' under both SD and LD conditions. Decreased ''OsGI'' expression led to early flowering under LD conditions and late flowering under SD conditions relative to the wild type, and that this early flowering phenotype under LD conditions was the opposite of that of the Arabidopsis ''gi'' mutants, confirming the opposite role of the ''GI'' gene in flowering under LD conditions in rice and Arabidopsis<ref name="ref5" />.<br />
<br />
===Mutation===<br />
*The single ''osgi-1'' mutation extended the time to flowering only under SD conditions. Under LD conditions the single ''hd1'' mutation induced early flowering, but ''osgi-1'' plants exhibited no phenotypic change<ref name="ref4" /><br />
<br />
===Evolution===<br />
* There is some evolutionary conservation between rice circadian clock–related genes and those associated with Arabidopsis circadian clocks, where the circadian clock consists of interlocked subloops, it is also possible that ''Os-GI'' is a member of a subloop in rice circadian clocks<ref name="ref4" />.But there is a divergence in the photoperiodic pathways of these two species arises from the molecular mechanisms controlling the expression of ''FT/Hd3a'' by ''CO/Hd1(Se1)''<ref name="ref5" />(figure 3)<br />
<br />
==Labs working on this gene==<br />
Please input related labs here.<br />
<br />
==Knowledge Extension==<br />
*The rice orthologues of the two key regulators of flowering time in Arabidopsis, ''CO'' and ''FT'', were recently shown to be important in the photoperiodic control of flowering in rice. The rice ''Hd1(Se1)''gene, which is an orthologue of the Arabidopsis ''CO'' gene, was required for the suppression of flowering under LD conditions and for the promotion of flowering under SD conditions. Furthermore, the rice ''Hd3a'' gene, an orthologue of the Arabidopsis ''FT'' gene, was shown to be an activator of flowering in rice. It was previously demonstrated that ''CO'' controls the flowering time by integrating signals from the circadian clock and light to regulate ''FT'' expression in Arabidopsis<ref name="ref5" />.Unlike Arabidopsis ''FT'', expression of ''Hd3a'' mRNA is more or less opposite to that of ''Hd1'' under inductive short-day conditions<ref name="ref6" />.<br />
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==References==<br />
Please input cited references here.<br />
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==Structured Information==<br />
{{JaponicaGene|<br />
GeneName = Os01g0182600|<br />
Description = GIGANTEA protein|<br />
Version = NM_001048755.1 GI:115434923 GeneID:4325329|<br />
Length = 9083 bp|<br />
Definition = Oryza sativa Japonica Group Os01g0182600, complete gene.|<br />
Source = Oryza sativa Japonica Group<br />
<br />
ORGANISM Oryza sativa Japonica Group<br />
Eukaryota; Viridiplantae; Streptophyta; Embryophyta; Tracheophyta;<br />
Spermatophyta; Magnoliophyta; Liliopsida; Poales; Poaceae; BEP<br />
clade; Ehrhartoideae; Oryzeae; Oryza.<br />
|<br />
Chromosome = [[:category:Japonica Chromosome 1|Chromosome 1]]|<br />
AP = Chromosome 1:4328362..4337444|<br />
CDS = 4328725..4329030,4329123..4329215,4329333..4329551,4329752..4329820,4329925..4331421<br>,4332220..4332429,4332563..4332713,4332791..4333031,4333898..4333956<br>,4334379..4334653,4334813..4334955,4335034..4335088,4335181..4335258<br>,4335351..4335437|<br />
GCID = <gbrowseImage1><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage1>|<br />
GSID = <gbrowseImage2><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage2>|<br />
CDNA = <cdnaseq>atgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactgcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcatagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcggcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctga</cdnaseq>|<br />
AA = <aaseq>MSASNEKWIDGLQFSSLFWPPPQDSQQKQAQILAYVEYFGQFTA DSEQFPEDIAQLIQSCYPSKEKRLVDEVLATFVLHHPEHGHAVVHPILSRIIDGTLSY DRNGFPFMSFISLFSHTSEKEYSEQWALACGEILRVLTHYNRPIFKVDHQHSEAECSS TSDQASSCESMEKRANGSPRNEPDRKPLRPLSPWITDILLAAPLGIRSDYFRWCGGVM GKYAAGGELKPPTTAYSRGSGKHPQLMPSTPRWAVANGAGVILSVCDEEVARYETANL TAAAVPALLLPPPTTPLDEHLVAGLPPLEPYARLFHRYYAIATPSATQRLLFGLLEAP PSWAPDALDAAVQLVELLRAAEDYDSGMRLPKNWMHLHFLRAIGTAMSMRAGIAADTS AALLFRILSQPTLLFPPLRHAEGVELHHEPLGGYVSSYKRQLEVPASEATIDATAQGI ASMLCAHGPDVEWRICTIWEAAYGLLPLSSSAVDLPEIVVAAPLQPPTLSWSLYLPLL KVFEYLPRGSPSEACLMRIFVATVEAILRRTFPSETSEQSRKPRSQSKNLAVAELRTM IHSLFVESCASMDLASRLLFVVLTVCVSHQALPGGSKRPTGSDNHSSEEVTNDSRLTN GRNRCKKRQGPVATFDSYVLAAVCALSCELQLFPFISKNGNHSNLKDSIKIVIPGKTT GISNELHNSISSAILHTRRILGILEALFSLKPSSVGTSWSYSSNEIVAAAMVAAHVSE LFRRSRPCLNALSALKQCKWDAEISTRASSLYHLIDLHGKTVTSIVNKAEPLEAHLTL TPVKKDEPPIEEKNINSSDGGALEKKDASRSHRKNGFARPLLKCAEDVILNGDVASTS GKAIASLQVEASDLANFLTMDRNGGYRGSQTLLRSVLSEKQELCFSVVSLLWQKLIAS PEMQMSAESTSAHQGWRKVVDALCDIVSASPTKASAAIVLQAEKDLQPWIARDDEQGQ KMWRVNQRIVKLIAELMRNHDSPEALVILASASDLLLRATDGMLVDGEACTLPQLELL EVTARAVHLIVEWGDSGVSVADGLSNLLKCRLSTTIRCLSHPSAHVRALSMSVLRDIL NSGQINSSKLIQGEHRNGIQSPTYQCLAASIINWQADVERCIEWEAHSRRATGLTLAF LTAAAKELGCPLTC</aaseq>|<br />
DNA = <dnaseqindica>8415..8720#8230..8322#7894..8112#7625..7693#6024..7520#5016..5225#4732..4882#4414..4654#3489..3547#2792..3066#2490..2632#2357..2411#2187..2264#2008..2094#ccgagccgtttctctccctctctctctcctcctccttcccctgcgcctcctccaccgcccgtgtcgccgccgcgcgcgtctatcccctcgccgccgccgccgtctacgccgcgtgaggtccggattggttgaaaggctgatcgaggtaatgcttttttatttttcccgtgcggttttagtttttctgtgctgtgcttggtggttttttctttaaaactatttttaggctggttttgatgatggcctttctaaggttgtcgtgtatggggggcgaggtggttcgggtggatttgtggtgttatgttgaggggatttttgtttggggggagagagttttggtggtagatgcggttgggattttgaagacgtgaaagggaaattagttttgaaatttgacggcgaggttgagctgattagtattgtaataattcgtggcatcggatagatcaacgcgagatggatgcttattactgcgaatttcaggaggtttttgtggtttttttaggcgagtagcgttgattctagctggattttgctgcttcgtggaggcgtttagatgtggtgcatacggctgtggatgtaatttgtgcggggataattactgaaaattaattcgcagtaagctagctctgctttaatattttttcccctgaaacagtaacaagggatcccttcctcatttcgttaacagtggaagtgaatacagacaccaaccgatgagcagagacacctgtaattaaacaccaaaagaagaacctgttttgttcctatatattttccatttctatgctttaggtaatatactttctggttcggtgtcatagtacccgaaatagaatctttgcttgccaaaaaaagaaagaataacaagatagaacaacatcattaaattctgtcttagacaatacatttataatcgaccaacaaaaacagaccgatgagtgacgtggaataacaaatgaagtataaatgtcacgaaaaacacatctattgattttgtgtgataagccataggagcctacgaggaaccaagtgaatggttctgtacaccacatccattagcattaccataaatttgcctatttctatgatgccttccttaacatatttatcgacatgtgagcttgcaccataactaaatgttatggtaaactttaccttataaatgattgcaacatgtaccttttcatttcgataattaaaaacgagaaagcttccacctttttcctagatatattttttcgtttgattatgaatttagttgtacttgacaataaaataatgcgcttgtaagattatgcaattttcgttgcattacctccctctcttttcatttgtcagtttttttctactccctccgtttcatattttaagtcgtttgacttttttcctagtcagactttgttaagtttgacaaagtttatagtaaaatttagcaatatctaaaatattaaattggtttcatcaaatctaacattgaatatattttgataatatgttttttttgtgttgaaaatacttctatatttttctataaacttagtcaaacttggaagaaatttcagtaagaaaaaagtcaaacggcttataatatgaaacggagggagtaaatttttttttgcattaattaaaattcatgtattttcctccttttttggtttttgttgtgcgatatattacagtgtagagactctgaagctcatatggttggcattggttgatatgataacgtgtcacgttgtgagatgcagacccccgttatgggggttgattgaatgaagtggatagcactcttataactttcttgctcgaagtttcttgataccaaggcaagtagcttataagaagatttggactttggtggcataaatggatgaactaactggcggcttcaacgtggtgttgattacaggtgcttttattgctaactggagcagtttgtgtagtttctttctttctctttgagaattctctcagtttcttgggcaggctcttgagctaccaagtatgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggtaacattattcctatttgattcttgttagaaattctatatgttgaacatacttgcctcaatgggaaacttggttgcgtctcctctctgaaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcaggtactgtatctctaagcatgtttagtatcttctgtgataaaatctttctccagtgtgttaacttttgcattctttttcttgcatgcttgcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattaggtaatttgatccagtgatcaatacatgttccacaaacaattgaagaaagagtttgacatatttttctcatatttccagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgaggtaaggcccctactctaatggttagttccaatttattggttcatttctctgtgtttgataagattttgcctctgctgtgcatttcaatattatatctaaactgtgctgcttatagcttatcaaactactcttgctcataaagatgcaacttttatgcagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgagtcataataaatgataacatattattgttctgtggactgaaaatctgaaatcctcctttcttgagaaatacaacctcaattttatggtgatattgacaagaaggaagagcgcatataatttaaagttaaaacacggaaaataatagaagtatgtgagcagaacgcaggagcaaggggtggaactcataacaagttcaaacttcatgcatgttcataattccccaatgcaagtatggcaagaaaaatagacaacttttaaaagtatgctaatggaactgcatgcattgtgtaaacttgcttttgtcaagcatgttggaacatactatatatttgctgctgatgatgaaaaatttgcttgagtactcttctattgctaatggacgcttaccattccctgttttgtcttcaaatgcatcaggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactggttggtaaaatcagcaatggaattttctatcgtgaattttttataattattgtaacattaattactcccatgtctttttattctgttgagataccatgtgacaatgtctacaattgcatattttgcattctttggtagatcagttcccttcttaagatccccaaagggaaactggagtatcaataacttactccctcaaataaaaaaaaagtgatactttaggcataaacacattcttcaaaacaaaactttgaatagcagtttcttttgtaatatattaataaagcatggaaaaattataatgtgaagaatgtacattcagggacaaatatactcatcatttttgcatggttgatcttaatgtccatagagagattgatcagaccaaaagtttatttttgacaggagtgagtatggattataataggagtgaatggcaatatgctgtgaaagtgagcattttatatttgtttactactagcatttggtgacaaagcaaatgatgtatattgtaaatcaaagtaaacaagtagaacatgtcctgtggcgacaagacataacctacaaattaaagttagcctgattgtttattttactgggagcacattttttctagattctagagtacatattattaaaaaaggtgtgatatgtttgcaaattactttttaatggaagtttgtaaattactttgaaacttgcattcagtttttttgcttcttaaaaaacatagtgcatatgggctttatgagctcatactatgcatgtttttctgcattggagtgggacctttatctttttcttgcaactcggatgctaatgcttattataaacacaagttactaacatgtgcttgggtcattcagcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcataggtaagctcttatgatcctgtccactgttgtaccatgtgtactttccaaatgattgctaacatgctatttttcgtcagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcgggtatatcatacatctctctggctgttgtttcagtgctatcaatcaagatgacacatcatgcaatattatcataacagcacattatgtgttttgaatattttattgattgtcatagctttgttttatgttttagcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcaggtatgcagtgtttcctgcacctacttgctacctattctgtatttgatgtattcaatttcctttttatcagttttctcgaaatatatgctaataggcttgtcactaaacggctacattatccagactctgttctgaatttttttttcattaagtactgttagctgccagattacatcatcatgactaaagtagtttagctacaatattgcaaagctagatttgtctagcctctattgttttgaatagaaaaggcaggtatgcagtgtttcctgcacctacttgctacctattctgtatttgatgtattcaatttcctttttatcagttttctcgaaatatatgctaataggcttgtcactaaacggctacattatccagactctgttctgaatttttttttcattaagtactgttagctgccagattacatcatcatgactaaagtagtttagctacaatattgcaaagctagatttgtctagcctctattgttttgaatagaaaattaaatcttttggtcatttttttgtaaccttttaggagtctctgtatgcttgtcgtcaacatgctatgataagtttatatcagttcttttggagctatgatgtattcttgtatattttatgactgctaggatattatgttaaaactgttttaaaggtatcatcaaacaagttaatatagatattgctacctaccaaagttggaatttgattatgtcgtaaaatatatgcaccctacatgttttagcttcatattgacttatattttcatatgtgctccttcaactgcacaaagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggtatggtcattcctgcttataagataccagaataatgctatgaagaccgtccttggttgtcagcattcttatacacaatgataatctttgtttttcatgggtaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggtcaagatattgttattgttatatttacatacttgttatcatatttaaaagcttcatattgtatgtacctttgaaaaaaaaagttccatactgtagcttcttttgttctcatttcttggtttcaaagcatcagcatcattaccgatataaaatatacccatttcctcagtcatcttccccgactcttgtatctttcacaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagaggtaacaccttcattcatgaggtccttaggtgcaaatcccagtgaagaataatcagttctactttgtatggttcatctcgcttactgaccggttcatctcgggatcaattacgagcagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaaggcatgtcctcttctctgatctcactcacgttcatgccaaacttactcatcatcatccatgatccatcatcctgactgaatcttgatatgcagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctgacaaggccacaccatgactgtcactgcaagcagctgctgtaggatcagcgagtaggaaaggatgacttgccggccagtttcctgagatgtgatttaagactgatattagctgatgttcccaagcatgatacggggcttgctcccaagatgtgatttttactttacttttggttaatgtgctaccgctgacattagatgttcaagcatatgaaaactgcttgtgctgtaacttgtatgtctgtaacgtaaaaatctttccatatggtaaccggggaagcagaggtatatccttttctgctgttgtatctatttccagtttcctgctgaacttgcaaatgaatattcagcaatttccgttttgctc</dnaseqindica>|<br />
Link = [http://www.ncbi.nlm.nih.gov/nuccore/NM_001048755.1 RefSeq:Os01g0182600]|<br />
}}<br />
[[Category:Genes]]<br />
[[Category:Japonica mRNA]]<br />
[[Category:Oryza Sativa Japonica Group]]<br />
[[Category:Japonica Genes]]<br />
[[Category:Japonica Chromosome 1]]<br />
[[Category:Chromosome 1]]</div>Mikahttps://ngdc.cncb.ac.cn/ricewiki/index.php?title=Os01g0182600&diff=183710Os01g01826002014-06-10T15:01:58Z<p>Mika: </p>
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==Annotated Information==<br />
===Function===<br />
*''OsGI'',a ''GI'' ortholog, regulates flowering time in response to photoperiodic conditions. Mutations in this gene delay flowering in inductive SDs (short-days),whereas flowering is marginally induced under LDs(long-days).In comparison, over expression of ''OsGI'' in transgenic rice causes late flowering in both SDs and LDs.In addition, ''OsGI'' has function to regulate ''Hd1'', a gene that also shows circadian rhythm<ref name="ref1" />. ''Os-GI'' affects gene expression of approximately 75% of genes among 27,201 genes and is required for strong amplitudes and proper phase-setting of global gene expression under natural field conditions<ref name="ref2" />. <br />
*The cDNA of ''OsGI'' firstly found in ''se5'' mutant that is completely photoperiod-insensitive, the expression patterns of rice and Arabidopsis ''GI'' gene are both photoperiodically regulated and are very similar under LD and SD conditions(FIG1)<ref name="ref3" />. ''GI'' is required for phase-dependent ''CO''transcription through the interaction with FKF1, which leads to cycling DNA binding with one finger (CDF) degradation and consequent derepression of CO. In addition, it has been known that GI protein interacts with ZEITLUPE (ZTL) F-box protein to control the diurnal rhythm of TOC1 protein in regulation of the Arabidopsis circadian clock<ref name="ref4" />. <br />
*''Os-GI'' also has function to maintain bioactive GA level through the regulation of the GA-deactivating enzyme genes in rice, ''Os-GI'' represses major ''OsGA2ox'' genes in order to maintain a proper bioactive GA level, ''osgi-1'' plants showed a semi-dwarf phenotype with elevated ''OsGA2ox'' gene expression(FIG2) <ref name="ref4" />.<br />
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===Expression===<br />
*''GI'' encodes a nuclear protein that defines the proper amplitude and period length of circadian rhythms. ''GI'' expression is regulated by the circadian clock, with a peak in transcript levels at 8 to 10 h after dawn. The ''GI'' protein acts between the circadian oscillator and ''CO'' to promote flowering by increasing ''CO'' and ''FT'' mRNA abundance<ref name="ref1" />. The expression of the ''OsGI'' messenger RNA was circadian-controlled and that its temporal expression pattern was very similar to that of ''GI'' under both SD and LD conditions. Decreased ''OsGI'' expression led to early flowering under LD conditions and late flowering under SD conditions relative to the wild type, and that this early flowering phenotype under LD conditions was the opposite of that of the Arabidopsis ''gi'' mutants, confirming the opposite role of the ''GI'' gene in flowering under LD conditions in rice and Arabidopsis<ref name="ref5" />.<br />
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===Mutation===<br />
*The single ''osgi-1'' mutation extended the time to flowering only under SD conditions. Under LD conditions the single ''hd1'' mutation induced early flowering, but ''osgi-1'' plants exhibited no phenotypic change<ref name="ref4" /><br />
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===Evolution===<br />
* There is some evolutionary conservation between rice circadian clock–related genes and those associated with Arabidopsis circadian clocks, where the circadian clock consists of interlocked subloops, it is also possible that ''Os-GI'' is a member of a subloop in rice circadian clocks<ref name="ref4" />.But there is a divergence in the photoperiodic pathways of these two species arises from the molecular mechanisms controlling the expression of ''FT/Hd3a'' by ''CO/Hd1(Se1)''<ref name="ref5" />(figure 3)<br />
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==Labs working on this gene==<br />
Please input related labs here.<br />
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==Knowledge Extension==<br />
*The rice orthologues of the two key regulators of flowering time in Arabidopsis, ''CO'' and ''FT'', were recently shown to be important in the photoperiodic control of flowering in rice. The rice ''Hd1(Se1)''gene, which is an orthologue of the Arabidopsis ''CO'' gene, was required for the suppression of flowering under LD conditions and for the promotion of flowering under SD conditions. Furthermore, the rice ''Hd3a'' gene, an orthologue of the Arabidopsis ''FT'' gene, was shown to be an activator of flowering in rice. It was previously demonstrated that ''CO'' controls the flowering time by integrating signals from the circadian clock and light to regulate ''FT'' expression in Arabidopsis<ref name="ref5" />.Unlike Arabidopsis ''FT'', expression of ''Hd3a'' mRNA is more or less opposite to that of ''Hd1'' under inductive short-day conditions<ref name="ref6" />.<br />
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==References==<br />
Please input cited references here.<br />
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==Structured Information==<br />
{{JaponicaGene|<br />
GeneName = Os01g0182600|<br />
Description = GIGANTEA protein|<br />
Version = NM_001048755.1 GI:115434923 GeneID:4325329|<br />
Length = 9083 bp|<br />
Definition = Oryza sativa Japonica Group Os01g0182600, complete gene.|<br />
Source = Oryza sativa Japonica Group<br />
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ORGANISM Oryza sativa Japonica Group<br />
Eukaryota; Viridiplantae; Streptophyta; Embryophyta; Tracheophyta;<br />
Spermatophyta; Magnoliophyta; Liliopsida; Poales; Poaceae; BEP<br />
clade; Ehrhartoideae; Oryzeae; Oryza.<br />
|<br />
Chromosome = [[:category:Japonica Chromosome 1|Chromosome 1]]|<br />
AP = Chromosome 1:4328362..4337444|<br />
CDS = 4328725..4329030,4329123..4329215,4329333..4329551,4329752..4329820,4329925..4331421<br>,4332220..4332429,4332563..4332713,4332791..4333031,4333898..4333956<br>,4334379..4334653,4334813..4334955,4335034..4335088,4335181..4335258<br>,4335351..4335437|<br />
GCID = <gbrowseImage1><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage1>|<br />
GSID = <gbrowseImage2><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage2>|<br />
CDNA = <cdnaseq>atgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactgcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcatagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcggcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctga</cdnaseq>|<br />
AA = <aaseq>MSASNEKWIDGLQFSSLFWPPPQDSQQKQAQILAYVEYFGQFTA DSEQFPEDIAQLIQSCYPSKEKRLVDEVLATFVLHHPEHGHAVVHPILSRIIDGTLSY DRNGFPFMSFISLFSHTSEKEYSEQWALACGEILRVLTHYNRPIFKVDHQHSEAECSS TSDQASSCESMEKRANGSPRNEPDRKPLRPLSPWITDILLAAPLGIRSDYFRWCGGVM GKYAAGGELKPPTTAYSRGSGKHPQLMPSTPRWAVANGAGVILSVCDEEVARYETANL TAAAVPALLLPPPTTPLDEHLVAGLPPLEPYARLFHRYYAIATPSATQRLLFGLLEAP PSWAPDALDAAVQLVELLRAAEDYDSGMRLPKNWMHLHFLRAIGTAMSMRAGIAADTS AALLFRILSQPTLLFPPLRHAEGVELHHEPLGGYVSSYKRQLEVPASEATIDATAQGI ASMLCAHGPDVEWRICTIWEAAYGLLPLSSSAVDLPEIVVAAPLQPPTLSWSLYLPLL KVFEYLPRGSPSEACLMRIFVATVEAILRRTFPSETSEQSRKPRSQSKNLAVAELRTM IHSLFVESCASMDLASRLLFVVLTVCVSHQALPGGSKRPTGSDNHSSEEVTNDSRLTN GRNRCKKRQGPVATFDSYVLAAVCALSCELQLFPFISKNGNHSNLKDSIKIVIPGKTT GISNELHNSISSAILHTRRILGILEALFSLKPSSVGTSWSYSSNEIVAAAMVAAHVSE LFRRSRPCLNALSALKQCKWDAEISTRASSLYHLIDLHGKTVTSIVNKAEPLEAHLTL TPVKKDEPPIEEKNINSSDGGALEKKDASRSHRKNGFARPLLKCAEDVILNGDVASTS GKAIASLQVEASDLANFLTMDRNGGYRGSQTLLRSVLSEKQELCFSVVSLLWQKLIAS PEMQMSAESTSAHQGWRKVVDALCDIVSASPTKASAAIVLQAEKDLQPWIARDDEQGQ KMWRVNQRIVKLIAELMRNHDSPEALVILASASDLLLRATDGMLVDGEACTLPQLELL EVTARAVHLIVEWGDSGVSVADGLSNLLKCRLSTTIRCLSHPSAHVRALSMSVLRDIL NSGQINSSKLIQGEHRNGIQSPTYQCLAASIINWQADVERCIEWEAHSRRATGLTLAF LTAAAKELGCPLTC</aaseq>|<br />
DNA = <dnaseqindica>8415..8720#8230..8322#7894..8112#7625..7693#6024..7520#5016..5225#4732..4882#4414..4654#3489..3547#2792..3066#2490..2632#2357..2411#2187..2264#2008..2094#ccgagccgtttctctccctctctctctcctcctccttcccctgcgcctcctccaccgcccgtgtcgccgccgcgcgcgtctatcccctcgccgccgccgccgtctacgccgcgtgaggtccggattggttgaaaggctgatcgaggtaatgcttttttatttttcccgtgcggttttagtttttctgtgctgtgcttggtggttttttctttaaaactatttttaggctggttttgatgatggcctttctaaggttgtcgtgtatggggggcgaggtggttcgggtggatttgtggtgttatgttgaggggatttttgtttggggggagagagttttggtggtagatgcggttgggattttgaagacgtgaaagggaaattagttttgaaatttgacggcgaggttgagctgattagtattgtaataattcgtggcatcggatagatcaacgcgagatggatgcttattactgcgaatttcaggaggtttttgtggtttttttaggcgagtagcgttgattctagctggattttgctgcttcgtggaggcgtttagatgtggtgcatacggctgtggatgtaatttgtgcggggataattactgaaaattaattcgcagtaagctagctctgctttaatattttttcccctgaaacagtaacaagggatcccttcctcatttcgttaacagtggaagtgaatacagacaccaaccgatgagcagagacacctgtaattaaacaccaaaagaagaacctgttttgttcctatatattttccatttctatgctttaggtaatatactttctggttcggtgtcatagtacccgaaatagaatctttgcttgccaaaaaaagaaagaataacaagatagaacaacatcattaaattctgtcttagacaatacatttataatcgaccaacaaaaacagaccgatgagtgacgtggaataacaaatgaagtataaatgtcacgaaaaacacatctattgattttgtgtgataagccataggagcctacgaggaaccaagtgaatggttctgtacaccacatccattagcattaccataaatttgcctatttctatgatgccttccttaacatatttatcgacatgtgagcttgcaccataactaaatgttatggtaaactttaccttataaatgattgcaacatgtaccttttcatttcgataattaaaaacgagaaagcttccacctttttcctagatatattttttcgtttgattatgaatttagttgtacttgacaataaaataatgcgcttgtaagattatgcaattttcgttgcattacctccctctcttttcatttgtcagtttttttctactccctccgtttcatattttaagtcgtttgacttttttcctagtcagactttgttaagtttgacaaagtttatagtaaaatttagcaatatctaaaatattaaattggtttcatcaaatctaacattgaatatattttgataatatgttttttttgtgttgaaaatacttctatatttttctataaacttagtcaaacttggaagaaatttcagtaagaaaaaagtcaaacggcttataatatgaaacggagggagtaaatttttttttgcattaattaaaattcatgtattttcctccttttttggtttttgttgtgcgatatattacagtgtagagactctgaagctcatatggttggcattggttgatatgataacgtgtcacgttgtgagatgcagacccccgttatgggggttgattgaatgaagtggatagcactcttataactttcttgctcgaagtttcttgataccaaggcaagtagcttataagaagatttggactttggtggcataaatggatgaactaactggcggcttcaacgtggtgttgattacaggtgcttttattgctaactggagcagtttgtgtagtttctttctttctctttgagaattctctcagtttcttgggcaggctcttgagctaccaagtatgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggtaacattattcctatttgattcttgttagaaattctatatgttgaacatacttgcctcaatgggaaacttggttgcgtctcctctctgaaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcaggtactgtatctctaagcatgtttagtatcttctgtgataaaatctttctccagtgtgttaacttttgcattctttttcttgcatgcttgcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattaggtaatttgatccagtgatcaatacatgttccacaaacaattgaagaaagagtttgacatatttttctcatatttccagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgaggtaaggcccctactctaatggttagttccaatttattggttcatttctctgtgtttgataagattttgcctctgctgtgcatttcaatattatatctaaactgtgctgcttatagcttatcaaactactcttgctcataaagatgcaacttttatgcagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgagtcataataaatgataacatattattgttctgtggactgaaaatctgaaatcctcctttcttgagaaatacaacctcaattttatggtgatattgacaagaaggaagagcgcatataatttaaagttaaaacacggaaaataatagaagtatgtgagcagaacgcaggagcaaggggtggaactcataacaagttcaaacttcatgcatgttcataattccccaatgcaagtatggcaagaaaaatagacaacttttaaaagtatgctaatggaactgcatgcattgtgtaaacttgcttttgtcaagcatgttggaacatactatatatttgctgctgatgatgaaaaatttgcttgagtactcttctattgctaatggacgcttaccattccctgttttgtcttcaaatgcatcaggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactggttggtaaaatcagcaatggaattttctatcgtgaattttttataattattgtaacattaattactcccatgtctttttattctgttgagataccatgtgacaatgtctacaattgcatattttgcattctttggtagatcagttcccttcttaagatccccaaagggaaactggagtatcaataacttactccctcaaataaaaaaaaagtgatactttaggcataaacacattcttcaaaacaaaactttgaatagcagtttcttttgtaatatattaataaagcatggaaaaattataatgtgaagaatgtacattcagggacaaatatactcatcatttttgcatggttgatcttaatgtccatagagagattgatcagaccaaaagtttatttttgacaggagtgagtatggattataataggagtgaatggcaatatgctgtgaaagtgagcattttatatttgtttactactagcatttggtgacaaagcaaatgatgtatattgtaaatcaaagtaaacaagtagaacatgtcctgtggcgacaagacataacctacaaattaaagttagcctgattgtttattttactgggagcacattttttctagattctagagtacatattattaaaaaaggtgtgatatgtttgcaaattactttttaatggaagtttgtaaattactttgaaacttgcattcagtttttttgcttcttaaaaaacatagtgcatatgggctttatgagctcatactatgcatgtttttctgcattggagtgggacctttatctttttcttgcaactcggatgctaatgcttattataaacacaagttactaacatgtgcttgggtcattcagcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcataggtaagctcttatgatcctgtccactgttgtaccatgtgtactttccaaatgattgctaacatgctatttttcgtcagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtac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/>
Link = [http://www.ncbi.nlm.nih.gov/nuccore/NM_001048755.1 RefSeq:Os01g0182600]|<br />
}}<br />
[[Category:Genes]]<br />
[[Category:Japonica mRNA]]<br />
[[Category:Oryza Sativa Japonica Group]]<br />
[[Category:Japonica Genes]]<br />
[[Category:Japonica Chromosome 1]]<br />
[[Category:Chromosome 1]]</div>Mikahttps://ngdc.cncb.ac.cn/ricewiki/index.php?title=Os01g0182600&diff=183709Os01g01826002014-06-10T15:00:46Z<p>Mika: </p>
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<div>Please input one-sentence summary here.<br />
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==Annotated Information==<br />
===Function===<br />
*''OsGI'',a ''GI'' ortholog, regulates flowering time in response to photoperiodic conditions. Mutations in this gene delay flowering in inductive SDs (short-days),whereas flowering is marginally induced under LDs(long-days).In comparison, over expression of ''OsGI'' in transgenic rice causes late flowering in both SDs and LDs.In addition, ''OsGI'' has function to regulate ''Hd1'', a gene that also shows circadian rhythm<ref name="ref1" />. ''Os-GI'' affects gene expression of approximately 75% of genes among 27,201 genes and is required for strong amplitudes and proper phase-setting of global gene expression under natural field conditions<ref name="ref2" />. <br />
*The cDNA of ''OsGI'' firstly found in ''se5'' mutant that is completely photoperiod-insensitive, the expression patterns of rice and Arabidopsis ''GI'' gene are both photoperiodically regulated and are very similar under LD and SD conditions(FIG1)<ref name="ref3" />. ''GI'' is required for phase-dependent ''CO''transcription through the interaction with FKF1, which leads to cycling DNA binding with one finger (CDF) degradation and consequent derepression of CO. In addition, it has been known that GI protein interacts with ZEITLUPE (ZTL) F-box protein to control the diurnal rhythm of TOC1 protein in regulation of the Arabidopsis circadian clock<ref name="ref4" />. <br />
*''Os-GI'' also has function to maintain bioactive GA level through the regulation of the GA-deactivating enzyme genes in rice, ''Os-GI'' represses major ''OsGA2ox'' genes in order to maintain a proper bioactive GA level, ''osgi-1'' plants showed a semi-dwarf phenotype with elevated ''OsGA2ox'' gene expression(FIG2) <ref name="ref4" />.<br />
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===Expression===<br />
*''GI'' encodes a nuclear protein that defines the proper amplitude and period length of circadian rhythms. ''GI'' expression is regulated by the circadian clock, with a peak in transcript levels at 8 to 10 h after dawn. The ''GI'' protein acts between the circadian oscillator and ''CO'' to promote flowering by increasing ''CO'' and ''FT'' mRNA abundance<ref name="ref1" />. The expression of the ''OsGI'' messenger RNA was circadian-controlled and that its temporal expression pattern was very similar to that of ''GI'' under both SD and LD conditions. Decreased ''OsGI'' expression led to early flowering under LD conditions and late flowering under SD conditions relative to the wild type, and that this early flowering phenotype under LD conditions was the opposite of that of the Arabidopsis ''gi'' mutants, confirming the opposite role of the ''GI'' gene in flowering under LD conditions in rice and Arabidopsis<ref name="ref5" />.<br />
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===Mutation===<br />
*The single ''osgi-1'' mutation extended the time to flowering only under SD conditions. Under LD conditions the single ''hd1'' mutation induced early flowering, but ''osgi-1'' plants exhibited no phenotypic change<ref name="ref4" /><br />
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===Evolution===<br />
* There is some evolutionary conservation between rice circadian clock–related genes and those associated with Arabidopsis circadian clocks, where the circadian clock consists of interlocked subloops, it is also possible that ''Os-GI'' is a member of a subloop in rice circadian clocks<ref name="ref4" />.But there is a divergence in the photoperiodic pathways of these two species arises from the molecular mechanisms controlling the expression of ''FT/Hd3a'' by ''CO/Hd1(Se1)''<ref name="ref5" />(figure 3)<br />
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==Labs working on this gene==<br />
Please input related labs here.<br />
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==Knowledge Extension==<br />
*The rice orthologues of the two key regulators of flowering time in Arabidopsis, ''CO'' and ''FT'', were recently shown to be important in the photoperiodic control of flowering in rice. The rice ''Hd1(Se1)''gene, which is an orthologue of the Arabidopsis ''CO'' gene, was required for the suppression of flowering under LD conditions and for the promotion of flowering under SD conditions. Furthermore, the rice ''Hd3a'' gene, an orthologue of the Arabidopsis ''FT'' gene, was shown to be an activator of flowering in rice. It was previously demonstrated that ''CO'' controls the flowering time by integrating signals from the circadian clock and light to regulate ''FT'' expression in Arabidopsis<ref name="ref5" />.Unlike Arabidopsis ''FT'', expression of ''Hd3a'' mRNA is more or less opposite to that of ''Hd1'' under inductive short-day conditions<ref name="ref6" />.<br />
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==References==<br />
Please input cited references here.<br />
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==Structured Information==<br />
{{JaponicaGene|<br />
GeneName = Os01g0182600|<br />
Description = GIGANTEA protein|<br />
Version = NM_001048755.1 GI:115434923 GeneID:4325329|<br />
Length = 9083 bp|<br />
Definition = Oryza sativa Japonica Group Os01g0182600, complete gene.|<br />
Source = Oryza sativa Japonica Group<br />
<br />
ORGANISM Oryza sativa Japonica Group<br />
Eukaryota; Viridiplantae; Streptophyta; Embryophyta; Tracheophyta;<br />
Spermatophyta; Magnoliophyta; Liliopsida; Poales; Poaceae; BEP<br />
clade; Ehrhartoideae; Oryzeae; Oryza.<br />
|<br />
Chromosome = [[:category:Japonica Chromosome 1|Chromosome 1]]|<br />
AP = Chromosome 1:4328362..4337444|<br />
CDS = 4328725..4329030,4329123..4329215,4329333..4329551,4329752..4329820,4329925..4331421<br>,4332220..4332429,4332563..4332713,4332791..4333031,4333898..4333956<br>,4334379..4334653,4334813..4334955,4335034..4335088,4335181..4335258<br>,4335351..4335437|<br />
GCID = <gbrowseImage1><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage1>|<br />
GSID = <gbrowseImage2><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage2>|<br />
CDNA = <cdnaseq>atgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactgcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcatagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcggcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctga</cdnaseq>|<br />
AA = <aaseq>MSASNEKWIDGLQFSSLFWPPPQDSQQKQAQILAYVEYFGQFTA DSEQFPEDIAQLIQSCYPSKEKRLVDEVLATFVLHHPEHGHAVVHPILSRIIDGTLSY DRNGFPFMSFISLFSHTSEKEYSEQWALACGEILRVLTHYNRPIFKVDHQHSEAECSS TSDQASSCESMEKRANGSPRNEPDRKPLRPLSPWITDILLAAPLGIRSDYFRWCGGVM GKYAAGGELKPPTTAYSRGSGKHPQLMPSTPRWAVANGAGVILSVCDEEVARYETANL TAAAVPALLLPPPTTPLDEHLVAGLPPLEPYARLFHRYYAIATPSATQRLLFGLLEAP PSWAPDALDAAVQLVELLRAAEDYDSGMRLPKNWMHLHFLRAIGTAMSMRAGIAADTS AALLFRILSQPTLLFPPLRHAEGVELHHEPLGGYVSSYKRQLEVPASEATIDATAQGI ASMLCAHGPDVEWRICTIWEAAYGLLPLSSSAVDLPEIVVAAPLQPPTLSWSLYLPLL KVFEYLPRGSPSEACLMRIFVATVEAILRRTFPSETSEQSRKPRSQSKNLAVAELRTM IHSLFVESCASMDLASRLLFVVLTVCVSHQALPGGSKRPTGSDNHSSEEVTNDSRLTN GRNRCKKRQGPVATFDSYVLAAVCALSCELQLFPFISKNGNHSNLKDSIKIVIPGKTT GISNELHNSISSAILHTRRILGILEALFSLKPSSVGTSWSYSSNEIVAAAMVAAHVSE LFRRSRPCLNALSALKQCKWDAEISTRASSLYHLIDLHGKTVTSIVNKAEPLEAHLTL TPVKKDEPPIEEKNINSSDGGALEKKDASRSHRKNGFARPLLKCAEDVILNGDVASTS GKAIASLQVEASDLANFLTMDRNGGYRGSQTLLRSVLSEKQELCFSVVSLLWQKLIAS PEMQMSAESTSAHQGWRKVVDALCDIVSASPTKASAAIVLQAEKDLQPWIARDDEQGQ KMWRVNQRIVKLIAELMRNHDSPEALVILASASDLLLRATDGMLVDGEACTLPQLELL EVTARAVHLIVEWGDSGVSVADGLSNLLKCRLSTTIRCLSHPSAHVRALSMSVLRDIL NSGQINSSKLIQGEHRNGIQSPTYQCLAASIINWQADVERCIEWEAHSRRATGLTLAF LTAAAKELGCPLTC</aaseq>|<br />
DNA = <dnaseqindica>8415..8720#8230..8322#7894..8112#7625..7693#6024..7520#5016..5225#4732..4882#4414..4654#3489..3547#2792..3066#2490..2632#2357..2411#2187..2264#2008..2094#ccgagccgtttctctccctctctctctcctcctccttcccctgcgcctcctccaccgcccgtgtcgccgccgcgcgcgtctatcccctcgccgccgccgccgtctacgccgcgtgaggtccggattggttgaaaggctgatcgaggtaatgcttttttatttttcccgtgcggttttagtttttctgtgctgtgcttggtggttttttctttaaaactatttttaggctggttttgatgatggcctttctaaggttgtcgtgtatggggggcgaggtggttcgggtggatttgtggtgttatgttgaggggatttttgtttggggggagagagttttggtggtagatgcggttgggattttgaagacgtgaaagggaaattagttttgaaatttgacggcgaggttgagctgattagtattgtaataattcgtggcatcggatagatcaacgcgagatggatgcttattactgcgaatttcaggaggtttttgtggtttttttaggcgagtagcgttgattctagctggattttgctgcttcgtggaggcgtttagatgtggtgcatacggctgtggatgtaatttgtgcggggataattactgaaaattaattcgcagtaagctagctctgctttaatattttttcccctgaaacagtaacaagggatcccttcctcatttcgttaacagtggaagtgaatacagacaccaaccgatgagcagagacacctgtaattaaacaccaaaagaagaacctgttttgttcctatatattttccatttctatgctttaggtaatatactttctggttcggtgtcatagtacccgaaatagaatctttgcttgccaaaaaaagaaagaataacaagatagaacaacatcattaaattctgtcttagacaatacatttataatcgaccaacaaaaacagaccgatgagtgacgtggaataacaaatgaagtataaatgtcacgaaaaacacatctattgattttgtgtgataagccataggagcctacgaggaaccaagtgaatggttctgtacaccacatccattagcattaccataaatttgcctatttctatgatgccttccttaacatatttatcgacatgtgagcttgcaccataactaaatgttatggtaaactttaccttataaatgattgcaacatgtaccttttcatttcgataattaaaaacgagaaagcttccacctttttcctagatatattttttcgtttgattatgaatttagttgtacttgacaataaaataatgcgcttgtaagattatgcaattttcgttgcattacctccctctcttttcatttgtcagtttttttctactccctccgtttcatattttaagtcgtttgacttttttcctagtcagactttgttaagtttgacaaagtttatagtaaaatttagcaatatctaaaatattaaattggtttcatcaaatctaacattgaatatattttgataatatgttttttttgtgttgaaaatacttctatatttttctataaacttagtcaaacttggaagaaatttcagtaagaaaaaagtcaaacggcttataatatgaaacggagggagtaaatttttttttgcattaattaaaattcatgtattttcctccttttttggtttttgttgtgcgatatattacagtgtagagactctgaagctcatatggttggcattggttgatatgataacgtgtcacgttgtgagatgcagacccccgttatgggggttgattgaatgaagtggatagcactcttataactttcttgctcgaagtttcttgataccaaggcaagtagcttataagaagatttggactttggtggcataaatggatgaactaactggcggcttcaacgtggtgttgattacaggtgcttttattgctaactggagcagtttgtgtagtttctttctttctctttgagaattctctcagtttcttgggcaggctcttgagctaccaagtatgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggtaacattattcctatttgattcttgttagaaattctatatgttgaacatacttgcctcaatgggaaacttggttgcgtctcctctctgaaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcaggtactgtatctctaagcatgtttagtatcttctgtgataaaatctttctccagtgtgttaacttttgcattctttttcttgcatgcttgcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattaggtaatttgatccagtgatcaatacatgttccacaaacaattgaagaaagagtttgacatatttttctcatatttccagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgaggtaaggcccctactctaatggttagttccaatttattggttcatttctctgtgtttgataagattttgcctctgctgtgcatttcaatattatatctaaactgtgctgcttatagcttatcaaactactcttgctcataaagatgcaacttttatgcagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgagtcataataaatgataacatattattgttctgtggactgaaaatctgaaatcctcctttcttgagaaatacaacctcaattttatggtgatattgacaagaaggaagagcgcatataatttaaagttaaaacacggaaaataatagaagtatgtgagcagaacgcaggagcaaggggtggaactcataacaagttcaaacttcatgcatgttcataattccccaatgcaagtatggcaagaaaaatagacaacttttaaaagtatgctaatggaactgcatgcattgtgtaaacttgcttttgtcaagcatgttggaacatactatatatttgctgctgatgatgaaaaatttgcttgagtactcttctattgctaatggacgcttaccattccctgttttgtcttcaaatgcatcaggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactggttggtaaaatcagcaatggaattttctatcgtgaattttttataattattgtaacattaattactcccatgtctttttattctgttgagataccatgtgacaatgtctacaattgcatattttgcattctttggtagatcagttcccttcttaagatccccaaagggaaactggagtatcaataacttactccctcaaataaaaaaaaagtgatactttaggcataaacacattcttcaaaacaaaactttgaatagcagtttcttttgtaatatattaataaagcatggaaaaattataatgtgaagaatgtacattcagggacaaatatactcatcatttttgcatggttgatcttaatgtccatagagagattgatcagaccaaaagtttatttttgacaggagtgagtatggattataataggagtgaatggcaatatgctgtgaaagtgagcattttatatttgtttactactagcatttggtgacaaagcaaatgatgtatattgtaaatcaaagtaaacaagtagaacatgtcctgtggcgacaagacataacctacaaattaaagttagcctgattgtttattttactgggagcacattttttctagattctagagtacatattattaaaaaaggtgtgatatgtttgcaaattactttttaatggaagtttgtaaattactttgaaacttgcattcagtttttttgcttcttaaaaaacatagtgcatatgggctttatgagctcatactatgcatgtttttctgcattggagtgggacctttatctttttcttgcaactcggatgctaatgcttattataaacacaagttactaacatgtgcttgggtcattcagcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcataggtaagctcttatgatcctgtccactgttgtaccatgtgtactttccaaatgattgctaacatgctatttttcgtcagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcgggtatatcatacatctctctggctgttgtttcagtgctatcaatcaagatgacacatcatgcaatattatcataacagcacattatgtgttttgaatattttattgattgtcatagctttgttttatgttttagcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcaggtatgcagtgtttcctgcacctacttgctacctattctgtatttgatgtattcaatttcctttttatcagttttctcgaaatatatgctaataggcttgtcactaaacggctacattatccagactctgttctgaatttttttttcattaagtactgttagctgccagattacatcatcatgactaaagtagtttagctacaatattgcaaagctagatttgtctagcctctattgttttgaatagaaaaggcaggtatgcagtgtttcctgcacctacttgctacctattctgtatttgatgtattcaatttcctttttatcagttttctcgaaatatatgctaataggcttgtcactaaacggctacattatccagactctgttctgaatttttttttcattaagtactgttagctgccagattacatcatcatgactaaagtagtttagctacaatattgcaaagctagatttgtctagcctctattgttttgaatagaaaattaaatcttttggtcatttttttgtaaccttttaggagtctctgtatgcttgtcgtcaacatgctatgataagtttatatcagttcttttggagctatgatgtattcttgtatattttatgactgctaggatattatgttaaaactgttttaaaggtatcatcaaacaagttaatatagatattgctacctaccaaagttggaatttgattatgtcgtaaaatatatgcaccctacatgttttagcttcatattgacttatattttcatatgtgctccttcaactgcacaaagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggtatggtcattcctgcttataagataccagaataatgctatgaagaccgtccttggttgtcagcattcttatacacaatgataatctttgtttttcatgggtaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggtcaagatattgttattgttatatttacatacttgttatcatatttaaaagcttcatattgtatgtacctttgaaaaaaaaagttccatactgtagcttcttttgttctcatttcttggtttcaaagcatcagcatcattaccgatataaaatatacccatttcctcagtcatcttccccgactcttgtatctttcacaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagaggtaacaccttcattcatgaggtccttaggtgcaaatcccagtgaagaataatcagttctactttgtatggttcatctcgcttactgaccggttcatctcgggatcaattacgagcagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaaggcatgtcctcttctctgatctcactcacgttcatgccaaacttactcatcatcatccatgatccatcatcctgactgaatcttgatatgcagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctgacaaggccacaccatgactgtcactgcaagcagctgctgtaggatcagcgagtaggaaaggatgacttgccggccagtttcctgagatgtgatttaagactgatattagctgatgttcccaagcatgatacggggcttgctcccaagatgtgatttttactttacttttggttaatgtgctaccgctgacattagatgttcaagcatatgaaaactgcttgtgctgtaacttgtatgtctgtaacgtaaaaatctttccatatggtaaccggggaagcagaggtatatccttttctgctgttgtatctatttccagtttcctgctgaacttgcaaatgaatattcagcaatttccgttttgctc</dnaseqindica>|<br />
Link = [http://www.ncbi.nlm.nih.gov/nuccore/NM_001048755.1 RefSeq:Os01g0182600]|<br />
}}<br />
[[Category:Genes]]<br />
[[Category:Japonica mRNA]]<br />
[[Category:Oryza Sativa Japonica Group]]<br />
[[Category:Japonica Genes]]<br />
[[Category:Japonica Chromosome 1]]<br />
[[Category:Chromosome 1]]</div>Mikahttps://ngdc.cncb.ac.cn/ricewiki/index.php?title=Os01g0182600&diff=183708Os01g01826002014-06-10T14:59:57Z<p>Mika: </p>
<hr />
<div>Please input one-sentence summary here.<br />
<br />
==Annotated Information==<br />
===Function===<br />
*''OsGI'',a ''GI'' ortholog, regulates flowering time in response to photoperiodic conditions. Mutations in this gene delay flowering in inductive SDs (short-days),whereas flowering is marginally induced under LDs(long-days).In comparison, over expression of ''OsGI'' in transgenic rice causes late flowering in both SDs and LDs.In addition, ''OsGI'' has function to regulate ''Hd1'', a gene that also shows circadian rhythm<ref name="ref1" />. ''Os-GI'' affects gene expression of approximately 75% of genes among 27,201 genes and is required for strong amplitudes and proper phase-setting of global gene expression under natural field conditions<ref name="ref2" />. <br />
*The cDNA of ''OsGI'' firstly found in ''se5'' mutant that is completely photoperiod-insensitive, the expression patterns of rice and Arabidopsis ''GI'' gene are both photoperiodically regulated and are very similar under LD and SD conditions(FIG1)<ref name="ref3" />. ''GI'' is required for phase-dependent ''CO''transcription through the interaction with FKF1, which leads to cycling DNA binding with one finger (CDF) degradation and consequent derepression of CO. In addition, it has been known that GI protein interacts with ZEITLUPE (ZTL) F-box protein to control the diurnal rhythm of TOC1 protein in regulation of the Arabidopsis circadian clock<ref name="ref4" />. <br />
*''Os-GI'' also has function to maintain bioactive GA level through the regulation of the GA-deactivating enzyme genes in rice, ''Os-GI'' represses major ''OsGA2ox'' genes in order to maintain a proper bioactive GA level, ''osgi-1'' plants showed a semi-dwarf phenotype with elevated ''OsGA2ox'' gene expression(FIG2) <ref name="ref4" />.<br />
<br />
===Expression===<br />
*''GI'' encodes a nuclear protein that defines the proper amplitude and period length of circadian rhythms. ''GI'' expression is regulated by the circadian clock, with a peak in transcript levels at 8 to 10 h after dawn. The ''GI'' protein acts between the circadian oscillator and ''CO'' to promote flowering by increasing ''CO'' and ''FT'' mRNA abundance<ref name="ref1" />. The expression of the ''OsGI'' messenger RNA was circadian-controlled and that its temporal expression pattern was very similar to that of ''GI'' under both SD and LD conditions. Decreased ''OsGI'' expression led to early flowering under LD conditions and late flowering under SD conditions relative to the wild type, and that this early flowering phenotype under LD conditions was the opposite of that of the Arabidopsis ''gi'' mutants, confirming the opposite role of the ''GI'' gene in flowering under LD conditions in rice and Arabidopsis<ref name="ref5" />.<br />
<br />
==Mutation==<br />
*The single ''osgi-1'' mutation extended the time to flowering only under SD conditions. Under LD conditions the single ''hd1'' mutation induced early flowering, but ''osgi-1'' plants exhibited no phenotypic change<ref name="ref4" /><br />
<br />
===Evolution===<br />
* There is some evolutionary conservation between rice circadian clock–related genes and those associated with Arabidopsis circadian clocks, where the circadian clock consists of interlocked subloops, it is also possible that ''Os-GI'' is a member of a subloop in rice circadian clocks<ref name="ref4" />.But there is a divergence in the photoperiodic pathways of these two species arises from the molecular mechanisms controlling the expression of ''FT/Hd3a'' by ''CO/Hd1(Se1)''<ref name="ref5" />(figure 3)<br />
<br />
==Labs working on this gene==<br />
Please input related labs here.<br />
<br />
==Knowledge Extension==<br />
*The rice orthologues of the two key regulators of flowering time in Arabidopsis, ''CO'' and ''FT'', were recently shown to be important in the photoperiodic control of flowering in rice. The rice ''Hd1(Se1)''gene, which is an orthologue of the Arabidopsis ''CO'' gene, was required for the suppression of flowering under LD conditions and for the promotion of flowering under SD conditions. Furthermore, the rice ''Hd3a'' gene, an orthologue of the Arabidopsis ''FT'' gene, was shown to be an activator of flowering in rice. It was previously demonstrated that ''CO'' controls the flowering time by integrating signals from the circadian clock and light to regulate ''FT'' expression in Arabidopsis<ref name="ref5" />.Unlike Arabidopsis ''FT'', expression of ''Hd3a'' mRNA is more or less opposite to that of ''Hd1'' under inductive short-day conditions<ref name="ref6" />.<br />
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==References==<br />
Please input cited references here.<br />
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==Structured Information==<br />
{{JaponicaGene|<br />
GeneName = Os01g0182600|<br />
Description = GIGANTEA protein|<br />
Version = NM_001048755.1 GI:115434923 GeneID:4325329|<br />
Length = 9083 bp|<br />
Definition = Oryza sativa Japonica Group Os01g0182600, complete gene.|<br />
Source = Oryza sativa Japonica Group<br />
<br />
ORGANISM Oryza sativa Japonica Group<br />
Eukaryota; Viridiplantae; Streptophyta; Embryophyta; Tracheophyta;<br />
Spermatophyta; Magnoliophyta; Liliopsida; Poales; Poaceae; BEP<br />
clade; Ehrhartoideae; Oryzeae; Oryza.<br />
|<br />
Chromosome = [[:category:Japonica Chromosome 1|Chromosome 1]]|<br />
AP = Chromosome 1:4328362..4337444|<br />
CDS = 4328725..4329030,4329123..4329215,4329333..4329551,4329752..4329820,4329925..4331421<br>,4332220..4332429,4332563..4332713,4332791..4333031,4333898..4333956<br>,4334379..4334653,4334813..4334955,4335034..4335088,4335181..4335258<br>,4335351..4335437|<br />
GCID = <gbrowseImage1><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage1>|<br />
GSID = <gbrowseImage2><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage2>|<br />
CDNA = <cdnaseq>atgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactgcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcatagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcggcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctga</cdnaseq>|<br />
AA = <aaseq>MSASNEKWIDGLQFSSLFWPPPQDSQQKQAQILAYVEYFGQFTA DSEQFPEDIAQLIQSCYPSKEKRLVDEVLATFVLHHPEHGHAVVHPILSRIIDGTLSY DRNGFPFMSFISLFSHTSEKEYSEQWALACGEILRVLTHYNRPIFKVDHQHSEAECSS TSDQASSCESMEKRANGSPRNEPDRKPLRPLSPWITDILLAAPLGIRSDYFRWCGGVM GKYAAGGELKPPTTAYSRGSGKHPQLMPSTPRWAVANGAGVILSVCDEEVARYETANL TAAAVPALLLPPPTTPLDEHLVAGLPPLEPYARLFHRYYAIATPSATQRLLFGLLEAP PSWAPDALDAAVQLVELLRAAEDYDSGMRLPKNWMHLHFLRAIGTAMSMRAGIAADTS AALLFRILSQPTLLFPPLRHAEGVELHHEPLGGYVSSYKRQLEVPASEATIDATAQGI ASMLCAHGPDVEWRICTIWEAAYGLLPLSSSAVDLPEIVVAAPLQPPTLSWSLYLPLL KVFEYLPRGSPSEACLMRIFVATVEAILRRTFPSETSEQSRKPRSQSKNLAVAELRTM IHSLFVESCASMDLASRLLFVVLTVCVSHQALPGGSKRPTGSDNHSSEEVTNDSRLTN GRNRCKKRQGPVATFDSYVLAAVCALSCELQLFPFISKNGNHSNLKDSIKIVIPGKTT GISNELHNSISSAILHTRRILGILEALFSLKPSSVGTSWSYSSNEIVAAAMVAAHVSE LFRRSRPCLNALSALKQCKWDAEISTRASSLYHLIDLHGKTVTSIVNKAEPLEAHLTL TPVKKDEPPIEEKNINSSDGGALEKKDASRSHRKNGFARPLLKCAEDVILNGDVASTS GKAIASLQVEASDLANFLTMDRNGGYRGSQTLLRSVLSEKQELCFSVVSLLWQKLIAS PEMQMSAESTSAHQGWRKVVDALCDIVSASPTKASAAIVLQAEKDLQPWIARDDEQGQ KMWRVNQRIVKLIAELMRNHDSPEALVILASASDLLLRATDGMLVDGEACTLPQLELL EVTARAVHLIVEWGDSGVSVADGLSNLLKCRLSTTIRCLSHPSAHVRALSMSVLRDIL NSGQINSSKLIQGEHRNGIQSPTYQCLAASIINWQADVERCIEWEAHSRRATGLTLAF LTAAAKELGCPLTC</aaseq>|<br />
DNA = <dnaseqindica>8415..8720#8230..8322#7894..8112#7625..7693#6024..7520#5016..5225#4732..4882#4414..4654#3489..3547#2792..3066#2490..2632#2357..2411#2187..2264#2008..2094#ccgagccgtttctctccctctctctctcctcctccttcccctgcgcctcctccaccgcccgtgtcgccgccgcgcgcgtctatcccctcgccgccgccgccgtctacgccgcgtgaggtccggattggttgaaaggctgatcgaggtaatgcttttttatttttcccgtgcggttttagtttttctgtgctgtgcttggtggttttttctttaaaactatttttaggctggttttgatgatggcctttctaaggttgtcgtgtatggggggcgaggtggttcgggtggatttgtggtgttatgttgaggggatttttgtttggggggagagagttttggtggtagatgcggttgggattttgaagacgtgaaagggaaattagttttgaaatttgacggcgaggttgagctgattagtattgtaataattcgtggcatcggatagatcaacgcgagatggatgcttattactgcgaatttcaggaggtttttgtggtttttttaggcgagtagcgttgattctagctggattttgctgcttcgtggaggcgtttagatgtggtgcatacggctgtggatgtaatttgtgcggggataattactgaaaattaattcgcagtaagctagctctgctttaatattttttcccctgaaacagtaacaagggatcccttcctcatttcgttaacagtggaagtgaatacagacaccaaccgatgagcagagacacctgtaattaaacaccaaaagaagaacctgttttgttcctatatattttccatttctatgctttaggtaatatactttctggttcggtgtcatagtacccgaaatagaatctttgcttgccaaaaaaagaaagaataacaagatagaacaacatcattaaattctgtcttagacaatacatttataatcgaccaacaaaaacagaccgatgagtgacgtggaataacaaatgaagtataaatgtcacgaaaaacacatctattgattttgtgtgataagccataggagcctacgaggaaccaagtgaatggttctgtacaccacatccattagcattaccataaatttgcctatttctatgatgccttccttaacatatttatcgacatgtgagcttgcaccataactaaatgttatggtaaactttaccttataaatgattgcaacatgtaccttttcatttcgataattaaaaacgagaaagcttccacctttttcctagatatattttttcgtttgattatgaatttagttgtacttgacaataaaataatgcgcttgtaagattatgcaattttcgttgcattacctccctctcttttcatttgtcagtttttttctactccctccgtttcatattttaagtcgtttgacttttttcctagtcagactttgttaagtttgacaaagtttatagtaaaatttagcaatatctaaaatattaaattggtttcatcaaatctaacattgaatatattttgataatatgttttttttgtgttgaaaatacttctatatttttctataaacttagtcaaacttggaagaaatttcagtaagaaaaaagtcaaacggcttataatatgaaacggagggagtaaatttttttttgcattaattaaaattcatgtattttcctccttttttggtttttgttgtgcgatatattacagtgtagagactctgaagctcatatggttggcattggttgatatgataacgtgtcacgttgtgagatgcagacccccgttatgggggttgattgaatgaagtggatagcactcttataactttcttgctcgaagtttcttgataccaaggcaagtagcttataagaagatttggactttggtggcataaatggatgaactaactggcggcttcaacgtggtgttgattacaggtgcttttattgctaactggagcagtttgtgtagtttctttctttctctttgagaattctctcagtttcttgggcaggctcttgagctaccaagtatgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggtaacattattcctatttgattcttgttagaaattctatatgttgaacatacttgcctcaatgggaaacttggttgcgtctcctctctgaaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcaggtactgtatctctaagcatgtttagtatcttctgtgataaaatctttctccagtgtgttaacttttgcattctttttcttgcatgcttgcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattaggtaatttgatccagtgatcaatacatgttccacaaacaattgaagaaagagtttgacatatttttctcatatttccagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgaggtaaggcccctactctaatggttagttccaatttattggttcatttctctgtgtttgataagattttgcctctgctgtgcatttcaatattatatctaaactgtgctgcttatagcttatcaaactactcttgctcataaagatgcaacttttatgcagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgagtcataataaatgataacatattattgttctgtggactgaaaatctgaaatcctcctttcttgagaaatacaacctcaattttatggtgatattgacaagaaggaagagcgcatataatttaaagttaaaacacggaaaataatagaagtatgtgagcagaacgcaggagcaaggggtggaactcataacaagttcaaacttcatgcatgttcataattccccaatgcaagtatggcaagaaaaatagacaacttttaaaagtatgctaatggaactgcatgcattgtgtaaacttgcttttgtcaagcatgttggaacatactatatatttgctgctgatgatgaaaaatttgcttgagtactcttctattgctaatggacgcttaccattccctgttttgtcttcaaatgcatcaggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactggttggtaaaatcagcaatggaattttctatcgtgaattttttataattattgtaacattaattactcccatgtctttttattctgttgagataccatgtgacaatgtctacaattgcatattttgcattctttggtagatcagttcccttcttaagatccccaaagggaaactggagtatcaataacttactccctcaaataaaaaaaaagtgatactttaggcataaacacattcttcaaaacaaaactttgaatagcagtttcttttgtaatatattaataaagcatggaaaaattataatgtgaagaatgtacattcagggacaaatatactcatcatttttgcatggttgatcttaatgtccatagagagattgatcagaccaaaagtttatttttgacaggagtgagtatggattataataggagtgaatggcaatatgctgtgaaagtgagcattttatatttgtttactactagcatttggtgacaaagcaaatgatgtatattgtaaatcaaagtaaacaagtagaacatgtcctgtggcgacaagacataacctacaaattaaagttagcctgattgtttattttactgggagcacattttttctagattctagagtacatattattaaaaaaggtgtgatatgtttgcaaattactttttaatggaagtttgtaaattactttgaaacttgcattcagtttttttgcttcttaaaaaacatagtgcatatgggctttatgagctcatactatgcatgtttttctgcattggagtgggacctttatctttttcttgcaactcggatgctaatgcttattataaacacaagttactaacatgtgcttgggtcattcagcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcataggtaagctcttatgatcctgtccactgttgtaccatgtgtactttccaaatgattgctaacatgctatttttcgtcagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcgggtatatcatacatctctctggctgttgtttcagtgctatcaatcaagatgacacatcatgcaatattatcataacagcacattatgtgttttgaatattttattgattgtcatagctttgttttatgttttagcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcaggtatgcagtgtttcctgcacctacttgctacctattctgtatttgatgtattcaatttcctttttatcagttttctcgaaatatatgctaataggcttgtcactaaacggctacattatccagactctgttctgaatttttttttcattaagtactgttagctgccagattacatcatcatgactaaagtagtttagctacaatattgcaaagctagatttgtctagcctctattgttttgaatagaaaaggcaggtatgcagtgtttcctgcacctacttgctacctattctgtatttgatgtattcaatttcctttttatcagttttctcgaaatatatgctaataggcttgtcactaaacggctacattatccagactctgttctgaatttttttttcattaagtactgttagctgccagattacatcatcatgactaaagtagtttagctacaatattgcaaagctagatttgtctagcctctattgttttgaatagaaaattaaatcttttggtcatttttttgtaaccttttaggagtctctgtatgcttgtcgtcaacatgctatgataagtttatatcagttcttttggagctatgatgtattcttgtatattttatgactgctaggatattatgttaaaactgttttaaaggtatcatcaaacaagttaatatagatattgctacctaccaaagttggaatttgattatgtcgtaaaatatatgcaccctacatgttttagcttcatattgacttatattttcatatgtgctccttcaactgcacaaagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggtatggtcattcctgcttataagataccagaataatgctatgaagaccgtccttggttgtcagcattcttatacacaatgataatctttgtttttcatgggtaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggtcaagatattgttattgttatatttacatacttgttatcatatttaaaagcttcatattgtatgtacctttgaaaaaaaaagttccatactgtagcttcttttgttctcatttcttggtttcaaagcatcagcatcattaccgatataaaatatacccatttcctcagtcatcttccccgactcttgtatctttcacaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagaggtaacaccttcattcatgaggtccttaggtgcaaatcccagtgaagaataatcagttctactttgtatggttcatctcgcttactgaccggttcatctcgggatcaattacgagcagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaaggcatgtcctcttctctgatctcactcacgttcatgccaaacttactcatcatcatccatgatccatcatcctgactgaatcttgatatgcagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctgacaaggccacaccatgactgtcactgcaagcagctgctgtaggatcagcgagtaggaaaggatgacttgccggccagtttcctgagatgtgatttaagactgatattagctgatgttcccaagcatgatacggggcttgctcccaagatgtgatttttactttacttttggttaatgtgctaccgctgacattagatgttcaagcatatgaaaactgcttgtgctgtaacttgtatgtctgtaacgtaaaaatctttccatatggtaaccggggaagcagaggtatatccttttctgctgttgtatctatttccagtttcctgctgaacttgcaaatgaatattcagcaatttccgttttgctc</dnaseqindica>|<br />
Link = [http://www.ncbi.nlm.nih.gov/nuccore/NM_001048755.1 RefSeq:Os01g0182600]|<br />
}}<br />
[[Category:Genes]]<br />
[[Category:Japonica mRNA]]<br />
[[Category:Oryza Sativa Japonica Group]]<br />
[[Category:Japonica Genes]]<br />
[[Category:Japonica Chromosome 1]]<br />
[[Category:Chromosome 1]]</div>Mikahttps://ngdc.cncb.ac.cn/ricewiki/index.php?title=Os01g0182600&diff=183706Os01g01826002014-06-10T14:57:39Z<p>Mika: Undo revision 183705 by Mika (talk)</p>
<hr />
<div>The gene of ''Gigantea(OsGI)''is solo ortholog of ''GI'' in the Arabidopsis circadian system and related to photoperiodic control of flowering.<br />
<br />
==Annotated Information==<br />
===Function===<br />
*''OsGI'',a ''GI'' ortholog, regulates flowering time in response to photoperiodic conditions. Mutations in this gene delay flowering in inductive SDs (short-days),whereas flowering is marginally induced under LDs(long-days).In comparison, over expression of ''OsGI'' in transgenic rice causes late flowering in both SDs and LDs.In addition, ''OsGI'' has function to regulate ''Hd1'', a gene that also shows circadian rhythm<ref name="ref1" />. ''Os-GI'' affects gene expression of approximately 75% of genes among 27,201 genes and is required for strong amplitudes and proper phase-setting of global gene expression under natural field conditions<ref name="ref2" />. <br />
*The cDNA of ''OsGI'' firstly found in ''se5'' mutant that is completely photoperiod-insensitive, the expression patterns of rice and Arabidopsis ''GI'' gene are both photoperiodically regulated and are very similar under LD and SD conditions(FIG1)<ref name="ref3" />. ''GI'' is required for phase-dependent ''CO''transcription through the interaction with FKF1, which leads to cycling DNA binding with one finger (CDF) degradation and consequent derepression of CO. In addition, it has been known that GI protein interacts with ZEITLUPE (ZTL) F-box protein to control the diurnal rhythm of TOC1 protein in regulation of the Arabidopsis circadian clock<ref name="ref4" />. <br />
*''Os-GI'' also has function to maintain bioactive GA level through the regulation of the GA-deactivating enzyme genes in rice, ''Os-GI'' represses major ''OsGA2ox'' genes in order to maintain a proper bioactive GA level, ''osgi-1'' plants showed a semi-dwarf phenotype with elevated ''OsGA2ox'' gene expression(FIG2) <ref name="ref4" />.<br />
<br />
===Expression===<br />
*''GI'' encodes a nuclear protein that defines the proper amplitude and period length of circadian rhythms. ''GI'' expression is regulated by the circadian clock, with a peak in transcript levels at 8 to 10 h after dawn. The ''GI'' protein acts between the circadian oscillator and ''CO'' to promote flowering by increasing ''CO'' and ''FT'' mRNA abundance<ref name="ref1" />. The expression of the ''OsGI'' messenger RNA was circadian-controlled and that its temporal expression pattern was very similar to that of ''GI'' under both SD and LD conditions. Decreased ''OsGI'' expression led to early flowering under LD conditions and late flowering under SD conditions relative to the wild type, and that this early flowering phenotype under LD conditions was the opposite of that of the Arabidopsis ''gi'' mutants, confirming the opposite role of the ''GI'' gene in flowering under LD conditions in rice and Arabidopsis<ref name="ref5" />.<br />
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===Mutation===<br />
*The single ''osgi-1'' mutation extended the time to flowering only under SD conditions. Under LD conditions the single ''hd1'' mutation induced early flowering, but ''osgi-1'' plants exhibited no phenotypic change<ref name="ref4" /><br />
<br />
===Evolution===<br />
* There is some evolutionary conservation between rice circadian clock–related genes and those associated with Arabidopsis circadian clocks, where the circadian clock consists of interlocked subloops, it is also possible that ''Os-GI'' is a member of a subloop in rice circadian clocks<ref name="ref4" />.But there is a divergence in the photoperiodic pathways of these two species arises from the molecular mechanisms controlling the expression of ''FT/Hd3a'' by ''CO/Hd1(Se1)''<ref name="ref5" />(figure 3)<br />
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==Labs working on this gene==<br />
Please input related labs here.<br />
<br />
==Knowledge Extension==<br />
*The rice orthologues of the two key regulators of flowering time in Arabidopsis, ''CO'' and ''FT'', were recently shown to be important in the photoperiodic control of flowering in rice. The rice ''Hd1(Se1)''gene, which is an orthologue of the Arabidopsis ''CO'' gene, was required for the suppression of flowering under LD conditions and for the promotion of flowering under SD conditions. Furthermore, the rice ''Hd3a'' gene, an orthologue of the Arabidopsis ''FT'' gene, was shown to be an activator of flowering in rice. It was previously demonstrated that ''CO'' controls the flowering time by integrating signals from the circadian clock and light to regulate ''FT'' expression in Arabidopsis<ref name="ref5" />.Unlike Arabidopsis ''FT'', expression of ''Hd3a'' mRNA is more or less opposite to that of ''Hd1'' under inductive short-day conditions<ref name="ref6" />.<br />
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==References==<br />
<references><br />
<ref name="ref1"> Song Lim Kim,Shinyoung Lee,Hyo Jung Kim,Hong Gil Nam,Gynheung An(2007) ''OsMADS51'' is a short-day flowering promoter that functions upstream of ''Ehd1'', ''OsMADS14'', and ''Hd3a''.Plant Physiology 145(4):1484-1494.</ref><br />
<br />
==Structured Information==<br />
{{JaponicaGene|<br />
GeneName = Os01g0182600|<br />
Description = GIGANTEA protein|<br />
Version = NM_001048755.1 GI:115434923 GeneID:4325329|<br />
Length = 9083 bp|<br />
Definition = Oryza sativa Japonica Group Os01g0182600, complete gene.|<br />
Source = Oryza sativa Japonica Group<br />
<br />
ORGANISM Oryza sativa Japonica Group<br />
Eukaryota; Viridiplantae; Streptophyta; Embryophyta; Tracheophyta;<br />
Spermatophyta; Magnoliophyta; Liliopsida; Poales; Poaceae; BEP<br />
clade; Ehrhartoideae; Oryzeae; Oryza.<br />
|<br />
Chromosome = [[:category:Japonica Chromosome 1|Chromosome 1]]|<br />
AP = Chromosome 1:4328362..4337444|<br />
CDS = 4328725..4329030,4329123..4329215,4329333..4329551,4329752..4329820,4329925..4331421<br>,4332220..4332429,4332563..4332713,4332791..4333031,4333898..4333956<br>,4334379..4334653,4334813..4334955,4335034..4335088,4335181..4335258<br>,4335351..4335437|<br />
GCID = <gbrowseImage1><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage1>|<br />
GSID = <gbrowseImage2><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage2>|<br />
CDNA = <cdnaseq>atgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactgcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcatagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcggcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctga</cdnaseq>|<br />
AA = <aaseq>MSASNEKWIDGLQFSSLFWPPPQDSQQKQAQILAYVEYFGQFTA DSEQFPEDIAQLIQSCYPSKEKRLVDEVLATFVLHHPEHGHAVVHPILSRIIDGTLSY DRNGFPFMSFISLFSHTSEKEYSEQWALACGEILRVLTHYNRPIFKVDHQHSEAECSS TSDQASSCESMEKRANGSPRNEPDRKPLRPLSPWITDILLAAPLGIRSDYFRWCGGVM GKYAAGGELKPPTTAYSRGSGKHPQLMPSTPRWAVANGAGVILSVCDEEVARYETANL TAAAVPALLLPPPTTPLDEHLVAGLPPLEPYARLFHRYYAIATPSATQRLLFGLLEAP PSWAPDALDAAVQLVELLRAAEDYDSGMRLPKNWMHLHFLRAIGTAMSMRAGIAADTS AALLFRILSQPTLLFPPLRHAEGVELHHEPLGGYVSSYKRQLEVPASEATIDATAQGI ASMLCAHGPDVEWRICTIWEAAYGLLPLSSSAVDLPEIVVAAPLQPPTLSWSLYLPLL KVFEYLPRGSPSEACLMRIFVATVEAILRRTFPSETSEQSRKPRSQSKNLAVAELRTM IHSLFVESCASMDLASRLLFVVLTVCVSHQALPGGSKRPTGSDNHSSEEVTNDSRLTN GRNRCKKRQGPVATFDSYVLAAVCALSCELQLFPFISKNGNHSNLKDSIKIVIPGKTT GISNELHNSISSAILHTRRILGILEALFSLKPSSVGTSWSYSSNEIVAAAMVAAHVSE LFRRSRPCLNALSALKQCKWDAEISTRASSLYHLIDLHGKTVTSIVNKAEPLEAHLTL TPVKKDEPPIEEKNINSSDGGALEKKDASRSHRKNGFARPLLKCAEDVILNGDVASTS GKAIASLQVEASDLANFLTMDRNGGYRGSQTLLRSVLSEKQELCFSVVSLLWQKLIAS PEMQMSAESTSAHQGWRKVVDALCDIVSASPTKASAAIVLQAEKDLQPWIARDDEQGQ KMWRVNQRIVKLIAELMRNHDSPEALVILASASDLLLRATDGMLVDGEACTLPQLELL EVTARAVHLIVEWGDSGVSVADGLSNLLKCRLSTTIRCLSHPSAHVRALSMSVLRDIL NSGQINSSKLIQGEHRNGIQSPTYQCLAASIINWQADVERCIEWEAHSRRATGLTLAF LTAAAKELGCPLTC</aaseq>|<br />
DNA = <dnaseqindica>8415..8720#8230..8322#7894..8112#7625..7693#6024..7520#5016..5225#4732..4882#4414..4654#3489..3547#2792..3066#2490..2632#2357..2411#2187..2264#2008..2094#ccgagccgtttctctccctctctctctcctcctccttcccctgcgcctcctccaccgcccgtgtcgccgccgcgcgcgtctatcccctcgccgccgccgccgtctacgccgcgtgaggtccggattggttgaaaggctgatcgaggtaatgcttttttatttttcccgtgcggttttagtttttctgtgctgtgcttggtggttttttctttaaaactatttttaggctggttttgatgatggcctttctaaggttgtcgtgtatggggggcgaggtggttcgggtggatttgtggtgttatgttgaggggatttttgtttggggggagagagttttggtggtagatgcggttgggattttgaagacgtgaaagggaaattagttttgaaatttgacggcgaggttgagctgattagtattgtaataattcgtggcatcggatagatcaacgcgagatggatgcttattactgcgaatttcaggaggtttttgtggtttttttaggcgagtagcgttgattctagctggattttgctgcttcgtggaggcgtttagatgtggtgcatacggctgtggatgtaatttgtgcggggataattactgaaaattaattcgcagtaagctagctctgctttaatattttttcccctgaaacagtaacaagggatcccttcctcatttcgttaacagtggaagtgaatacagacaccaaccgatgagcagagacacctgtaattaaacaccaaaagaagaacctgttttgttcctatatattttccatttctatgctttaggtaatatactttctggttcggtgtcatagtacccgaaatagaatctttgcttgccaaaaaaagaaagaataacaagatagaacaacatcattaaattctgtcttagacaatacatttataatcgaccaacaaaaacagaccgatgagtgacgtggaataacaaatgaagtataaatgtcacgaaaaacacatctattgattttgtgtgataagccataggagcctacgaggaaccaagtgaatggttctgtacaccacatccattagcattaccataaatttgcctatttctatgatgccttccttaacatatttatcgacatgtgagcttgcaccataactaaatgttatggtaaactttaccttataaatgattgcaacatgtaccttttcatttcgataattaaaaacgagaaagcttccacctttttcctagatatattttttcgtttgattatgaatttagttgtacttgacaataaaataatgcgcttgtaagattatgcaattttcgttgcattacctccctctcttttcatttgtcagtttttttctactccctccgtttcatattttaagtcgtttgacttttttcctagtcagactttgttaagtttgacaaagtttatagtaaaatttagcaatatctaaaatattaaattggtttcatcaaatctaacattgaatatattttgataatatgttttttttgtgttgaaaatacttctatatttttctataaacttagtcaaacttggaagaaatttcagtaagaaaaaagtcaaacggcttataatatgaaacggagggagtaaatttttttttgcattaattaaaattcatgtattttcctccttttttggtttttgttgtgcgatatattacagtgtagagactctgaagctcatatggttggcattggttgatatgataacgtgtcacgttgtgagatgcagacccccgttatgggggttgattgaatgaagtggatagcactcttataactttcttgctcgaagtttcttgataccaaggcaagtagcttataagaagatttggactttggtggcataaatggatgaactaactggcggcttcaacgtggtgttgattacaggtgcttttattgctaactggagcagtttgtgtagtttctttctttctctttgagaattctctcagtttcttgggcaggctcttgagctaccaagtatgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggtaacattattcctatttgattcttgttagaaattctatatgttgaacatacttgcctcaatgggaaacttggttgcgtctcctctctgaaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcaggtactgtatctctaagcatgtttagtatcttctgtgataaaatctttctccagtgtgttaacttttgcattctttttcttgcatgcttgcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattaggtaatttgatccagtgatcaatacatgttccacaaacaattgaagaaagagtttgacatatttttctcatatttccagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgaggtaaggcccctactctaatggttagttccaatttattggttcatttctctgtgtttgataagattttgcctctgctgtgcatttcaatattatatctaaactgtgctgcttatagcttatcaaactactcttgctcataaagatgcaacttttatgcagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgagtcataataaatgataacatattattgttctgtggactgaaaatctgaaatcctcctttcttgagaaatacaacctcaattttatggtgatattgacaagaaggaagagcgcatataatttaaagttaaaacacggaaaataatagaagtatgtgagcagaacgcaggagcaaggggtggaactcataacaagttcaaacttcatgcatgttcataattccccaatgcaagtatggcaagaaaaatagacaacttttaaaagtatgctaatggaactgcatgcattgtgtaaacttgcttttgtcaagcatgttggaacatactatatatttgctgctgatgatgaaaaatttgcttgagtactcttctattgctaatggacgcttaccattccctgttttgtcttcaaatgcatcaggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactggttggtaaaatcagcaatggaattttctatcgtgaattttttataattattgtaacattaattactcccatgtctttttattctgttgagataccatgtgacaatgtctacaattgcatattttgcattctttggtagatcagttcccttcttaagatccccaaagggaaactggagtatcaataacttactccctcaaataaaaaaaaagtgatactttaggcataaacacattcttcaaaacaaaactttgaatagcagtttcttttgtaatatattaataaagcatggaaaaattataatgtgaagaatgtacattcagggacaaatatactcatcatttttgcatggttgatcttaatgtccatagagagattgatcagaccaaaagtttatttttgacaggagtgagtatggattataataggagtgaatggcaatatgctgtgaaagtgagcattttatatttgtttactactagcatttggtgacaaagcaaatgatgtatattgtaaatcaaagtaaacaagtagaacatgtcctgtggcgacaagacataacctacaaattaaagttagcctgattgtttattttactgggagcacattttttctagattctagagtacatattattaaaaaaggtgtgatatgtttgcaaattactttttaatggaagtttgtaaattactttgaaacttgcattcagtttttttgcttcttaaaaaacatagtgcatatgggctttatgagctcatactatgcatgtttttctgcattggagtgggacctttatctttttcttgcaactcggatgctaatgcttattataaacacaagttactaacatgtgcttgggtcattcagcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcataggtaagctcttatgatcctgtccactgttgtaccatgtgtactttccaaatgattgctaacatgctatttttcgtcagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtac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/>
Link = [http://www.ncbi.nlm.nih.gov/nuccore/NM_001048755.1 RefSeq:Os01g0182600]|<br />
}}<br />
[[Category:Genes]]<br />
[[Category:Japonica mRNA]]<br />
[[Category:Oryza Sativa Japonica Group]]<br />
[[Category:Japonica Genes]]<br />
[[Category:Japonica Chromosome 1]]<br />
[[Category:Chromosome 1]]</div>Mikahttps://ngdc.cncb.ac.cn/ricewiki/index.php?title=Os01g0182600&diff=183705Os01g01826002014-06-10T14:56:52Z<p>Mika: Undo revision 183704 by Mika (talk)</p>
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<div>The gene of ''Gigantea(OsGI)''is solo ortholog of ''GI'' in the Arabidopsis circadian system and related to photoperiodic control of flowering.<br />
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==Annotated Information==<br />
===Function===<br />
*''OsGI'',a ''GI'' ortholog, regulates flowering time in response to photoperiodic conditions. Mutations in this gene delay flowering in inductive SDs (short-days),whereas flowering is marginally induced under LDs(long-days).In comparison, over expression of ''OsGI'' in transgenic rice causes late flowering in both SDs and LDs.In addition, ''OsGI'' has function to regulate ''Hd1'', a gene that also shows circadian rhythm<ref name="ref1" />. ''Os-GI'' affects gene expression of approximately 75% of genes among 27,201 genes and is required for strong amplitudes and proper phase-setting of global gene expression under natural field conditions<ref name="ref2" />. <br />
*The cDNA of ''OsGI'' firstly found in ''se5'' mutant that is completely photoperiod-insensitive, the expression patterns of rice and Arabidopsis ''GI'' gene are both photoperiodically regulated and are very similar under LD and SD conditions(FIG1)<ref name="ref3" />. ''GI'' is required for phase-dependent ''CO''transcription through the interaction with FKF1, which leads to cycling DNA binding with one finger (CDF) degradation and consequent derepression of CO. In addition, it has been known that GI protein interacts with ZEITLUPE (ZTL) F-box protein to control the diurnal rhythm of TOC1 protein in regulation of the Arabidopsis circadian clock<ref name="ref4" />. <br />
*''Os-GI'' also has function to maintain bioactive GA level through the regulation of the GA-deactivating enzyme genes in rice, ''Os-GI'' represses major ''OsGA2ox'' genes in order to maintain a proper bioactive GA level, ''osgi-1'' plants showed a semi-dwarf phenotype with elevated ''OsGA2ox'' gene expression(FIG2) <ref name="ref4" />.<br />
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===Expression===<br />
*''GI'' encodes a nuclear protein that defines the proper amplitude and period length of circadian rhythms. ''GI'' expression is regulated by the circadian clock, with a peak in transcript levels at 8 to 10 h after dawn. The ''GI'' protein acts between the circadian oscillator and ''CO'' to promote flowering by increasing ''CO'' and ''FT'' mRNA abundance<ref name="ref1" />. The expression of the ''OsGI'' messenger RNA was circadian-controlled and that its temporal expression pattern was very similar to that of ''GI'' under both SD and LD conditions. Decreased ''OsGI'' expression led to early flowering under LD conditions and late flowering under SD conditions relative to the wild type, and that this early flowering phenotype under LD conditions was the opposite of that of the Arabidopsis ''gi'' mutants, confirming the opposite role of the ''GI'' gene in flowering under LD conditions in rice and Arabidopsis<ref name="ref5" />.<br />
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===Mutation===<br />
*The single ''osgi-1'' mutation extended the time to flowering only under SD conditions. Under LD conditions the single ''hd1'' mutation induced early flowering, but ''osgi-1'' plants exhibited no phenotypic change<ref name="ref4" /><br />
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===Evolution===<br />
* There is some evolutionary conservation between rice circadian clock–related genes and those associated with Arabidopsis circadian clocks, where the circadian clock consists of interlocked subloops, it is also possible that ''Os-GI'' is a member of a subloop in rice circadian clocks<ref name="ref4" />.But there is a divergence in the photoperiodic pathways of these two species arises from the molecular mechanisms controlling the expression of ''FT/Hd3a'' by ''CO/Hd1(Se1)''<ref name="ref5" />(figure 3)<br />
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==Labs working on this gene==<br />
Please input related labs here.<br />
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==Knowledge Extension==<br />
*The rice orthologues of the two key regulators of flowering time in Arabidopsis, ''CO'' and ''FT'', were recently shown to be important in the photoperiodic control of flowering in rice. The rice ''Hd1(Se1)''gene, which is an orthologue of the Arabidopsis ''CO'' gene, was required for the suppression of flowering under LD conditions and for the promotion of flowering under SD conditions. Furthermore, the rice ''Hd3a'' gene, an orthologue of the Arabidopsis ''FT'' gene, was shown to be an activator of flowering in rice. It was previously demonstrated that ''CO'' controls the flowering time by integrating signals from the circadian clock and light to regulate ''FT'' expression in Arabidopsis<ref name="ref5" />.Unlike Arabidopsis ''FT'', expression of ''Hd3a'' mRNA is more or less opposite to that of ''Hd1'' under inductive short-day conditions<ref name="ref6" />.<br />
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==References==<br />
<references><br />
<ref name="ref1"> Song Lim Kim,Shinyoung Lee,Hyo Jung Kim,Hong Gil Nam,Gynheung An(2007) ''OsMADS51'' is a short-day flowering promoter that functions upstream of ''Ehd1'', ''OsMADS14'', and ''Hd3a''.Plant Physiology 145(4):1484-1494.</ref><br />
<ref name="ref2">Spielmeyer W, Ellis MH, Chandler PM (2002) Semidwarf (sd-1), "green revolution" rice, contains a defective gibberellin 20-oxidase gene. Proc Natl Acad Sci U S A 99: 9043-9048.</ref><br />
<ref name="ref3">Monna L, Kitazawa N, Yoshino R, Suzuki J, Masuda H, et al. (2002) Positional cloning of rice semidwarfing gene, sd-1: rice "green revolution gene" encodes a mutant enzyme involved in gibberellin synthesis. DNA Res 9: 11-17.</ref><br />
<ref name="ref4">Hedden P. (2003) The genes of the Green Revolution. Trends Genet 19: 5-9.</ref><br />
<ref name="ref5">Peng J, Richards DE, Hartley NM, Murphy GP, Devos KM, et al. (1999) 'Green revolution' genes encode mutant gibberellin response modulators. Nature 400: 256-261.</ref><br />
<ref name="ref6">Asano K, Hirano K, Ueguchi-Tanaka M, Angeles-Shim RB, Komura T, et al. (2009) Isolation and characterization of dominant dwarf mutants, ''Slr1-d'', in rice. Mol Genet Genomics 281: 223-231.</ref><br />
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==Structured Information==<br />
{{JaponicaGene|<br />
GeneName = Os01g0182600|<br />
Description = GIGANTEA protein|<br />
Version = NM_001048755.1 GI:115434923 GeneID:4325329|<br />
Length = 9083 bp|<br />
Definition = Oryza sativa Japonica Group Os01g0182600, complete gene.|<br />
Source = Oryza sativa Japonica Group<br />
<br />
ORGANISM Oryza sativa Japonica Group<br />
Eukaryota; Viridiplantae; Streptophyta; Embryophyta; Tracheophyta;<br />
Spermatophyta; Magnoliophyta; Liliopsida; Poales; Poaceae; BEP<br />
clade; Ehrhartoideae; Oryzeae; Oryza.<br />
|<br />
Chromosome = [[:category:Japonica Chromosome 1|Chromosome 1]]|<br />
AP = Chromosome 1:4328362..4337444|<br />
CDS = 4328725..4329030,4329123..4329215,4329333..4329551,4329752..4329820,4329925..4331421<br>,4332220..4332429,4332563..4332713,4332791..4333031,4333898..4333956<br>,4334379..4334653,4334813..4334955,4335034..4335088,4335181..4335258<br>,4335351..4335437|<br />
GCID = <gbrowseImage1><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage1>|<br />
GSID = <gbrowseImage2><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage2>|<br />
CDNA = <cdnaseq>atgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactgcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcatagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcggcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctga</cdnaseq>|<br />
AA = <aaseq>MSASNEKWIDGLQFSSLFWPPPQDSQQKQAQILAYVEYFGQFTA DSEQFPEDIAQLIQSCYPSKEKRLVDEVLATFVLHHPEHGHAVVHPILSRIIDGTLSY DRNGFPFMSFISLFSHTSEKEYSEQWALACGEILRVLTHYNRPIFKVDHQHSEAECSS TSDQASSCESMEKRANGSPRNEPDRKPLRPLSPWITDILLAAPLGIRSDYFRWCGGVM GKYAAGGELKPPTTAYSRGSGKHPQLMPSTPRWAVANGAGVILSVCDEEVARYETANL TAAAVPALLLPPPTTPLDEHLVAGLPPLEPYARLFHRYYAIATPSATQRLLFGLLEAP PSWAPDALDAAVQLVELLRAAEDYDSGMRLPKNWMHLHFLRAIGTAMSMRAGIAADTS AALLFRILSQPTLLFPPLRHAEGVELHHEPLGGYVSSYKRQLEVPASEATIDATAQGI ASMLCAHGPDVEWRICTIWEAAYGLLPLSSSAVDLPEIVVAAPLQPPTLSWSLYLPLL KVFEYLPRGSPSEACLMRIFVATVEAILRRTFPSETSEQSRKPRSQSKNLAVAELRTM IHSLFVESCASMDLASRLLFVVLTVCVSHQALPGGSKRPTGSDNHSSEEVTNDSRLTN GRNRCKKRQGPVATFDSYVLAAVCALSCELQLFPFISKNGNHSNLKDSIKIVIPGKTT GISNELHNSISSAILHTRRILGILEALFSLKPSSVGTSWSYSSNEIVAAAMVAAHVSE LFRRSRPCLNALSALKQCKWDAEISTRASSLYHLIDLHGKTVTSIVNKAEPLEAHLTL TPVKKDEPPIEEKNINSSDGGALEKKDASRSHRKNGFARPLLKCAEDVILNGDVASTS GKAIASLQVEASDLANFLTMDRNGGYRGSQTLLRSVLSEKQELCFSVVSLLWQKLIAS PEMQMSAESTSAHQGWRKVVDALCDIVSASPTKASAAIVLQAEKDLQPWIARDDEQGQ KMWRVNQRIVKLIAELMRNHDSPEALVILASASDLLLRATDGMLVDGEACTLPQLELL EVTARAVHLIVEWGDSGVSVADGLSNLLKCRLSTTIRCLSHPSAHVRALSMSVLRDIL NSGQINSSKLIQGEHRNGIQSPTYQCLAASIINWQADVERCIEWEAHSRRATGLTLAF LTAAAKELGCPLTC</aaseq>|<br />
DNA = <dnaseqindica>8415..8720#8230..8322#7894..8112#7625..7693#6024..7520#5016..5225#4732..4882#4414..4654#3489..3547#2792..3066#2490..2632#2357..2411#2187..2264#2008..2094#ccgagccgtttctctccctctctctctcctcctccttcccctgcgcctcctccaccgcccgtgtcgccgccgcgcgcgtctatcccctcgccgccgccgccgtctacgccgcgtgaggtccggattggttgaaaggctgatcgaggtaatgcttttttatttttcccgtgcggttttagtttttctgtgctgtgcttggtggttttttctttaaaactatttttaggctggttttgatgatggcctttctaaggttgtcgtgtatggggggcgaggtggttcgggtggatttgtggtgttatgttgaggggatttttgtttggggggagagagttttggtggtagatgcggttgggattttgaagacgtgaaagggaaattagttttgaaatttgacggcgaggttgagctgattagtattgtaataattcgtggcatcggatagatcaacgcgagatggatgcttattactgcgaatttcaggaggtttttgtggtttttttaggcgagtagcgttgattctagctggattttgctgcttcgtggaggcgtttagatgtggtgcatacggctgtggatgtaatttgtgcggggataattactgaaaattaattcgcagtaagctagctctgctttaatattttttcccctgaaacagtaacaagggatcccttcctcatttcgttaacagtggaagtgaatacagacaccaaccgatgagcagagacacctgtaattaaacaccaaaagaagaacctgttttgttcctatatattttccatttctatgctttaggtaatatactttctggttcggtgtcatagtacccgaaatagaatctttgcttgccaaaaaaagaaagaataacaagatagaacaacatcattaaattctgtcttagacaatacatttataatcgaccaacaaaaacagaccgatgagtgacgtggaataacaaatgaagtataaatgtcacgaaaaacacatctattgattttgtgtgataagccataggagcctacgaggaaccaagtgaatggttctgtacaccacatccattagcattaccataaatttgcctatttctatgatgccttccttaacatatttatcgacatgtgagcttgcaccataactaaatgttatggtaaactttaccttataaatgattgcaacatgtaccttttcatttcgataattaaaaacgagaaagcttccacctttttcctagatatattttttcgtttgattatgaatttagttgtacttgacaataaaataatgcgcttgtaagattatgcaattttcgttgcattacctccctctcttttcatttgtcagtttttttctactccctccgtttcatattttaagtcgtttgacttttttcctagtcagactttgttaagtttgacaaagtttatagtaaaatttagcaatatctaaaatattaaattggtttcatcaaatctaacattgaatatattttgataatatgttttttttgtgttgaaaatacttctatatttttctataaacttagtcaaacttggaagaaatttcagtaagaaaaaagtcaaacggcttataatatgaaacggagggagtaaatttttttttgcattaattaaaattcatgtattttcctccttttttggtttttgttgtgcgatatattacagtgtagagactctgaagctcatatggttggcattggttgatatgataacgtgtcacgttgtgagatgcagacccccgttatgggggttgattgaatgaagtggatagcactcttataactttcttgctcgaagtttcttgataccaaggcaagtagcttataagaagatttggactttggtggcataaatggatgaactaactggcggcttcaacgtggtgttgattacaggtgcttttattgctaactggagcagtttgtgtagtttctttctttctctttgagaattctctcagtttcttgggcaggctcttgagctaccaagtatgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggtaacattattcctatttgattcttgttagaaattctatatgttgaacatacttgcctcaatgggaaacttggttgcgtctcctctctgaaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcaggtactgtatctctaagcatgtttagtatcttctgtgataaaatctttctccagtgtgttaacttttgcattctttttcttgcatgcttgcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattaggtaatttgatccagtgatcaatacatgttccacaaacaattgaagaaagagtttgacatatttttctcatatttccagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgaggtaaggcccctactctaatggttagttccaatttattggttcatttctctgtgtttgataagattttgcctctgctgtgcatttcaatattatatctaaactgtgctgcttatagcttatcaaactactcttgctcataaagatgcaacttttatgcagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgagtcataataaatgataacatattattgttctgtggactgaaaatctgaaatcctcctttcttgagaaatacaacctcaattttatggtgatattgacaagaaggaagagcgcatataatttaaagttaaaacacggaaaataatagaagtatgtgagcagaacgcaggagcaaggggtggaactcataacaagttcaaacttcatgcatgttcataattccccaatgcaagtatggcaagaaaaatagacaacttttaaaagtatgctaatggaactgcatgcattgtgtaaacttgcttttgtcaagcatgttggaacatactatatatttgctgctgatgatgaaaaatttgcttgagtactcttctattgctaatggacgcttaccattccctgttttgtcttcaaatgcatcaggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactggttggtaaaatcagcaatggaattttctatcgtgaattttttataattattgtaacattaattactcccatgtctttttattctgttgagataccatgtgacaatgtctacaattgcatattttgcattctttggtagatcagttcccttcttaagatccccaaagggaaactggagtatcaataacttactccctcaaataaaaaaaaagtgatactttaggcataaacacattcttcaaaacaaaactttgaatagcagtttcttttgtaatatattaataaagcatggaaaaattataatgtgaagaatgtacattcagggacaaatatactcatcatttttgcatggttgatcttaatgtccatagagagattgatcagaccaaaagtttatttttgacaggagtgagtatggattataataggagtgaatggcaatatgctgtgaaagtgagcattttatatttgtttactactagcatttggtgacaaagcaaatgatgtatattgtaaatcaaagtaaacaagtagaacatgtcctgtggcgacaagacataacctacaaattaaagttagcctgattgtttattttactgggagcacattttttctagattctagagtacatattattaaaaaaggtgtgatatgtttgcaaattactttttaatggaagtttgtaaattactttgaaacttgcattcagtttttttgcttcttaaaaaacatagtgcatatgggctttatgagctcatactatgcatgtttttctgcattggagtgggacctttatctttttcttgcaactcggatgctaatgcttattataaacacaagttactaacatgtgcttgggtcattcagcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcataggtaagctcttatgatcctgtccactgttgtaccatgtgtactttccaaatgattgctaacatgctatttttcgtcagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcgggtatatcatacatctctctggctgttgtttcagtgctatcaatcaagatgacacatcatgcaatattatcataacagcacattatgtgttttgaatattttattgattgtcatagctttgttttatgttttagcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcaggtatgcagtgtttcctgcacctacttgctacctattctgtatttgatgtattcaatttcctttttatcagttttctcgaaatatatgctaataggcttgtcactaaacggctacattatccagactctgttctgaatttttttttcattaagtactgttagctgccagattacatcatcatgactaaagtagtttagctacaatattgcaaagctagatttgtctagcctctattgttttgaatagaaaaggcaggtatgcagtgtttcctgcacctacttgctacctattctgtatttgatgtattcaatttcctttttatcagttttctcgaaatatatgctaataggcttgtcactaaacggctacattatccagactctgttctgaatttttttttcattaagtactgttagctgccagattacatcatcatgactaaagtagtttagctacaatattgcaaagctagatttgtctagcctctattgttttgaatagaaaattaaatcttttggtcatttttttgtaaccttttaggagtctctgtatgcttgtcgtcaacatgctatgataagtttatatcagttcttttggagctatgatgtattcttgtatattttatgactgctaggatattatgttaaaactgttttaaaggtatcatcaaacaagttaatatagatattgctacctaccaaagttggaatttgattatgtcgtaaaatatatgcaccctacatgttttagcttcatattgacttatattttcatatgtgctccttcaactgcacaaagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggtatggtcattcctgcttataagataccagaataatgctatgaagaccgtccttggttgtcagcattcttatacacaatgataatctttgtttttcatgggtaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggtcaagatattgttattgttatatttacatacttgttatcatatttaaaagcttcatattgtatgtacctttgaaaaaaaaagttccatactgtagcttcttttgttctcatttcttggtttcaaagcatcagcatcattaccgatataaaatatacccatttcctcagtcatcttccccgactcttgtatctttcacaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagaggtaacaccttcattcatgaggtccttaggtgcaaatcccagtgaagaataatcagttctactttgtatggttcatctcgcttactgaccggttcatctcgggatcaattacgagcagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaaggcatgtcctcttctctgatctcactcacgttcatgccaaacttactcatcatcatccatgatccatcatcctgactgaatcttgatatgcagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctgacaaggccacaccatgactgtcactgcaagcagctgctgtaggatcagcgagtaggaaaggatgacttgccggccagtttcctgagatgtgatttaagactgatattagctgatgttcccaagcatgatacggggcttgctcccaagatgtgatttttactttacttttggttaatgtgctaccgctgacattagatgttcaagcatatgaaaactgcttgtgctgtaacttgtatgtctgtaacgtaaaaatctttccatatggtaaccggggaagcagaggtatatccttttctgctgttgtatctatttccagtttcctgctgaacttgcaaatgaatattcagcaatttccgttttgctc</dnaseqindica>|<br />
Link = [http://www.ncbi.nlm.nih.gov/nuccore/NM_001048755.1 RefSeq:Os01g0182600]|<br />
}}<br />
[[Category:Genes]]<br />
[[Category:Japonica mRNA]]<br />
[[Category:Oryza Sativa Japonica Group]]<br />
[[Category:Japonica Genes]]<br />
[[Category:Japonica Chromosome 1]]<br />
[[Category:Chromosome 1]]</div>Mikahttps://ngdc.cncb.ac.cn/ricewiki/index.php?title=Os01g0182600&diff=183704Os01g01826002014-06-10T14:56:42Z<p>Mika: Undo revision 183703 by Mika (talk)</p>
<hr />
<div>The gene of ''Gigantea(OsGI)''is solo ortholog of ''GI'' in the Arabidopsis circadian system and related to photoperiodic control of flowering.<br />
<br />
==Annotated Information==<br />
===Function===<br />
*''OsGI'',a ''GI'' ortholog, regulates flowering time in response to photoperiodic conditions. Mutations in this gene delay flowering in inductive SDs (short-days),whereas flowering is marginally induced under LDs(long-days).In comparison, over expression of ''OsGI'' in transgenic rice causes late flowering in both SDs and LDs.In addition, ''OsGI'' has function to regulate ''Hd1'', a gene that also shows circadian rhythm<ref name="ref1" />. ''Os-GI'' affects gene expression of approximately 75% of genes among 27,201 genes and is required for strong amplitudes and proper phase-setting of global gene expression under natural field conditions<ref name="ref2" />. <br />
*The cDNA of ''OsGI'' firstly found in ''se5'' mutant that is completely photoperiod-insensitive, the expression patterns of rice and Arabidopsis ''GI'' gene are both photoperiodically regulated and are very similar under LD and SD conditions(FIG1)<ref name="ref3" />. ''GI'' is required for phase-dependent ''CO''transcription through the interaction with FKF1, which leads to cycling DNA binding with one finger (CDF) degradation and consequent derepression of CO. In addition, it has been known that GI protein interacts with ZEITLUPE (ZTL) F-box protein to control the diurnal rhythm of TOC1 protein in regulation of the Arabidopsis circadian clock<ref name="ref4" />. <br />
*''Os-GI'' also has function to maintain bioactive GA level through the regulation of the GA-deactivating enzyme genes in rice, ''Os-GI'' represses major ''OsGA2ox'' genes in order to maintain a proper bioactive GA level, ''osgi-1'' plants showed a semi-dwarf phenotype with elevated ''OsGA2ox'' gene expression(FIG2) <ref name="ref4" />.<br />
<br />
===Expression===<br />
*''GI'' encodes a nuclear protein that defines the proper amplitude and period length of circadian rhythms. ''GI'' expression is regulated by the circadian clock, with a peak in transcript levels at 8 to 10 h after dawn. The ''GI'' protein acts between the circadian oscillator and ''CO'' to promote flowering by increasing ''CO'' and ''FT'' mRNA abundance<ref name="ref1" />. The expression of the ''OsGI'' messenger RNA was circadian-controlled and that its temporal expression pattern was very similar to that of ''GI'' under both SD and LD conditions. Decreased ''OsGI'' expression led to early flowering under LD conditions and late flowering under SD conditions relative to the wild type, and that this early flowering phenotype under LD conditions was the opposite of that of the Arabidopsis ''gi'' mutants, confirming the opposite role of the ''GI'' gene in flowering under LD conditions in rice and Arabidopsis<ref name="ref5" />.<br />
<br />
===Mutation===<br />
*The single ''osgi-1'' mutation extended the time to flowering only under SD conditions. Under LD conditions the single ''hd1'' mutation induced early flowering, but ''osgi-1'' plants exhibited no phenotypic change<ref name="ref4" /><br />
<br />
===Evolution===<br />
* There is some evolutionary conservation between rice circadian clock–related genes and those associated with Arabidopsis circadian clocks, where the circadian clock consists of interlocked subloops, it is also possible that ''Os-GI'' is a member of a subloop in rice circadian clocks<ref name="ref4" />.But there is a divergence in the photoperiodic pathways of these two species arises from the molecular mechanisms controlling the expression of ''FT/Hd3a'' by ''CO/Hd1(Se1)''<ref name="ref5" />(figure 3)<br />
<br />
==Labs working on this gene==<br />
Please input related labs here.<br />
<br />
==Knowledge Extension==<br />
*The rice orthologues of the two key regulators of flowering time in Arabidopsis, ''CO'' and ''FT'', were recently shown to be important in the photoperiodic control of flowering in rice. The rice ''Hd1(Se1)''gene, which is an orthologue of the Arabidopsis ''CO'' gene, was required for the suppression of flowering under LD conditions and for the promotion of flowering under SD conditions. Furthermore, the rice ''Hd3a'' gene, an orthologue of the Arabidopsis ''FT'' gene, was shown to be an activator of flowering in rice. It was previously demonstrated that ''CO'' controls the flowering time by integrating signals from the circadian clock and light to regulate ''FT'' expression in Arabidopsis<ref name="ref5" />.Unlike Arabidopsis ''FT'', expression of ''Hd3a'' mRNA is more or less opposite to that of ''Hd1'' under inductive short-day conditions<ref name="ref6" />.<br />
<br />
==References==<br />
<references><br />
<ref name="ref1"> Song Lim Kim,Shinyoung Lee,Hyo Jung Kim,Hong Gil Nam,Gynheung An(2007) ''OsMADS51'' is a short-day flowering promoter that functions upstream of ''Ehd1'', ''OsMADS14'', and ''Hd3a''.Plant Physiology 145(4):1484-1494.</ref><br />
<br />
==Structured Information==<br />
{{JaponicaGene|<br />
GeneName = Os01g0182600|<br />
Description = GIGANTEA protein|<br />
Version = NM_001048755.1 GI:115434923 GeneID:4325329|<br />
Length = 9083 bp|<br />
Definition = Oryza sativa Japonica Group Os01g0182600, complete gene.|<br />
Source = Oryza sativa Japonica Group<br />
<br />
ORGANISM Oryza sativa Japonica Group<br />
Eukaryota; Viridiplantae; Streptophyta; Embryophyta; Tracheophyta;<br />
Spermatophyta; Magnoliophyta; Liliopsida; Poales; Poaceae; BEP<br />
clade; Ehrhartoideae; Oryzeae; Oryza.<br />
|<br />
Chromosome = [[:category:Japonica Chromosome 1|Chromosome 1]]|<br />
AP = Chromosome 1:4328362..4337444|<br />
CDS = 4328725..4329030,4329123..4329215,4329333..4329551,4329752..4329820,4329925..4331421<br>,4332220..4332429,4332563..4332713,4332791..4333031,4333898..4333956<br>,4334379..4334653,4334813..4334955,4335034..4335088,4335181..4335258<br>,4335351..4335437|<br />
GCID = <gbrowseImage1><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage1>|<br />
GSID = <gbrowseImage2><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage2>|<br />
CDNA = <cdnaseq>atgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactgcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcatagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcggcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctga</cdnaseq>|<br />
AA = <aaseq>MSASNEKWIDGLQFSSLFWPPPQDSQQKQAQILAYVEYFGQFTA DSEQFPEDIAQLIQSCYPSKEKRLVDEVLATFVLHHPEHGHAVVHPILSRIIDGTLSY DRNGFPFMSFISLFSHTSEKEYSEQWALACGEILRVLTHYNRPIFKVDHQHSEAECSS TSDQASSCESMEKRANGSPRNEPDRKPLRPLSPWITDILLAAPLGIRSDYFRWCGGVM GKYAAGGELKPPTTAYSRGSGKHPQLMPSTPRWAVANGAGVILSVCDEEVARYETANL TAAAVPALLLPPPTTPLDEHLVAGLPPLEPYARLFHRYYAIATPSATQRLLFGLLEAP PSWAPDALDAAVQLVELLRAAEDYDSGMRLPKNWMHLHFLRAIGTAMSMRAGIAADTS AALLFRILSQPTLLFPPLRHAEGVELHHEPLGGYVSSYKRQLEVPASEATIDATAQGI ASMLCAHGPDVEWRICTIWEAAYGLLPLSSSAVDLPEIVVAAPLQPPTLSWSLYLPLL KVFEYLPRGSPSEACLMRIFVATVEAILRRTFPSETSEQSRKPRSQSKNLAVAELRTM IHSLFVESCASMDLASRLLFVVLTVCVSHQALPGGSKRPTGSDNHSSEEVTNDSRLTN GRNRCKKRQGPVATFDSYVLAAVCALSCELQLFPFISKNGNHSNLKDSIKIVIPGKTT GISNELHNSISSAILHTRRILGILEALFSLKPSSVGTSWSYSSNEIVAAAMVAAHVSE LFRRSRPCLNALSALKQCKWDAEISTRASSLYHLIDLHGKTVTSIVNKAEPLEAHLTL TPVKKDEPPIEEKNINSSDGGALEKKDASRSHRKNGFARPLLKCAEDVILNGDVASTS GKAIASLQVEASDLANFLTMDRNGGYRGSQTLLRSVLSEKQELCFSVVSLLWQKLIAS PEMQMSAESTSAHQGWRKVVDALCDIVSASPTKASAAIVLQAEKDLQPWIARDDEQGQ KMWRVNQRIVKLIAELMRNHDSPEALVILASASDLLLRATDGMLVDGEACTLPQLELL EVTARAVHLIVEWGDSGVSVADGLSNLLKCRLSTTIRCLSHPSAHVRALSMSVLRDIL NSGQINSSKLIQGEHRNGIQSPTYQCLAASIINWQADVERCIEWEAHSRRATGLTLAF LTAAAKELGCPLTC</aaseq>|<br />
DNA = <dnaseqindica>8415..8720#8230..8322#7894..8112#7625..7693#6024..7520#5016..5225#4732..4882#4414..4654#3489..3547#2792..3066#2490..2632#2357..2411#2187..2264#2008..2094#ccgagccgtttctctccctctctctctcctcctccttcccctgcgcctcctccaccgcccgtgtcgccgccgcgcgcgtctatcccctcgccgccgccgccgtctacgccgcgtgaggtccggattggttgaaaggctgatcgaggtaatgcttttttatttttcccgtgcggttttagtttttctgtgctgtgcttggtggttttttctttaaaactatttttaggctggttttgatgatggcctttctaaggttgtcgtgtatggggggcgaggtggttcgggtggatttgtggtgttatgttgaggggatttttgtttggggggagagagttttggtggtagatgcggttgggattttgaagacgtgaaagggaaattagttttgaaatttgacggcgaggttgagctgattagtattgtaataattcgtggcatcggatagatcaacgcgagatggatgcttattactgcgaatttcaggaggtttttgtggtttttttaggcgagtagcgttgattctagctggattttgctgcttcgtggaggcgtttagatgtggtgcatacggctgtggatgtaatttgtgcggggataattactgaaaattaattcgcagtaagctagctctgctttaatattttttcccctgaaacagtaacaagggatcccttcctcatttcgttaacagtggaagtgaatacagacaccaaccgatgagcagagacacctgtaattaaacaccaaaagaagaacctgttttgttcctatatattttccatttctatgctttaggtaatatactttctggttcggtgtcatagtacccgaaatagaatctttgcttgccaaaaaaagaaagaataacaagatagaacaacatcattaaattctgtcttagacaatacatttataatcgaccaacaaaaacagaccgatgagtgacgtggaataacaaatgaagtataaatgtcacgaaaaacacatctattgattttgtgtgataagccataggagcctacgaggaaccaagtgaatggttctgtacaccacatccattagcattaccataaatttgcctatttctatgatgccttccttaacatatttatcgacatgtgagcttgcaccataactaaatgttatggtaaactttaccttataaatgattgcaacatgtaccttttcatttcgataattaaaaacgagaaagcttccacctttttcctagatatattttttcgtttgattatgaatttagttgtacttgacaataaaataatgcgcttgtaagattatgcaattttcgttgcattacctccctctcttttcatttgtcagtttttttctactccctccgtttcatattttaagtcgtttgacttttttcctagtcagactttgttaagtttgacaaagtttatagtaaaatttagcaatatctaaaatattaaattggtttcatcaaatctaacattgaatatattttgataatatgttttttttgtgttgaaaatacttctatatttttctataaacttagtcaaacttggaagaaatttcagtaagaaaaaagtcaaacggcttataatatgaaacggagggagtaaatttttttttgcattaattaaaattcatgtattttcctccttttttggtttttgttgtgcgatatattacagtgtagagactctgaagctcatatggttggcattggttgatatgataacgtgtcacgttgtgagatgcagacccccgttatgggggttgattgaatgaagtggatagcactcttataactttcttgctcgaagtttcttgataccaaggcaagtagcttataagaagatttggactttggtggcataaatggatgaactaactggcggcttcaacgtggtgttgattacaggtgcttttattgctaactggagcagtttgtgtagtttctttctttctctttgagaattctctcagtttcttgggcaggctcttgagctaccaagtatgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggtaacattattcctatttgattcttgttagaaattctatatgttgaacatacttgcctcaatgggaaacttggttgcgtctcctctctgaaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcaggtactgtatctctaagcatgtttagtatcttctgtgataaaatctttctccagtgtgttaacttttgcattctttttcttgcatgcttgcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattaggtaatttgatccagtgatcaatacatgttccacaaacaattgaagaaagagtttgacatatttttctcatatttccagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgaggtaaggcccctactctaatggttagttccaatttattggttcatttctctgtgtttgataagattttgcctctgctgtgcatttcaatattatatctaaactgtgctgcttatagcttatcaaactactcttgctcataaagatgcaacttttatgcagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgagtcataataaatgataacatattattgttctgtggactgaaaatctgaaatcctcctttcttgagaaatacaacctcaattttatggtgatattgacaagaaggaagagcgcatataatttaaagttaaaacacggaaaataatagaagtatgtgagcagaacgcaggagcaaggggtggaactcataacaagttcaaacttcatgcatgttcataattccccaatgcaagtatggcaagaaaaatagacaacttttaaaagtatgctaatggaactgcatgcattgtgtaaacttgcttttgtcaagcatgttggaacatactatatatttgctgctgatgatgaaaaatttgcttgagtactcttctattgctaatggacgcttaccattccctgttttgtcttcaaatgcatcaggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactggttggtaaaatcagcaatggaattttctatcgtgaattttttataattattgtaacattaattactcccatgtctttttattctgttgagataccatgtgacaatgtctacaattgcatattttgcattctttggtagatcagttcccttcttaagatccccaaagggaaactggagtatcaataacttactccctcaaataaaaaaaaagtgatactttaggcataaacacattcttcaaaacaaaactttgaatagcagtttcttttgtaatatattaataaagcatggaaaaattataatgtgaagaatgtacattcagggacaaatatactcatcatttttgcatggttgatcttaatgtccatagagagattgatcagaccaaaagtttatttttgacaggagtgagtatggattataataggagtgaatggcaatatgctgtgaaagtgagcattttatatttgtttactactagcatttggtgacaaagcaaatgatgtatattgtaaatcaaagtaaacaagtagaacatgtcctgtggcgacaagacataacctacaaattaaagttagcctgattgtttattttactgggagcacattttttctagattctagagtacatattattaaaaaaggtgtgatatgtttgcaaattactttttaatggaagtttgtaaattactttgaaacttgcattcagtttttttgcttcttaaaaaacatagtgcatatgggctttatgagctcatactatgcatgtttttctgcattggagtgggacctttatctttttcttgcaactcggatgctaatgcttattataaacacaagttactaacatgtgcttgggtcattcagcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcataggtaagctcttatgatcctgtccactgttgtaccatgtgtactttccaaatgattgctaacatgctatttttcgtcagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcgggtatatcatacatctctctggctgttgtttcagtgctatcaatcaagatgacacatcatgcaatattatcataacagcacattatgtgttttgaatattttattgattgtcatagctttgttttatgttttagcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcaggtatgcagtgtttcctgcacctacttgctacctattctgtatttgatgtattcaatttcctttttatcagttttctcgaaatatatgctaataggcttgtcactaaacggctacattatccagactctgttctgaatttttttttcattaagtactgttagctgccagattacatcatcatgactaaagtagtttagctacaatattgcaaagctagatttgtctagcctctattgttttgaatagaaaaggcaggtatgcagtgtttcctgcacctacttgctacctattctgtatttgatgtattcaatttcctttttatcagttttctcgaaatatatgctaataggcttgtcactaaacggctacattatccagactctgttctgaatttttttttcattaagtactgttagctgccagattacatcatcatgactaaagtagtttagctacaatattgcaaagctagatttgtctagcctctattgttttgaatagaaaattaaatcttttggtcatttttttgtaaccttttaggagtctctgtatgcttgtcgtcaacatgctatgataagtttatatcagttcttttggagctatgatgtattcttgtatattttatgactgctaggatattatgttaaaactgttttaaaggtatcatcaaacaagttaatatagatattgctacctaccaaagttggaatttgattatgtcgtaaaatatatgcaccctacatgttttagcttcatattgacttatattttcatatgtgctccttcaactgcacaaagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggtatggtcattcctgcttataagataccagaataatgctatgaagaccgtccttggttgtcagcattcttatacacaatgataatctttgtttttcatgggtaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggtcaagatattgttattgttatatttacatacttgttatcatatttaaaagcttcatattgtatgtacctttgaaaaaaaaagttccatactgtagcttcttttgttctcatttcttggtttcaaagcatcagcatcattaccgatataaaatatacccatttcctcagtcatcttccccgactcttgtatctttcacaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagaggtaacaccttcattcatgaggtccttaggtgcaaatcccagtgaagaataatcagttctactttgtatggttcatctcgcttactgaccggttcatctcgggatcaattacgagcagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaaggcatgtcctcttctctgatctcactcacgttcatgccaaacttactcatcatcatccatgatccatcatcctgactgaatcttgatatgcagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctgacaaggccacaccatgactgtcactgcaagcagctgctgtaggatcagcgagtaggaaaggatgacttgccggccagtttcctgagatgtgatttaagactgatattagctgatgttcccaagcatgatacggggcttgctcccaagatgtgatttttactttacttttggttaatgtgctaccgctgacattagatgttcaagcatatgaaaactgcttgtgctgtaacttgtatgtctgtaacgtaaaaatctttccatatggtaaccggggaagcagaggtatatccttttctgctgttgtatctatttccagtttcctgctgaacttgcaaatgaatattcagcaatttccgttttgctc</dnaseqindica>|<br />
Link = [http://www.ncbi.nlm.nih.gov/nuccore/NM_001048755.1 RefSeq:Os01g0182600]|<br />
}}<br />
[[Category:Genes]]<br />
[[Category:Japonica mRNA]]<br />
[[Category:Oryza Sativa Japonica Group]]<br />
[[Category:Japonica Genes]]<br />
[[Category:Japonica Chromosome 1]]<br />
[[Category:Chromosome 1]]</div>Mikahttps://ngdc.cncb.ac.cn/ricewiki/index.php?title=Os01g0182600&diff=183703Os01g01826002014-06-10T14:56:14Z<p>Mika: Undo revision 183702 by Mika (talk)</p>
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<div>The gene of ''Gigantea(OsGI)''is solo ortholog of ''GI'' in the Arabidopsis circadian system and related to photoperiodic control of flowering.<br />
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==Annotated Information==<br />
===Function===<br />
*''OsGI'',a ''GI'' ortholog, regulates flowering time in response to photoperiodic conditions. Mutations in this gene delay flowering in inductive SDs (short-days),whereas flowering is marginally induced under LDs(long-days).In comparison, over expression of ''OsGI'' in transgenic rice causes late flowering in both SDs and LDs.In addition, ''OsGI'' has function to regulate ''Hd1'', a gene that also shows circadian rhythm<ref name="ref1" />. ''Os-GI'' affects gene expression of approximately 75% of genes among 27,201 genes and is required for strong amplitudes and proper phase-setting of global gene expression under natural field conditions<ref name="ref2" />. <br />
*The cDNA of ''OsGI'' firstly found in ''se5'' mutant that is completely photoperiod-insensitive, the expression patterns of rice and Arabidopsis ''GI'' gene are both photoperiodically regulated and are very similar under LD and SD conditions(FIG1)<ref name="ref3" />. ''GI'' is required for phase-dependent ''CO''transcription through the interaction with FKF1, which leads to cycling DNA binding with one finger (CDF) degradation and consequent derepression of CO. In addition, it has been known that GI protein interacts with ZEITLUPE (ZTL) F-box protein to control the diurnal rhythm of TOC1 protein in regulation of the Arabidopsis circadian clock<ref name="ref4" />. <br />
*''Os-GI'' also has function to maintain bioactive GA level through the regulation of the GA-deactivating enzyme genes in rice, ''Os-GI'' represses major ''OsGA2ox'' genes in order to maintain a proper bioactive GA level, ''osgi-1'' plants showed a semi-dwarf phenotype with elevated ''OsGA2ox'' gene expression(FIG2) <ref name="ref4" />.<br />
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===Expression===<br />
*''GI'' encodes a nuclear protein that defines the proper amplitude and period length of circadian rhythms. ''GI'' expression is regulated by the circadian clock, with a peak in transcript levels at 8 to 10 h after dawn. The ''GI'' protein acts between the circadian oscillator and ''CO'' to promote flowering by increasing ''CO'' and ''FT'' mRNA abundance<ref name="ref1" />. The expression of the ''OsGI'' messenger RNA was circadian-controlled and that its temporal expression pattern was very similar to that of ''GI'' under both SD and LD conditions. Decreased ''OsGI'' expression led to early flowering under LD conditions and late flowering under SD conditions relative to the wild type, and that this early flowering phenotype under LD conditions was the opposite of that of the Arabidopsis ''gi'' mutants, confirming the opposite role of the ''GI'' gene in flowering under LD conditions in rice and Arabidopsis<ref name="ref5" />.<br />
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===Mutation===<br />
*The single ''osgi-1'' mutation extended the time to flowering only under SD conditions. Under LD conditions the single ''hd1'' mutation induced early flowering, but ''osgi-1'' plants exhibited no phenotypic change<ref name="ref4" /><br />
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===Evolution===<br />
* There is some evolutionary conservation between rice circadian clock–related genes and those associated with Arabidopsis circadian clocks, where the circadian clock consists of interlocked subloops, it is also possible that ''Os-GI'' is a member of a subloop in rice circadian clocks<ref name="ref4" />.But there is a divergence in the photoperiodic pathways of these two species arises from the molecular mechanisms controlling the expression of ''FT/Hd3a'' by ''CO/Hd1(Se1)''<ref name="ref5" />(figure 3)<br />
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==Labs working on this gene==<br />
Please input related labs here.<br />
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==Knowledge Extension==<br />
*The rice orthologues of the two key regulators of flowering time in Arabidopsis, ''CO'' and ''FT'', were recently shown to be important in the photoperiodic control of flowering in rice. The rice ''Hd1(Se1)''gene, which is an orthologue of the Arabidopsis ''CO'' gene, was required for the suppression of flowering under LD conditions and for the promotion of flowering under SD conditions. Furthermore, the rice ''Hd3a'' gene, an orthologue of the Arabidopsis ''FT'' gene, was shown to be an activator of flowering in rice. It was previously demonstrated that ''CO'' controls the flowering time by integrating signals from the circadian clock and light to regulate ''FT'' expression in Arabidopsis<ref name="ref5" />.Unlike Arabidopsis ''FT'', expression of ''Hd3a'' mRNA is more or less opposite to that of ''Hd1'' under inductive short-day conditions<ref name="ref6" />.<br />
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==References==<br />
<references><br />
<ref name="ref1"> Song Lim Kim,Shinyoung Lee,Hyo Jung Kim,Hong Gil Nam,Gynheung An(2007) ''OsMADS51'' is a short-day flowering promoter that functions upstream of ''Ehd1'', ''OsMADS14'', and ''Hd3a''.Plant Physiology 145(4):1484-1494.</ref><br />
<ref name="ref2">Spielmeyer W, Ellis MH, Chandler PM (2002) Semidwarf (sd-1), "green revolution" rice, contains a defective gibberellin 20-oxidase gene. Proc Natl Acad Sci U S A 99: 9043-9048.</ref><br />
<ref name="ref3">Monna L, Kitazawa N, Yoshino R, Suzuki J, Masuda H, et al. (2002) Positional cloning of rice semidwarfing gene, sd-1: rice "green revolution gene" encodes a mutant enzyme involved in gibberellin synthesis. DNA Res 9: 11-17.</ref><br />
<ref name="ref4">Hedden P. (2003) The genes of the Green Revolution. Trends Genet 19: 5-9.</ref><br />
<ref name="ref5">Peng J, Richards DE, Hartley NM, Murphy GP, Devos KM, et al. (1999) 'Green revolution' genes encode mutant gibberellin response modulators. Nature 400: 256-261.</ref><br />
<ref name="ref6">Asano K, Hirano K, Ueguchi-Tanaka M, Angeles-Shim RB, Komura T, et al. (2009) Isolation and characterization of dominant dwarf mutants, ''Slr1-d'', in rice. Mol Genet Genomics 281: 223-231.</ref><br />
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==Structured Information==<br />
{{JaponicaGene|<br />
GeneName = Os01g0182600|<br />
Description = GIGANTEA protein|<br />
Version = NM_001048755.1 GI:115434923 GeneID:4325329|<br />
Length = 9083 bp|<br />
Definition = Oryza sativa Japonica Group Os01g0182600, complete gene.|<br />
Source = Oryza sativa Japonica Group<br />
<br />
ORGANISM Oryza sativa Japonica Group<br />
Eukaryota; Viridiplantae; Streptophyta; Embryophyta; Tracheophyta;<br />
Spermatophyta; Magnoliophyta; Liliopsida; Poales; Poaceae; BEP<br />
clade; Ehrhartoideae; Oryzeae; Oryza.<br />
|<br />
Chromosome = [[:category:Japonica Chromosome 1|Chromosome 1]]|<br />
AP = Chromosome 1:4328362..4337444|<br />
CDS = 4328725..4329030,4329123..4329215,4329333..4329551,4329752..4329820,4329925..4331421<br>,4332220..4332429,4332563..4332713,4332791..4333031,4333898..4333956<br>,4334379..4334653,4334813..4334955,4335034..4335088,4335181..4335258<br>,4335351..4335437|<br />
GCID = <gbrowseImage1><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage1>|<br />
GSID = <gbrowseImage2><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage2>|<br />
CDNA = <cdnaseq>atgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactgcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcatagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcggcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctga</cdnaseq>|<br />
AA = <aaseq>MSASNEKWIDGLQFSSLFWPPPQDSQQKQAQILAYVEYFGQFTA DSEQFPEDIAQLIQSCYPSKEKRLVDEVLATFVLHHPEHGHAVVHPILSRIIDGTLSY DRNGFPFMSFISLFSHTSEKEYSEQWALACGEILRVLTHYNRPIFKVDHQHSEAECSS TSDQASSCESMEKRANGSPRNEPDRKPLRPLSPWITDILLAAPLGIRSDYFRWCGGVM GKYAAGGELKPPTTAYSRGSGKHPQLMPSTPRWAVANGAGVILSVCDEEVARYETANL TAAAVPALLLPPPTTPLDEHLVAGLPPLEPYARLFHRYYAIATPSATQRLLFGLLEAP PSWAPDALDAAVQLVELLRAAEDYDSGMRLPKNWMHLHFLRAIGTAMSMRAGIAADTS AALLFRILSQPTLLFPPLRHAEGVELHHEPLGGYVSSYKRQLEVPASEATIDATAQGI ASMLCAHGPDVEWRICTIWEAAYGLLPLSSSAVDLPEIVVAAPLQPPTLSWSLYLPLL KVFEYLPRGSPSEACLMRIFVATVEAILRRTFPSETSEQSRKPRSQSKNLAVAELRTM IHSLFVESCASMDLASRLLFVVLTVCVSHQALPGGSKRPTGSDNHSSEEVTNDSRLTN GRNRCKKRQGPVATFDSYVLAAVCALSCELQLFPFISKNGNHSNLKDSIKIVIPGKTT GISNELHNSISSAILHTRRILGILEALFSLKPSSVGTSWSYSSNEIVAAAMVAAHVSE LFRRSRPCLNALSALKQCKWDAEISTRASSLYHLIDLHGKTVTSIVNKAEPLEAHLTL TPVKKDEPPIEEKNINSSDGGALEKKDASRSHRKNGFARPLLKCAEDVILNGDVASTS GKAIASLQVEASDLANFLTMDRNGGYRGSQTLLRSVLSEKQELCFSVVSLLWQKLIAS PEMQMSAESTSAHQGWRKVVDALCDIVSASPTKASAAIVLQAEKDLQPWIARDDEQGQ KMWRVNQRIVKLIAELMRNHDSPEALVILASASDLLLRATDGMLVDGEACTLPQLELL EVTARAVHLIVEWGDSGVSVADGLSNLLKCRLSTTIRCLSHPSAHVRALSMSVLRDIL NSGQINSSKLIQGEHRNGIQSPTYQCLAASIINWQADVERCIEWEAHSRRATGLTLAF LTAAAKELGCPLTC</aaseq>|<br />
DNA = <dnaseqindica>8415..8720#8230..8322#7894..8112#7625..7693#6024..7520#5016..5225#4732..4882#4414..4654#3489..3547#2792..3066#2490..2632#2357..2411#2187..2264#2008..2094#ccgagccgtttctctccctctctctctcctcctccttcccctgcgcctcctccaccgcccgtgtcgccgccgcgcgcgtctatcccctcgccgccgccgccgtctacgccgcgtgaggtccggattggttgaaaggctgatcgaggtaatgcttttttatttttcccgtgcggttttagtttttctgtgctgtgcttggtggttttttctttaaaactatttttaggctggttttgatgatggcctttctaaggttgtcgtgtatggggggcgaggtggttcgggtggatttgtggtgttatgttgaggggatttttgtttggggggagagagttttggtggtagatgcggttgggattttgaagacgtgaaagggaaattagttttgaaatttgacggcgaggttgagctgattagtattgtaataattcgtggcatcggatagatcaacgcgagatggatgcttattactgcgaatttcaggaggtttttgtggtttttttaggcgagtagcgttgattctagctggattttgctgcttcgtggaggcgtttagatgtggtgcatacggctgtggatgtaatttgtgcggggataattactgaaaattaattcgcagtaagctagctctgctttaatattttttcccctgaaacagtaacaagggatcccttcctcatttcgttaacagtggaagtgaatacagacaccaaccgatgagcagagacacctgtaattaaacaccaaaagaagaacctgttttgttcctatatattttccatttctatgctttaggtaatatactttctggttcggtgtcatagtacccgaaatagaatctttgcttgccaaaaaaagaaagaataacaagatagaacaacatcattaaattctgtcttagacaatacatttataatcgaccaacaaaaacagaccgatgagtgacgtggaataacaaatgaagtataaatgtcacgaaaaacacatctattgattttgtgtgataagccataggagcctacgaggaaccaagtgaatggttctgtacaccacatccattagcattaccataaatttgcctatttctatgatgccttccttaacatatttatcgacatgtgagcttgcaccataactaaatgttatggtaaactttaccttataaatgattgcaacatgtaccttttcatttcgataattaaaaacgagaaagcttccacctttttcctagatatattttttcgtttgattatgaatttagttgtacttgacaataaaataatgcgcttgtaagattatgcaattttcgttgcattacctccctctcttttcatttgtcagtttttttctactccctccgtttcatattttaagtcgtttgacttttttcctagtcagactttgttaagtttgacaaagtttatagtaaaatttagcaatatctaaaatattaaattggtttcatcaaatctaacattgaatatattttgataatatgttttttttgtgttgaaaatacttctatatttttctataaacttagtcaaacttggaagaaatttcagtaagaaaaaagtcaaacggcttataatatgaaacggagggagtaaatttttttttgcattaattaaaattcatgtattttcctccttttttggtttttgttgtgcgatatattacagtgtagagactctgaagctcatatggttggcattggttgatatgataacgtgtcacgttgtgagatgcagacccccgttatgggggttgattgaatgaagtggatagcactcttataactttcttgctcgaagtttcttgataccaaggcaagtagcttataagaagatttggactttggtggcataaatggatgaactaactggcggcttcaacgtggtgttgattacaggtgcttttattgctaactggagcagtttgtgtagtttctttctttctctttgagaattctctcagtttcttgggcaggctcttgagctaccaagtatgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggtaacattattcctatttgattcttgttagaaattctatatgttgaacatacttgcctcaatgggaaacttggttgcgtctcctctctgaaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcaggtactgtatctctaagcatgtttagtatcttctgtgataaaatctttctccagtgtgttaacttttgcattctttttcttgcatgcttgcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattaggtaatttgatccagtgatcaatacatgttccacaaacaattgaagaaagagtttgacatatttttctcatatttccagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgaggtaaggcccctactctaatggttagttccaatttattggttcatttctctgtgtttgataagattttgcctctgctgtgcatttcaatattatatctaaactgtgctgcttatagcttatcaaactactcttgctcataaagatgcaacttttatgcagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgagtcataataaatgataacatattattgttctgtggactgaaaatctgaaatcctcctttcttgagaaatacaacctcaattttatggtgatattgacaagaaggaagagcgcatataatttaaagttaaaacacggaaaataatagaagtatgtgagcagaacgcaggagcaaggggtggaactcataacaagttcaaacttcatgcatgttcataattccccaatgcaagtatggcaagaaaaatagacaacttttaaaagtatgctaatggaactgcatgcattgtgtaaacttgcttttgtcaagcatgttggaacatactatatatttgctgctgatgatgaaaaatttgcttgagtactcttctattgctaatggacgcttaccattccctgttttgtcttcaaatgcatcaggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactggttggtaaaatcagcaatggaattttctatcgtgaattttttataattattgtaacattaattactcccatgtctttttattctgttgagataccatgtgacaatgtctacaattgcatattttgcattctttggtagatcagttcccttcttaagatccccaaagggaaactggagtatcaataacttactccctcaaataaaaaaaaagtgatactttaggcataaacacattcttcaaaacaaaactttgaatagcagtttcttttgtaatatattaataaagcatggaaaaattataatgtgaagaatgtacattcagggacaaatatactcatcatttttgcatggttgatcttaatgtccatagagagattgatcagaccaaaagtttatttttgacaggagtgagtatggattataataggagtgaatggcaatatgctgtgaaagtgagcattttatatttgtttactactagcatttggtgacaaagcaaatgatgtatattgtaaatcaaagtaaacaagtagaacatgtcctgtggcgacaagacataacctacaaattaaagttagcctgattgtttattttactgggagcacattttttctagattctagagtacatattattaaaaaaggtgtgatatgtttgcaaattactttttaatggaagtttgtaaattactttgaaacttgcattcagtttttttgcttcttaaaaaacatagtgcatatgggctttatgagctcatactatgcatgtttttctgcattggagtgggacctttatctttttcttgcaactcggatgctaatgcttattataaacacaagttactaacatgtgcttgggtcattcagcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcataggtaagctcttatgatcctgtccactgttgtaccatgtgtactttccaaatgattgctaacatgctatttttcgtcagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtac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/>
Link = [http://www.ncbi.nlm.nih.gov/nuccore/NM_001048755.1 RefSeq:Os01g0182600]|<br />
}}<br />
[[Category:Genes]]<br />
[[Category:Japonica mRNA]]<br />
[[Category:Oryza Sativa Japonica Group]]<br />
[[Category:Japonica Genes]]<br />
[[Category:Japonica Chromosome 1]]<br />
[[Category:Chromosome 1]]</div>Mikahttps://ngdc.cncb.ac.cn/ricewiki/index.php?title=Os01g0182600&diff=183702Os01g01826002014-06-10T14:53:23Z<p>Mika: /* References */</p>
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<div>The gene of ''Gigantea(OsGI)''is solo ortholog of ''GI'' in the Arabidopsis circadian system and related to photoperiodic control of flowering.<br />
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==Annotated Information==<br />
===Function===<br />
*''OsGI'',a ''GI'' ortholog, regulates flowering time in response to photoperiodic conditions. Mutations in this gene delay flowering in inductive SDs (short-days),whereas flowering is marginally induced under LDs(long-days).In comparison, over expression of ''OsGI'' in transgenic rice causes late flowering in both SDs and LDs.In addition, ''OsGI'' has function to regulate ''Hd1'', a gene that also shows circadian rhythm<ref name="ref1" />. ''Os-GI'' affects gene expression of approximately 75% of genes among 27,201 genes and is required for strong amplitudes and proper phase-setting of global gene expression under natural field conditions<ref name="ref2" />. <br />
*The cDNA of ''OsGI'' firstly found in ''se5'' mutant that is completely photoperiod-insensitive, the expression patterns of rice and Arabidopsis ''GI'' gene are both photoperiodically regulated and are very similar under LD and SD conditions(FIG1)<ref name="ref3" />. ''GI'' is required for phase-dependent ''CO''transcription through the interaction with FKF1, which leads to cycling DNA binding with one finger (CDF) degradation and consequent derepression of CO. In addition, it has been known that GI protein interacts with ZEITLUPE (ZTL) F-box protein to control the diurnal rhythm of TOC1 protein in regulation of the Arabidopsis circadian clock<ref name="ref4" />. <br />
*''Os-GI'' also has function to maintain bioactive GA level through the regulation of the GA-deactivating enzyme genes in rice, ''Os-GI'' represses major ''OsGA2ox'' genes in order to maintain a proper bioactive GA level, ''osgi-1'' plants showed a semi-dwarf phenotype with elevated ''OsGA2ox'' gene expression(FIG2) <ref name="ref4" />.<br />
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===Expression===<br />
*''GI'' encodes a nuclear protein that defines the proper amplitude and period length of circadian rhythms. ''GI'' expression is regulated by the circadian clock, with a peak in transcript levels at 8 to 10 h after dawn. The ''GI'' protein acts between the circadian oscillator and ''CO'' to promote flowering by increasing ''CO'' and ''FT'' mRNA abundance<ref name="ref1" />. The expression of the ''OsGI'' messenger RNA was circadian-controlled and that its temporal expression pattern was very similar to that of ''GI'' under both SD and LD conditions. Decreased ''OsGI'' expression led to early flowering under LD conditions and late flowering under SD conditions relative to the wild type, and that this early flowering phenotype under LD conditions was the opposite of that of the Arabidopsis ''gi'' mutants, confirming the opposite role of the ''GI'' gene in flowering under LD conditions in rice and Arabidopsis<ref name="ref5" />.<br />
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===Mutation===<br />
*The single ''osgi-1'' mutation extended the time to flowering only under SD conditions. Under LD conditions the single ''hd1'' mutation induced early flowering, but ''osgi-1'' plants exhibited no phenotypic change<ref name="ref4" /><br />
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===Evolution===<br />
* There is some evolutionary conservation between rice circadian clock–related genes and those associated with Arabidopsis circadian clocks, where the circadian clock consists of interlocked subloops, it is also possible that ''Os-GI'' is a member of a subloop in rice circadian clocks<ref name="ref4" />.But there is a divergence in the photoperiodic pathways of these two species arises from the molecular mechanisms controlling the expression of ''FT/Hd3a'' by ''CO/Hd1(Se1)''<ref name="ref5" />(figure 3)<br />
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==Labs working on this gene==<br />
Please input related labs here.<br />
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==Knowledge Extension==<br />
*The rice orthologues of the two key regulators of flowering time in Arabidopsis, ''CO'' and ''FT'', were recently shown to be important in the photoperiodic control of flowering in rice. The rice ''Hd1(Se1)''gene, which is an orthologue of the Arabidopsis ''CO'' gene, was required for the suppression of flowering under LD conditions and for the promotion of flowering under SD conditions. Furthermore, the rice ''Hd3a'' gene, an orthologue of the Arabidopsis ''FT'' gene, was shown to be an activator of flowering in rice. It was previously demonstrated that ''CO'' controls the flowering time by integrating signals from the circadian clock and light to regulate ''FT'' expression in Arabidopsis<ref name="ref5" />.Unlike Arabidopsis ''FT'', expression of ''Hd3a'' mRNA is more or less opposite to that of ''Hd1'' under inductive short-day conditions<ref name="ref6" />.<br />
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==References==<br />
<references><br />
<ref name="ref1"> Song Lim Kim,Shinyoung Lee,Hyo Jung Kim,Hong Gil Nam,Gynheung An(2007) ''OsMADS51'' is a short-day flowering promoter that functions upstream of ''Ehd1'', ''OsMADS14'', and ''Hd3a''.Plant Physiology 145(4):1484-1494.</ref><br />
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==Structured Information==<br />
{{JaponicaGene|<br />
GeneName = Os01g0182600|<br />
Description = GIGANTEA protein|<br />
Version = NM_001048755.1 GI:115434923 GeneID:4325329|<br />
Length = 9083 bp|<br />
Definition = Oryza sativa Japonica Group Os01g0182600, complete gene.|<br />
Source = Oryza sativa Japonica Group<br />
<br />
ORGANISM Oryza sativa Japonica Group<br />
Eukaryota; Viridiplantae; Streptophyta; Embryophyta; Tracheophyta;<br />
Spermatophyta; Magnoliophyta; Liliopsida; Poales; Poaceae; BEP<br />
clade; Ehrhartoideae; Oryzeae; Oryza.<br />
|<br />
Chromosome = [[:category:Japonica Chromosome 1|Chromosome 1]]|<br />
AP = Chromosome 1:4328362..4337444|<br />
CDS = 4328725..4329030,4329123..4329215,4329333..4329551,4329752..4329820,4329925..4331421<br>,4332220..4332429,4332563..4332713,4332791..4333031,4333898..4333956<br>,4334379..4334653,4334813..4334955,4335034..4335088,4335181..4335258<br>,4335351..4335437|<br />
GCID = <gbrowseImage1><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage1>|<br />
GSID = <gbrowseImage2><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage2>|<br />
CDNA = <cdnaseq>atgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactgcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcatagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcggcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctga</cdnaseq>|<br />
AA = <aaseq>MSASNEKWIDGLQFSSLFWPPPQDSQQKQAQILAYVEYFGQFTA DSEQFPEDIAQLIQSCYPSKEKRLVDEVLATFVLHHPEHGHAVVHPILSRIIDGTLSY DRNGFPFMSFISLFSHTSEKEYSEQWALACGEILRVLTHYNRPIFKVDHQHSEAECSS TSDQASSCESMEKRANGSPRNEPDRKPLRPLSPWITDILLAAPLGIRSDYFRWCGGVM GKYAAGGELKPPTTAYSRGSGKHPQLMPSTPRWAVANGAGVILSVCDEEVARYETANL TAAAVPALLLPPPTTPLDEHLVAGLPPLEPYARLFHRYYAIATPSATQRLLFGLLEAP PSWAPDALDAAVQLVELLRAAEDYDSGMRLPKNWMHLHFLRAIGTAMSMRAGIAADTS AALLFRILSQPTLLFPPLRHAEGVELHHEPLGGYVSSYKRQLEVPASEATIDATAQGI ASMLCAHGPDVEWRICTIWEAAYGLLPLSSSAVDLPEIVVAAPLQPPTLSWSLYLPLL KVFEYLPRGSPSEACLMRIFVATVEAILRRTFPSETSEQSRKPRSQSKNLAVAELRTM IHSLFVESCASMDLASRLLFVVLTVCVSHQALPGGSKRPTGSDNHSSEEVTNDSRLTN GRNRCKKRQGPVATFDSYVLAAVCALSCELQLFPFISKNGNHSNLKDSIKIVIPGKTT GISNELHNSISSAILHTRRILGILEALFSLKPSSVGTSWSYSSNEIVAAAMVAAHVSE LFRRSRPCLNALSALKQCKWDAEISTRASSLYHLIDLHGKTVTSIVNKAEPLEAHLTL TPVKKDEPPIEEKNINSSDGGALEKKDASRSHRKNGFARPLLKCAEDVILNGDVASTS GKAIASLQVEASDLANFLTMDRNGGYRGSQTLLRSVLSEKQELCFSVVSLLWQKLIAS PEMQMSAESTSAHQGWRKVVDALCDIVSASPTKASAAIVLQAEKDLQPWIARDDEQGQ KMWRVNQRIVKLIAELMRNHDSPEALVILASASDLLLRATDGMLVDGEACTLPQLELL EVTARAVHLIVEWGDSGVSVADGLSNLLKCRLSTTIRCLSHPSAHVRALSMSVLRDIL NSGQINSSKLIQGEHRNGIQSPTYQCLAASIINWQADVERCIEWEAHSRRATGLTLAF LTAAAKELGCPLTC</aaseq>|<br />
DNA = <dnaseqindica>8415..8720#8230..8322#7894..8112#7625..7693#6024..7520#5016..5225#4732..4882#4414..4654#3489..3547#2792..3066#2490..2632#2357..2411#2187..2264#2008..2094#ccgagccgtttctctccctctctctctcctcctccttcccctgcgcctcctccaccgcccgtgtcgccgccgcgcgcgtctatcccctcgccgccgccgccgtctacgccgcgtgaggtccggattggttgaaaggctgatcgaggtaatgcttttttatttttcccgtgcggttttagtttttctgtgctgtgcttggtggttttttctttaaaactatttttaggctggttttgatgatggcctttctaaggttgtcgtgtatggggggcgaggtggttcgggtggatttgtggtgttatgttgaggggatttttgtttggggggagagagttttggtggtagatgcggttgggattttgaagacgtgaaagggaaattagttttgaaatttgacggcgaggttgagctgattagtattgtaataattcgtggcatcggatagatcaacgcgagatggatgcttattactgcgaatttcaggaggtttttgtggtttttttaggcgagtagcgttgattctagctggattttgctgcttcgtggaggcgtttagatgtggtgcatacggctgtggatgtaatttgtgcggggataattactgaaaattaattcgcagtaagctagctctgctttaatattttttcccctgaaacagtaacaagggatcccttcctcatttcgttaacagtggaagtgaatacagacaccaaccgatgagcagagacacctgtaattaaacaccaaaagaagaacctgttttgttcctatatattttccatttctatgctttaggtaatatactttctggttcggtgtcatagtacccgaaatagaatctttgcttgccaaaaaaagaaagaataacaagatagaacaacatcattaaattctgtcttagacaatacatttataatcgaccaacaaaaacagaccgatgagtgacgtggaataacaaatgaagtataaatgtcacgaaaaacacatctattgattttgtgtgataagccataggagcctacgaggaaccaagtgaatggttctgtacaccacatccattagcattaccataaatttgcctatttctatgatgccttccttaacatatttatcgacatgtgagcttgcaccataactaaatgttatggtaaactttaccttataaatgattgcaacatgtaccttttcatttcgataattaaaaacgagaaagcttccacctttttcctagatatattttttcgtttgattatgaatttagttgtacttgacaataaaataatgcgcttgtaagattatgcaattttcgttgcattacctccctctcttttcatttgtcagtttttttctactccctccgtttcatattttaagtcgtttgacttttttcctagtcagactttgttaagtttgacaaagtttatagtaaaatttagcaatatctaaaatattaaattggtttcatcaaatctaacattgaatatattttgataatatgttttttttgtgttgaaaatacttctatatttttctataaacttagtcaaacttggaagaaatttcagtaagaaaaaagtcaaacggcttataatatgaaacggagggagtaaatttttttttgcattaattaaaattcatgtattttcctccttttttggtttttgttgtgcgatatattacagtgtagagactctgaagctcatatggttggcattggttgatatgataacgtgtcacgttgtgagatgcagacccccgttatgggggttgattgaatgaagtggatagcactcttataactttcttgctcgaagtttcttgataccaaggcaagtagcttataagaagatttggactttggtggcataaatggatgaactaactggcggcttcaacgtggtgttgattacaggtgcttttattgctaactggagcagtttgtgtagtttctttctttctctttgagaattctctcagtttcttgggcaggctcttgagctaccaagtatgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggtaacattattcctatttgattcttgttagaaattctatatgttgaacatacttgcctcaatgggaaacttggttgcgtctcctctctgaaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcaggtactgtatctctaagcatgtttagtatcttctgtgataaaatctttctccagtgtgttaacttttgcattctttttcttgcatgcttgcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattaggtaatttgatccagtgatcaatacatgttccacaaacaattgaagaaagagtttgacatatttttctcatatttccagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgaggtaaggcccctactctaatggttagttccaatttattggttcatttctctgtgtttgataagattttgcctctgctgtgcatttcaatattatatctaaactgtgctgcttatagcttatcaaactactcttgctcataaagatgcaacttttatgcagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgagtcataataaatgataacatattattgttctgtggactgaaaatctgaaatcctcctttcttgagaaatacaacctcaattttatggtgatattgacaagaaggaagagcgcatataatttaaagttaaaacacggaaaataatagaagtatgtgagcagaacgcaggagcaaggggtggaactcataacaagttcaaacttcatgcatgttcataattccccaatgcaagtatggcaagaaaaatagacaacttttaaaagtatgctaatggaactgcatgcattgtgtaaacttgcttttgtcaagcatgttggaacatactatatatttgctgctgatgatgaaaaatttgcttgagtactcttctattgctaatggacgcttaccattccctgttttgtcttcaaatgcatcaggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactggttggtaaaatcagcaatggaattttctatcgtgaattttttataattattgtaacattaattactcccatgtctttttattctgttgagataccatgtgacaatgtctacaattgcatattttgcattctttggtagatcagttcccttcttaagatccccaaagggaaactggagtatcaataacttactccctcaaataaaaaaaaagtgatactttaggcataaacacattcttcaaaacaaaactttgaatagcagtttcttttgtaatatattaataaagcatggaaaaattataatgtgaagaatgtacattcagggacaaatatactcatcatttttgcatggttgatcttaatgtccatagagagattgatcagaccaaaagtttatttttgacaggagtgagtatggattataataggagtgaatggcaatatgctgtgaaagtgagcattttatatttgtttactactagcatttggtgacaaagcaaatgatgtatattgtaaatcaaagtaaacaagtagaacatgtcctgtggcgacaagacataacctacaaattaaagttagcctgattgtttattttactgggagcacattttttctagattctagagtacatattattaaaaaaggtgtgatatgtttgcaaattactttttaatggaagtttgtaaattactttgaaacttgcattcagtttttttgcttcttaaaaaacatagtgcatatgggctttatgagctcatactatgcatgtttttctgcattggagtgggacctttatctttttcttgcaactcggatgctaatgcttattataaacacaagttactaacatgtgcttgggtcattcagcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcataggtaagctcttatgatcctgtccactgttgtaccatgtgtactttccaaatgattgctaacatgctatttttcgtcagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcgggtatatcatacatctctctggctgttgtttcagtgctatcaatcaagatgacacatcatgcaatattatcataacagcacattatgtgttttgaatattttattgattgtcatagctttgttttatgttttagcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcaggtatgcagtgtttcctgcacctacttgctacctattctgtatttgatgtattcaatttcctttttatcagttttctcgaaatatatgctaataggcttgtcactaaacggctacattatccagactctgttctgaatttttttttcattaagtactgttagctgccagattacatcatcatgactaaagtagtttagctacaatattgcaaagctagatttgtctagcctctattgttttgaatagaaaaggcaggtatgcagtgtttcctgcacctacttgctacctattctgtatttgatgtattcaatttcctttttatcagttttctcgaaatatatgctaataggcttgtcactaaacggctacattatccagactctgttctgaatttttttttcattaagtactgttagctgccagattacatcatcatgactaaagtagtttagctacaatattgcaaagctagatttgtctagcctctattgttttgaatagaaaattaaatcttttggtcatttttttgtaaccttttaggagtctctgtatgcttgtcgtcaacatgctatgataagtttatatcagttcttttggagctatgatgtattcttgtatattttatgactgctaggatattatgttaaaactgttttaaaggtatcatcaaacaagttaatatagatattgctacctaccaaagttggaatttgattatgtcgtaaaatatatgcaccctacatgttttagcttcatattgacttatattttcatatgtgctccttcaactgcacaaagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggtatggtcattcctgcttataagataccagaataatgctatgaagaccgtccttggttgtcagcattcttatacacaatgataatctttgtttttcatgggtaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggtcaagatattgttattgttatatttacatacttgttatcatatttaaaagcttcatattgtatgtacctttgaaaaaaaaagttccatactgtagcttcttttgttctcatttcttggtttcaaagcatcagcatcattaccgatataaaatatacccatttcctcagtcatcttccccgactcttgtatctttcacaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagaggtaacaccttcattcatgaggtccttaggtgcaaatcccagtgaagaataatcagttctactttgtatggttcatctcgcttactgaccggttcatctcgggatcaattacgagcagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaaggcatgtcctcttctctgatctcactcacgttcatgccaaacttactcatcatcatccatgatccatcatcctgactgaatcttgatatgcagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctgacaaggccacaccatgactgtcactgcaagcagctgctgtaggatcagcgagtaggaaaggatgacttgccggccagtttcctgagatgtgatttaagactgatattagctgatgttcccaagcatgatacggggcttgctcccaagatgtgatttttactttacttttggttaatgtgctaccgctgacattagatgttcaagcatatgaaaactgcttgtgctgtaacttgtatgtctgtaacgtaaaaatctttccatatggtaaccggggaagcagaggtatatccttttctgctgttgtatctatttccagtttcctgctgaacttgcaaatgaatattcagcaatttccgttttgctc</dnaseqindica>|<br />
Link = [http://www.ncbi.nlm.nih.gov/nuccore/NM_001048755.1 RefSeq:Os01g0182600]|<br />
}}<br />
[[Category:Genes]]<br />
[[Category:Japonica mRNA]]<br />
[[Category:Oryza Sativa Japonica Group]]<br />
[[Category:Japonica Genes]]<br />
[[Category:Japonica Chromosome 1]]<br />
[[Category:Chromosome 1]]</div>Mikahttps://ngdc.cncb.ac.cn/ricewiki/index.php?title=Os01g0182600&diff=183701Os01g01826002014-06-10T14:50:54Z<p>Mika: /* References */</p>
<hr />
<div>The gene of ''Gigantea(OsGI)''is solo ortholog of ''GI'' in the Arabidopsis circadian system and related to photoperiodic control of flowering.<br />
<br />
==Annotated Information==<br />
===Function===<br />
*''OsGI'',a ''GI'' ortholog, regulates flowering time in response to photoperiodic conditions. Mutations in this gene delay flowering in inductive SDs (short-days),whereas flowering is marginally induced under LDs(long-days).In comparison, over expression of ''OsGI'' in transgenic rice causes late flowering in both SDs and LDs.In addition, ''OsGI'' has function to regulate ''Hd1'', a gene that also shows circadian rhythm<ref name="ref1" />. ''Os-GI'' affects gene expression of approximately 75% of genes among 27,201 genes and is required for strong amplitudes and proper phase-setting of global gene expression under natural field conditions<ref name="ref2" />. <br />
*The cDNA of ''OsGI'' firstly found in ''se5'' mutant that is completely photoperiod-insensitive, the expression patterns of rice and Arabidopsis ''GI'' gene are both photoperiodically regulated and are very similar under LD and SD conditions(FIG1)<ref name="ref3" />. ''GI'' is required for phase-dependent ''CO''transcription through the interaction with FKF1, which leads to cycling DNA binding with one finger (CDF) degradation and consequent derepression of CO. In addition, it has been known that GI protein interacts with ZEITLUPE (ZTL) F-box protein to control the diurnal rhythm of TOC1 protein in regulation of the Arabidopsis circadian clock<ref name="ref4" />. <br />
*''Os-GI'' also has function to maintain bioactive GA level through the regulation of the GA-deactivating enzyme genes in rice, ''Os-GI'' represses major ''OsGA2ox'' genes in order to maintain a proper bioactive GA level, ''osgi-1'' plants showed a semi-dwarf phenotype with elevated ''OsGA2ox'' gene expression(FIG2) <ref name="ref4" />.<br />
<br />
===Expression===<br />
*''GI'' encodes a nuclear protein that defines the proper amplitude and period length of circadian rhythms. ''GI'' expression is regulated by the circadian clock, with a peak in transcript levels at 8 to 10 h after dawn. The ''GI'' protein acts between the circadian oscillator and ''CO'' to promote flowering by increasing ''CO'' and ''FT'' mRNA abundance<ref name="ref1" />. The expression of the ''OsGI'' messenger RNA was circadian-controlled and that its temporal expression pattern was very similar to that of ''GI'' under both SD and LD conditions. Decreased ''OsGI'' expression led to early flowering under LD conditions and late flowering under SD conditions relative to the wild type, and that this early flowering phenotype under LD conditions was the opposite of that of the Arabidopsis ''gi'' mutants, confirming the opposite role of the ''GI'' gene in flowering under LD conditions in rice and Arabidopsis<ref name="ref5" />.<br />
<br />
===Mutation===<br />
*The single ''osgi-1'' mutation extended the time to flowering only under SD conditions. Under LD conditions the single ''hd1'' mutation induced early flowering, but ''osgi-1'' plants exhibited no phenotypic change<ref name="ref4" /><br />
<br />
===Evolution===<br />
* There is some evolutionary conservation between rice circadian clock–related genes and those associated with Arabidopsis circadian clocks, where the circadian clock consists of interlocked subloops, it is also possible that ''Os-GI'' is a member of a subloop in rice circadian clocks<ref name="ref4" />.But there is a divergence in the photoperiodic pathways of these two species arises from the molecular mechanisms controlling the expression of ''FT/Hd3a'' by ''CO/Hd1(Se1)''<ref name="ref5" />(figure 3)<br />
<br />
==Labs working on this gene==<br />
Please input related labs here.<br />
<br />
==Knowledge Extension==<br />
*The rice orthologues of the two key regulators of flowering time in Arabidopsis, ''CO'' and ''FT'', were recently shown to be important in the photoperiodic control of flowering in rice. The rice ''Hd1(Se1)''gene, which is an orthologue of the Arabidopsis ''CO'' gene, was required for the suppression of flowering under LD conditions and for the promotion of flowering under SD conditions. Furthermore, the rice ''Hd3a'' gene, an orthologue of the Arabidopsis ''FT'' gene, was shown to be an activator of flowering in rice. It was previously demonstrated that ''CO'' controls the flowering time by integrating signals from the circadian clock and light to regulate ''FT'' expression in Arabidopsis<ref name="ref5" />.Unlike Arabidopsis ''FT'', expression of ''Hd3a'' mRNA is more or less opposite to that of ''Hd1'' under inductive short-day conditions<ref name="ref6" />.<br />
<br />
==References==<br />
<references><br />
<ref name="ref1"> Song Lim Kim,Shinyoung Lee,Hyo Jung Kim,Hong Gil Nam,Gynheung An(2007) ''OsMADS51'' is a short-day flowering promoter that functions upstream of ''Ehd1'', ''OsMADS14'', and ''Hd3a''.Plant Physiology 145(4):1484-1494.</ref><br />
<ref name="ref2">Spielmeyer W, Ellis MH, Chandler PM (2002) Semidwarf (sd-1), "green revolution" rice, contains a defective gibberellin 20-oxidase gene. Proc Natl Acad Sci U S A 99: 9043-9048.</ref><br />
<ref name="ref3">Monna L, Kitazawa N, Yoshino R, Suzuki J, Masuda H, et al. (2002) Positional cloning of rice semidwarfing gene, sd-1: rice "green revolution gene" encodes a mutant enzyme involved in gibberellin synthesis. DNA Res 9: 11-17.</ref><br />
<ref name="ref4">Hedden P. (2003) The genes of the Green Revolution. Trends Genet 19: 5-9.</ref><br />
<ref name="ref5">Peng J, Richards DE, Hartley NM, Murphy GP, Devos KM, et al. (1999) 'Green revolution' genes encode mutant gibberellin response modulators. Nature 400: 256-261.</ref><br />
<ref name="ref6">Asano K, Hirano K, Ueguchi-Tanaka M, Angeles-Shim RB, Komura T, et al. (2009) Isolation and characterization of dominant dwarf mutants, ''Slr1-d'', in rice. Mol Genet Genomics 281: 223-231.</ref><br />
<br />
==Structured Information==<br />
{{JaponicaGene|<br />
GeneName = Os01g0182600|<br />
Description = GIGANTEA protein|<br />
Version = NM_001048755.1 GI:115434923 GeneID:4325329|<br />
Length = 9083 bp|<br />
Definition = Oryza sativa Japonica Group Os01g0182600, complete gene.|<br />
Source = Oryza sativa Japonica Group<br />
<br />
ORGANISM Oryza sativa Japonica Group<br />
Eukaryota; Viridiplantae; Streptophyta; Embryophyta; Tracheophyta;<br />
Spermatophyta; Magnoliophyta; Liliopsida; Poales; Poaceae; BEP<br />
clade; Ehrhartoideae; Oryzeae; Oryza.<br />
|<br />
Chromosome = [[:category:Japonica Chromosome 1|Chromosome 1]]|<br />
AP = Chromosome 1:4328362..4337444|<br />
CDS = 4328725..4329030,4329123..4329215,4329333..4329551,4329752..4329820,4329925..4331421<br>,4332220..4332429,4332563..4332713,4332791..4333031,4333898..4333956<br>,4334379..4334653,4334813..4334955,4335034..4335088,4335181..4335258<br>,4335351..4335437|<br />
GCID = <gbrowseImage1><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage1>|<br />
GSID = <gbrowseImage2><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage2>|<br />
CDNA = <cdnaseq>atgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactgcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcatagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcggcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctga</cdnaseq>|<br />
AA = <aaseq>MSASNEKWIDGLQFSSLFWPPPQDSQQKQAQILAYVEYFGQFTA DSEQFPEDIAQLIQSCYPSKEKRLVDEVLATFVLHHPEHGHAVVHPILSRIIDGTLSY DRNGFPFMSFISLFSHTSEKEYSEQWALACGEILRVLTHYNRPIFKVDHQHSEAECSS TSDQASSCESMEKRANGSPRNEPDRKPLRPLSPWITDILLAAPLGIRSDYFRWCGGVM GKYAAGGELKPPTTAYSRGSGKHPQLMPSTPRWAVANGAGVILSVCDEEVARYETANL TAAAVPALLLPPPTTPLDEHLVAGLPPLEPYARLFHRYYAIATPSATQRLLFGLLEAP PSWAPDALDAAVQLVELLRAAEDYDSGMRLPKNWMHLHFLRAIGTAMSMRAGIAADTS AALLFRILSQPTLLFPPLRHAEGVELHHEPLGGYVSSYKRQLEVPASEATIDATAQGI ASMLCAHGPDVEWRICTIWEAAYGLLPLSSSAVDLPEIVVAAPLQPPTLSWSLYLPLL KVFEYLPRGSPSEACLMRIFVATVEAILRRTFPSETSEQSRKPRSQSKNLAVAELRTM IHSLFVESCASMDLASRLLFVVLTVCVSHQALPGGSKRPTGSDNHSSEEVTNDSRLTN GRNRCKKRQGPVATFDSYVLAAVCALSCELQLFPFISKNGNHSNLKDSIKIVIPGKTT GISNELHNSISSAILHTRRILGILEALFSLKPSSVGTSWSYSSNEIVAAAMVAAHVSE LFRRSRPCLNALSALKQCKWDAEISTRASSLYHLIDLHGKTVTSIVNKAEPLEAHLTL TPVKKDEPPIEEKNINSSDGGALEKKDASRSHRKNGFARPLLKCAEDVILNGDVASTS GKAIASLQVEASDLANFLTMDRNGGYRGSQTLLRSVLSEKQELCFSVVSLLWQKLIAS PEMQMSAESTSAHQGWRKVVDALCDIVSASPTKASAAIVLQAEKDLQPWIARDDEQGQ KMWRVNQRIVKLIAELMRNHDSPEALVILASASDLLLRATDGMLVDGEACTLPQLELL EVTARAVHLIVEWGDSGVSVADGLSNLLKCRLSTTIRCLSHPSAHVRALSMSVLRDIL NSGQINSSKLIQGEHRNGIQSPTYQCLAASIINWQADVERCIEWEAHSRRATGLTLAF LTAAAKELGCPLTC</aaseq>|<br />
DNA = <dnaseqindica>8415..8720#8230..8322#7894..8112#7625..7693#6024..7520#5016..5225#4732..4882#4414..4654#3489..3547#2792..3066#2490..2632#2357..2411#2187..2264#2008..2094#ccgagccgtttctctccctctctctctcctcctccttcccctgcgcctcctccaccgcccgtgtcgccgccgcgcgcgtctatcccctcgccgccgccgccgtctacgccgcgtgaggtccggattggttgaaaggctgatcgaggtaatgcttttttatttttcccgtgcggttttagtttttctgtgctgtgcttggtggttttttctttaaaactatttttaggctggttttgatgatggcctttctaaggttgtcgtgtatggggggcgaggtggttcgggtggatttgtggtgttatgttgaggggatttttgtttggggggagagagttttggtggtagatgcggttgggattttgaagacgtgaaagggaaattagttttgaaatttgacggcgaggttgagctgattagtattgtaataattcgtggcatcggatagatcaacgcgagatggatgcttattactgcgaatttcaggaggtttttgtggtttttttaggcgagtagcgttgattctagctggattttgctgcttcgtggaggcgtttagatgtggtgcatacggctgtggatgtaatttgtgcggggataattactgaaaattaattcgcagtaagctagctctgctttaatattttttcccctgaaacagtaacaagggatcccttcctcatttcgttaacagtggaagtgaatacagacaccaaccgatgagcagagacacctgtaattaaacaccaaaagaagaacctgttttgttcctatatattttccatttctatgctttaggtaatatactttctggttcggtgtcatagtacccgaaatagaatctttgcttgccaaaaaaagaaagaataacaagatagaacaacatcattaaattctgtcttagacaatacatttataatcgaccaacaaaaacagaccgatgagtgacgtggaataacaaatgaagtataaatgtcacgaaaaacacatctattgattttgtgtgataagccataggagcctacgaggaaccaagtgaatggttctgtacaccacatccattagcattaccataaatttgcctatttctatgatgccttccttaacatatttatcgacatgtgagcttgcaccataactaaatgttatggtaaactttaccttataaatgattgcaacatgtaccttttcatttcgataattaaaaacgagaaagcttccacctttttcctagatatattttttcgtttgattatgaatttagttgtacttgacaataaaataatgcgcttgtaagattatgcaattttcgttgcattacctccctctcttttcatttgtcagtttttttctactccctccgtttcatattttaagtcgtttgacttttttcctagtcagactttgttaagtttgacaaagtttatagtaaaatttagcaatatctaaaatattaaattggtttcatcaaatctaacattgaatatattttgataatatgttttttttgtgttgaaaatacttctatatttttctataaacttagtcaaacttggaagaaatttcagtaagaaaaaagtcaaacggcttataatatgaaacggagggagtaaatttttttttgcattaattaaaattcatgtattttcctccttttttggtttttgttgtgcgatatattacagtgtagagactctgaagctcatatggttggcattggttgatatgataacgtgtcacgttgtgagatgcagacccccgttatgggggttgattgaatgaagtggatagcactcttataactttcttgctcgaagtttcttgataccaaggcaagtagcttataagaagatttggactttggtggcataaatggatgaactaactggcggcttcaacgtggtgttgattacaggtgcttttattgctaactggagcagtttgtgtagtttctttctttctctttgagaattctctcagtttcttgggcaggctcttgagctaccaagtatgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggtaacattattcctatttgattcttgttagaaattctatatgttgaacatacttgcctcaatgggaaacttggttgcgtctcctctctgaaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcaggtactgtatctctaagcatgtttagtatcttctgtgataaaatctttctccagtgtgttaacttttgcattctttttcttgcatgcttgcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattaggtaatttgatccagtgatcaatacatgttccacaaacaattgaagaaagagtttgacatatttttctcatatttccagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgaggtaaggcccctactctaatggttagttccaatttattggttcatttctctgtgtttgataagattttgcctctgctgtgcatttcaatattatatctaaactgtgctgcttatagcttatcaaactactcttgctcataaagatgcaacttttatgcagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgagtcataataaatgataacatattattgttctgtggactgaaaatctgaaatcctcctttcttgagaaatacaacctcaattttatggtgatattgacaagaaggaagagcgcatataatttaaagttaaaacacggaaaataatagaagtatgtgagcagaacgcaggagcaaggggtggaactcataacaagttcaaacttcatgcatgttcataattccccaatgcaagtatggcaagaaaaatagacaacttttaaaagtatgctaatggaactgcatgcattgtgtaaacttgcttttgtcaagcatgttggaacatactatatatttgctgctgatgatgaaaaatttgcttgagtactcttctattgctaatggacgcttaccattccctgttttgtcttcaaatgcatcaggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactggttggtaaaatcagcaatggaattttctatcgtgaattttttataattattgtaacattaattactcccatgtctttttattctgttgagataccatgtgacaatgtctacaattgcatattttgcattctttggtagatcagttcccttcttaagatccccaaagggaaactggagtatcaataacttactccctcaaataaaaaaaaagtgatactttaggcataaacacattcttcaaaacaaaactttgaatagcagtttcttttgtaatatattaataaagcatggaaaaattataatgtgaagaatgtacattcagggacaaatatactcatcatttttgcatggttgatcttaatgtccatagagagattgatcagaccaaaagtttatttttgacaggagtgagtatggattataataggagtgaatggcaatatgctgtgaaagtgagcattttatatttgtttactactagcatttggtgacaaagcaaatgatgtatattgtaaatcaaagtaaacaagtagaacatgtcctgtggcgacaagacataacctacaaattaaagttagcctgattgtttattttactgggagcacattttttctagattctagagtacatattattaaaaaaggtgtgatatgtttgcaaattactttttaatggaagtttgtaaattactttgaaacttgcattcagtttttttgcttcttaaaaaacatagtgcatatgggctttatgagctcatactatgcatgtttttctgcattggagtgggacctttatctttttcttgcaactcggatgctaatgcttattataaacacaagttactaacatgtgcttgggtcattcagcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcataggtaagctcttatgatcctgtccactgttgtaccatgtgtactttccaaatgattgctaacatgctatttttcgtcagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtac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/>
Link = [http://www.ncbi.nlm.nih.gov/nuccore/NM_001048755.1 RefSeq:Os01g0182600]|<br />
}}<br />
[[Category:Genes]]<br />
[[Category:Japonica mRNA]]<br />
[[Category:Oryza Sativa Japonica Group]]<br />
[[Category:Japonica Genes]]<br />
[[Category:Japonica Chromosome 1]]<br />
[[Category:Chromosome 1]]</div>Mikahttps://ngdc.cncb.ac.cn/ricewiki/index.php?title=Os01g0182600&diff=183695Os01g01826002014-06-10T14:38:51Z<p>Mika: /* References */</p>
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<div>The gene of ''Gigantea(OsGI)''is solo ortholog of ''GI'' in the Arabidopsis circadian system and related to photoperiodic control of flowering.<br />
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==Annotated Information==<br />
===Function===<br />
*''OsGI'',a ''GI'' ortholog, regulates flowering time in response to photoperiodic conditions. Mutations in this gene delay flowering in inductive SDs (short-days),whereas flowering is marginally induced under LDs(long-days).In comparison, over expression of ''OsGI'' in transgenic rice causes late flowering in both SDs and LDs.In addition, ''OsGI'' has function to regulate ''Hd1'', a gene that also shows circadian rhythm<ref name="ref1" />. ''Os-GI'' affects gene expression of approximately 75% of genes among 27,201 genes and is required for strong amplitudes and proper phase-setting of global gene expression under natural field conditions<ref name="ref2" />. <br />
*The cDNA of ''OsGI'' firstly found in ''se5'' mutant that is completely photoperiod-insensitive, the expression patterns of rice and Arabidopsis ''GI'' gene are both photoperiodically regulated and are very similar under LD and SD conditions(FIG1)<ref name="ref3" />. ''GI'' is required for phase-dependent ''CO''transcription through the interaction with FKF1, which leads to cycling DNA binding with one finger (CDF) degradation and consequent derepression of CO. In addition, it has been known that GI protein interacts with ZEITLUPE (ZTL) F-box protein to control the diurnal rhythm of TOC1 protein in regulation of the Arabidopsis circadian clock<ref name="ref4" />. <br />
*''Os-GI'' also has function to maintain bioactive GA level through the regulation of the GA-deactivating enzyme genes in rice, ''Os-GI'' represses major ''OsGA2ox'' genes in order to maintain a proper bioactive GA level, ''osgi-1'' plants showed a semi-dwarf phenotype with elevated ''OsGA2ox'' gene expression(FIG2) <ref name="ref4" />.<br />
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===Expression===<br />
*''GI'' encodes a nuclear protein that defines the proper amplitude and period length of circadian rhythms. ''GI'' expression is regulated by the circadian clock, with a peak in transcript levels at 8 to 10 h after dawn. The ''GI'' protein acts between the circadian oscillator and ''CO'' to promote flowering by increasing ''CO'' and ''FT'' mRNA abundance<ref name="ref1" />. The expression of the ''OsGI'' messenger RNA was circadian-controlled and that its temporal expression pattern was very similar to that of ''GI'' under both SD and LD conditions. Decreased ''OsGI'' expression led to early flowering under LD conditions and late flowering under SD conditions relative to the wild type, and that this early flowering phenotype under LD conditions was the opposite of that of the Arabidopsis ''gi'' mutants, confirming the opposite role of the ''GI'' gene in flowering under LD conditions in rice and Arabidopsis<ref name="ref5" />.<br />
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===Mutation===<br />
*The single ''osgi-1'' mutation extended the time to flowering only under SD conditions. Under LD conditions the single ''hd1'' mutation induced early flowering, but ''osgi-1'' plants exhibited no phenotypic change<ref name="ref4" /><br />
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===Evolution===<br />
* There is some evolutionary conservation between rice circadian clock–related genes and those associated with Arabidopsis circadian clocks, where the circadian clock consists of interlocked subloops, it is also possible that ''Os-GI'' is a member of a subloop in rice circadian clocks<ref name="ref4" />.But there is a divergence in the photoperiodic pathways of these two species arises from the molecular mechanisms controlling the expression of ''FT/Hd3a'' by ''CO/Hd1(Se1)''<ref name="ref5" />(figure 3)<br />
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==Labs working on this gene==<br />
Please input related labs here.<br />
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==Knowledge Extension==<br />
*The rice orthologues of the two key regulators of flowering time in Arabidopsis, ''CO'' and ''FT'', were recently shown to be important in the photoperiodic control of flowering in rice. The rice ''Hd1(Se1)''gene, which is an orthologue of the Arabidopsis ''CO'' gene, was required for the suppression of flowering under LD conditions and for the promotion of flowering under SD conditions. Furthermore, the rice ''Hd3a'' gene, an orthologue of the Arabidopsis ''FT'' gene, was shown to be an activator of flowering in rice. It was previously demonstrated that ''CO'' controls the flowering time by integrating signals from the circadian clock and light to regulate ''FT'' expression in Arabidopsis<ref name="ref5" />.Unlike Arabidopsis ''FT'', expression of ''Hd3a'' mRNA is more or less opposite to that of ''Hd1'' under inductive short-day conditions<ref name="ref6" />.<br />
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==References==<br />
<references><br />
<ref name="ref1"> Song Lim Kim,Shinyoung Lee,Hyo Jung Kim,Hong Gil Nam,Gynheung An(2007) ''OsMADS51'' is a short-day flowering promoter that functions upstream of ''Ehd1'', ''OsMADS14'', and ''Hd3a''.Plant Physiology 145(4):1484-1494.</ref><br />
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==Structured Information==<br />
{{JaponicaGene|<br />
GeneName = Os01g0182600|<br />
Description = GIGANTEA protein|<br />
Version = NM_001048755.1 GI:115434923 GeneID:4325329|<br />
Length = 9083 bp|<br />
Definition = Oryza sativa Japonica Group Os01g0182600, complete gene.|<br />
Source = Oryza sativa Japonica Group<br />
<br />
ORGANISM Oryza sativa Japonica Group<br />
Eukaryota; Viridiplantae; Streptophyta; Embryophyta; Tracheophyta;<br />
Spermatophyta; Magnoliophyta; Liliopsida; Poales; Poaceae; BEP<br />
clade; Ehrhartoideae; Oryzeae; Oryza.<br />
|<br />
Chromosome = [[:category:Japonica Chromosome 1|Chromosome 1]]|<br />
AP = Chromosome 1:4328362..4337444|<br />
CDS = 4328725..4329030,4329123..4329215,4329333..4329551,4329752..4329820,4329925..4331421<br>,4332220..4332429,4332563..4332713,4332791..4333031,4333898..4333956<br>,4334379..4334653,4334813..4334955,4335034..4335088,4335181..4335258<br>,4335351..4335437|<br />
GCID = <gbrowseImage1><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage1>|<br />
GSID = <gbrowseImage2><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage2>|<br />
CDNA = <cdnaseq>atgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactgcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcatagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcggcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctga</cdnaseq>|<br />
AA = <aaseq>MSASNEKWIDGLQFSSLFWPPPQDSQQKQAQILAYVEYFGQFTA DSEQFPEDIAQLIQSCYPSKEKRLVDEVLATFVLHHPEHGHAVVHPILSRIIDGTLSY DRNGFPFMSFISLFSHTSEKEYSEQWALACGEILRVLTHYNRPIFKVDHQHSEAECSS TSDQASSCESMEKRANGSPRNEPDRKPLRPLSPWITDILLAAPLGIRSDYFRWCGGVM GKYAAGGELKPPTTAYSRGSGKHPQLMPSTPRWAVANGAGVILSVCDEEVARYETANL TAAAVPALLLPPPTTPLDEHLVAGLPPLEPYARLFHRYYAIATPSATQRLLFGLLEAP PSWAPDALDAAVQLVELLRAAEDYDSGMRLPKNWMHLHFLRAIGTAMSMRAGIAADTS AALLFRILSQPTLLFPPLRHAEGVELHHEPLGGYVSSYKRQLEVPASEATIDATAQGI ASMLCAHGPDVEWRICTIWEAAYGLLPLSSSAVDLPEIVVAAPLQPPTLSWSLYLPLL KVFEYLPRGSPSEACLMRIFVATVEAILRRTFPSETSEQSRKPRSQSKNLAVAELRTM IHSLFVESCASMDLASRLLFVVLTVCVSHQALPGGSKRPTGSDNHSSEEVTNDSRLTN GRNRCKKRQGPVATFDSYVLAAVCALSCELQLFPFISKNGNHSNLKDSIKIVIPGKTT GISNELHNSISSAILHTRRILGILEALFSLKPSSVGTSWSYSSNEIVAAAMVAAHVSE LFRRSRPCLNALSALKQCKWDAEISTRASSLYHLIDLHGKTVTSIVNKAEPLEAHLTL TPVKKDEPPIEEKNINSSDGGALEKKDASRSHRKNGFARPLLKCAEDVILNGDVASTS GKAIASLQVEASDLANFLTMDRNGGYRGSQTLLRSVLSEKQELCFSVVSLLWQKLIAS PEMQMSAESTSAHQGWRKVVDALCDIVSASPTKASAAIVLQAEKDLQPWIARDDEQGQ KMWRVNQRIVKLIAELMRNHDSPEALVILASASDLLLRATDGMLVDGEACTLPQLELL EVTARAVHLIVEWGDSGVSVADGLSNLLKCRLSTTIRCLSHPSAHVRALSMSVLRDIL NSGQINSSKLIQGEHRNGIQSPTYQCLAASIINWQADVERCIEWEAHSRRATGLTLAF LTAAAKELGCPLTC</aaseq>|<br />
DNA = <dnaseqindica>8415..8720#8230..8322#7894..8112#7625..7693#6024..7520#5016..5225#4732..4882#4414..4654#3489..3547#2792..3066#2490..2632#2357..2411#2187..2264#2008..2094#ccgagccgtttctctccctctctctctcctcctccttcccctgcgcctcctccaccgcccgtgtcgccgccgcgcgcgtctatcccctcgccgccgccgccgtctacgccgcgtgaggtccggattggttgaaaggctgatcgaggtaatgcttttttatttttcccgtgcggttttagtttttctgtgctgtgcttggtggttttttctttaaaactatttttaggctggttttgatgatggcctttctaaggttgtcgtgtatggggggcgaggtggttcgggtggatttgtggtgttatgttgaggggatttttgtttggggggagagagttttggtggtagatgcggttgggattttgaagacgtgaaagggaaattagttttgaaatttgacggcgaggttgagctgattagtattgtaataattcgtggcatcggatagatcaacgcgagatggatgcttattactgcgaatttcaggaggtttttgtggtttttttaggcgagtagcgttgattctagctggattttgctgcttcgtggaggcgtttagatgtggtgcatacggctgtggatgtaatttgtgcggggataattactgaaaattaattcgcagtaagctagctctgctttaatattttttcccctgaaacagtaacaagggatcccttcctcatttcgttaacagtggaagtgaatacagacaccaaccgatgagcagagacacctgtaattaaacaccaaaagaagaacctgttttgttcctatatattttccatttctatgctttaggtaatatactttctggttcggtgtcatagtacccgaaatagaatctttgcttgccaaaaaaagaaagaataacaagatagaacaacatcattaaattctgtcttagacaatacatttataatcgaccaacaaaaacagaccgatgagtgacgtggaataacaaatgaagtataaatgtcacgaaaaacacatctattgattttgtgtgataagccataggagcctacgaggaaccaagtgaatggttctgtacaccacatccattagcattaccataaatttgcctatttctatgatgccttccttaacatatttatcgacatgtgagcttgcaccataactaaatgttatggtaaactttaccttataaatgattgcaacatgtaccttttcatttcgataattaaaaacgagaaagcttccacctttttcctagatatattttttcgtttgattatgaatttagttgtacttgacaataaaataatgcgcttgtaagattatgcaattttcgttgcattacctccctctcttttcatttgtcagtttttttctactccctccgtttcatattttaagtcgtttgacttttttcctagtcagactttgttaagtttgacaaagtttatagtaaaatttagcaatatctaaaatattaaattggtttcatcaaatctaacattgaatatattttgataatatgttttttttgtgttgaaaatacttctatatttttctataaacttagtcaaacttggaagaaatttcagtaagaaaaaagtcaaacggcttataatatgaaacggagggagtaaatttttttttgcattaattaaaattcatgtattttcctccttttttggtttttgttgtgcgatatattacagtgtagagactctgaagctcatatggttggcattggttgatatgataacgtgtcacgttgtgagatgcagacccccgttatgggggttgattgaatgaagtggatagcactcttataactttcttgctcgaagtttcttgataccaaggcaagtagcttataagaagatttggactttggtggcataaatggatgaactaactggcggcttcaacgtggtgttgattacaggtgcttttattgctaactggagcagtttgtgtagtttctttctttctctttgagaattctctcagtttcttgggcaggctcttgagctaccaagtatgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggtaacattattcctatttgattcttgttagaaattctatatgttgaacatacttgcctcaatgggaaacttggttgcgtctcctctctgaaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcaggtactgtatctctaagcatgtttagtatcttctgtgataaaatctttctccagtgtgttaacttttgcattctttttcttgcatgcttgcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattaggtaatttgatccagtgatcaatacatgttccacaaacaattgaagaaagagtttgacatatttttctcatatttccagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgaggtaaggcccctactctaatggttagttccaatttattggttcatttctctgtgtttgataagattttgcctctgctgtgcatttcaatattatatctaaactgtgctgcttatagcttatcaaactactcttgctcataaagatgcaacttttatgcagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgagtcataataaatgataacatattattgttctgtggactgaaaatctgaaatcctcctttcttgagaaatacaacctcaattttatggtgatattgacaagaaggaagagcgcatataatttaaagttaaaacacggaaaataatagaagtatgtgagcagaacgcaggagcaaggggtggaactcataacaagttcaaacttcatgcatgttcataattccccaatgcaagtatggcaagaaaaatagacaacttttaaaagtatgctaatggaactgcatgcattgtgtaaacttgcttttgtcaagcatgttggaacatactatatatttgctgctgatgatgaaaaatttgcttgagtactcttctattgctaatggacgcttaccattccctgttttgtcttcaaatgcatcaggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactggttggtaaaatcagcaatggaattttctatcgtgaattttttataattattgtaacattaattactcccatgtctttttattctgttgagataccatgtgacaatgtctacaattgcatattttgcattctttggtagatcagttcccttcttaagatccccaaagggaaactggagtatcaataacttactccctcaaataaaaaaaaagtgatactttaggcataaacacattcttcaaaacaaaactttgaatagcagtttcttttgtaatatattaataaagcatggaaaaattataatgtgaagaatgtacattcagggacaaatatactcatcatttttgcatggttgatcttaatgtccatagagagattgatcagaccaaaagtttatttttgacaggagtgagtatggattataataggagtgaatggcaatatgctgtgaaagtgagcattttatatttgtttactactagcatttggtgacaaagcaaatgatgtatattgtaaatcaaagtaaacaagtagaacatgtcctgtggcgacaagacataacctacaaattaaagttagcctgattgtttattttactgggagcacattttttctagattctagagtacatattattaaaaaaggtgtgatatgtttgcaaattactttttaatggaagtttgtaaattactttgaaacttgcattcagtttttttgcttcttaaaaaacatagtgcatatgggctttatgagctcatactatgcatgtttttctgcattggagtgggacctttatctttttcttgcaactcggatgctaatgcttattataaacacaagttactaacatgtgcttgggtcattcagcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcataggtaagctcttatgatcctgtccactgttgtaccatgtgtactttccaaatgattgctaacatgctatttttcgtcagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcgggtatatcatacatctctctggctgttgtttcagtgctatcaatcaagatgacacatcatgcaatattatcataacagcacattatgtgttttgaatattttattgattgtcatagctttgttttatgttttagcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcaggtatgcagtgtttcctgcacctacttgctacctattctgtatttgatgtattcaatttcctttttatcagttttctcgaaatatatgctaataggcttgtcactaaacggctacattatccagactctgttctgaatttttttttcattaagtactgttagctgccagattacatcatcatgactaaagtagtttagctacaatattgcaaagctagatttgtctagcctctattgttttgaatagaaaaggcaggtatgcagtgtttcctgcacctacttgctacctattctgtatttgatgtattcaatttcctttttatcagttttctcgaaatatatgctaataggcttgtcactaaacggctacattatccagactctgttctgaatttttttttcattaagtactgttagctgccagattacatcatcatgactaaagtagtttagctacaatattgcaaagctagatttgtctagcctctattgttttgaatagaaaattaaatcttttggtcatttttttgtaaccttttaggagtctctgtatgcttgtcgtcaacatgctatgataagtttatatcagttcttttggagctatgatgtattcttgtatattttatgactgctaggatattatgttaaaactgttttaaaggtatcatcaaacaagttaatatagatattgctacctaccaaagttggaatttgattatgtcgtaaaatatatgcaccctacatgttttagcttcatattgacttatattttcatatgtgctccttcaactgcacaaagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggtatggtcattcctgcttataagataccagaataatgctatgaagaccgtccttggttgtcagcattcttatacacaatgataatctttgtttttcatgggtaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggtcaagatattgttattgttatatttacatacttgttatcatatttaaaagcttcatattgtatgtacctttgaaaaaaaaagttccatactgtagcttcttttgttctcatttcttggtttcaaagcatcagcatcattaccgatataaaatatacccatttcctcagtcatcttccccgactcttgtatctttcacaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagaggtaacaccttcattcatgaggtccttaggtgcaaatcccagtgaagaataatcagttctactttgtatggttcatctcgcttactgaccggttcatctcgggatcaattacgagcagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaaggcatgtcctcttctctgatctcactcacgttcatgccaaacttactcatcatcatccatgatccatcatcctgactgaatcttgatatgcagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctgacaaggccacaccatgactgtcactgcaagcagctgctgtaggatcagcgagtaggaaaggatgacttgccggccagtttcctgagatgtgatttaagactgatattagctgatgttcccaagcatgatacggggcttgctcccaagatgtgatttttactttacttttggttaatgtgctaccgctgacattagatgttcaagcatatgaaaactgcttgtgctgtaacttgtatgtctgtaacgtaaaaatctttccatatggtaaccggggaagcagaggtatatccttttctgctgttgtatctatttccagtttcctgctgaacttgcaaatgaatattcagcaatttccgttttgctc</dnaseqindica>|<br />
Link = [http://www.ncbi.nlm.nih.gov/nuccore/NM_001048755.1 RefSeq:Os01g0182600]|<br />
}}<br />
[[Category:Genes]]<br />
[[Category:Japonica mRNA]]<br />
[[Category:Oryza Sativa Japonica Group]]<br />
[[Category:Japonica Genes]]<br />
[[Category:Japonica Chromosome 1]]<br />
[[Category:Chromosome 1]]</div>Mikahttps://ngdc.cncb.ac.cn/ricewiki/index.php?title=Os01g0182600&diff=183694Os01g01826002014-06-10T14:37:57Z<p>Mika: /* References */</p>
<hr />
<div>The gene of ''Gigantea(OsGI)''is solo ortholog of ''GI'' in the Arabidopsis circadian system and related to photoperiodic control of flowering.<br />
<br />
==Annotated Information==<br />
===Function===<br />
*''OsGI'',a ''GI'' ortholog, regulates flowering time in response to photoperiodic conditions. Mutations in this gene delay flowering in inductive SDs (short-days),whereas flowering is marginally induced under LDs(long-days).In comparison, over expression of ''OsGI'' in transgenic rice causes late flowering in both SDs and LDs.In addition, ''OsGI'' has function to regulate ''Hd1'', a gene that also shows circadian rhythm<ref name="ref1" />. ''Os-GI'' affects gene expression of approximately 75% of genes among 27,201 genes and is required for strong amplitudes and proper phase-setting of global gene expression under natural field conditions<ref name="ref2" />. <br />
*The cDNA of ''OsGI'' firstly found in ''se5'' mutant that is completely photoperiod-insensitive, the expression patterns of rice and Arabidopsis ''GI'' gene are both photoperiodically regulated and are very similar under LD and SD conditions(FIG1)<ref name="ref3" />. ''GI'' is required for phase-dependent ''CO''transcription through the interaction with FKF1, which leads to cycling DNA binding with one finger (CDF) degradation and consequent derepression of CO. In addition, it has been known that GI protein interacts with ZEITLUPE (ZTL) F-box protein to control the diurnal rhythm of TOC1 protein in regulation of the Arabidopsis circadian clock<ref name="ref4" />. <br />
*''Os-GI'' also has function to maintain bioactive GA level through the regulation of the GA-deactivating enzyme genes in rice, ''Os-GI'' represses major ''OsGA2ox'' genes in order to maintain a proper bioactive GA level, ''osgi-1'' plants showed a semi-dwarf phenotype with elevated ''OsGA2ox'' gene expression(FIG2) <ref name="ref4" />.<br />
<br />
===Expression===<br />
*''GI'' encodes a nuclear protein that defines the proper amplitude and period length of circadian rhythms. ''GI'' expression is regulated by the circadian clock, with a peak in transcript levels at 8 to 10 h after dawn. The ''GI'' protein acts between the circadian oscillator and ''CO'' to promote flowering by increasing ''CO'' and ''FT'' mRNA abundance<ref name="ref1" />. The expression of the ''OsGI'' messenger RNA was circadian-controlled and that its temporal expression pattern was very similar to that of ''GI'' under both SD and LD conditions. Decreased ''OsGI'' expression led to early flowering under LD conditions and late flowering under SD conditions relative to the wild type, and that this early flowering phenotype under LD conditions was the opposite of that of the Arabidopsis ''gi'' mutants, confirming the opposite role of the ''GI'' gene in flowering under LD conditions in rice and Arabidopsis<ref name="ref5" />.<br />
<br />
===Mutation===<br />
*The single ''osgi-1'' mutation extended the time to flowering only under SD conditions. Under LD conditions the single ''hd1'' mutation induced early flowering, but ''osgi-1'' plants exhibited no phenotypic change<ref name="ref4" /><br />
<br />
===Evolution===<br />
* There is some evolutionary conservation between rice circadian clock–related genes and those associated with Arabidopsis circadian clocks, where the circadian clock consists of interlocked subloops, it is also possible that ''Os-GI'' is a member of a subloop in rice circadian clocks<ref name="ref4" />.But there is a divergence in the photoperiodic pathways of these two species arises from the molecular mechanisms controlling the expression of ''FT/Hd3a'' by ''CO/Hd1(Se1)''<ref name="ref5" />(figure 3)<br />
<br />
==Labs working on this gene==<br />
Please input related labs here.<br />
<br />
==Knowledge Extension==<br />
*The rice orthologues of the two key regulators of flowering time in Arabidopsis, ''CO'' and ''FT'', were recently shown to be important in the photoperiodic control of flowering in rice. The rice ''Hd1(Se1)''gene, which is an orthologue of the Arabidopsis ''CO'' gene, was required for the suppression of flowering under LD conditions and for the promotion of flowering under SD conditions. Furthermore, the rice ''Hd3a'' gene, an orthologue of the Arabidopsis ''FT'' gene, was shown to be an activator of flowering in rice. It was previously demonstrated that ''CO'' controls the flowering time by integrating signals from the circadian clock and light to regulate ''FT'' expression in Arabidopsis<ref name="ref5" />.Unlike Arabidopsis ''FT'', expression of ''Hd3a'' mRNA is more or less opposite to that of ''Hd1'' under inductive short-day conditions<ref name="ref6" />.<br />
<br />
==References==<br />
<ref name="ref1"> Song Lim Kim,Shinyoung Lee,Hyo Jung Kim,Hong Gil Nam,Gynheung An(2007) ''OsMADS51'' is a short-day flowering promoter that functions upstream of ''Ehd1'', ''OsMADS14'', and ''Hd3a''.Plant Physiology 145(4):1484-1494.</ref><br />
<br />
==Structured Information==<br />
{{JaponicaGene|<br />
GeneName = Os01g0182600|<br />
Description = GIGANTEA protein|<br />
Version = NM_001048755.1 GI:115434923 GeneID:4325329|<br />
Length = 9083 bp|<br />
Definition = Oryza sativa Japonica Group Os01g0182600, complete gene.|<br />
Source = Oryza sativa Japonica Group<br />
<br />
ORGANISM Oryza sativa Japonica Group<br />
Eukaryota; Viridiplantae; Streptophyta; Embryophyta; Tracheophyta;<br />
Spermatophyta; Magnoliophyta; Liliopsida; Poales; Poaceae; BEP<br />
clade; Ehrhartoideae; Oryzeae; Oryza.<br />
|<br />
Chromosome = [[:category:Japonica Chromosome 1|Chromosome 1]]|<br />
AP = Chromosome 1:4328362..4337444|<br />
CDS = 4328725..4329030,4329123..4329215,4329333..4329551,4329752..4329820,4329925..4331421<br>,4332220..4332429,4332563..4332713,4332791..4333031,4333898..4333956<br>,4334379..4334653,4334813..4334955,4335034..4335088,4335181..4335258<br>,4335351..4335437|<br />
GCID = <gbrowseImage1><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage1>|<br />
GSID = <gbrowseImage2><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage2>|<br />
CDNA = <cdnaseq>atgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactgcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcatagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcggcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctga</cdnaseq>|<br />
AA = <aaseq>MSASNEKWIDGLQFSSLFWPPPQDSQQKQAQILAYVEYFGQFTA DSEQFPEDIAQLIQSCYPSKEKRLVDEVLATFVLHHPEHGHAVVHPILSRIIDGTLSY DRNGFPFMSFISLFSHTSEKEYSEQWALACGEILRVLTHYNRPIFKVDHQHSEAECSS TSDQASSCESMEKRANGSPRNEPDRKPLRPLSPWITDILLAAPLGIRSDYFRWCGGVM GKYAAGGELKPPTTAYSRGSGKHPQLMPSTPRWAVANGAGVILSVCDEEVARYETANL TAAAVPALLLPPPTTPLDEHLVAGLPPLEPYARLFHRYYAIATPSATQRLLFGLLEAP PSWAPDALDAAVQLVELLRAAEDYDSGMRLPKNWMHLHFLRAIGTAMSMRAGIAADTS AALLFRILSQPTLLFPPLRHAEGVELHHEPLGGYVSSYKRQLEVPASEATIDATAQGI ASMLCAHGPDVEWRICTIWEAAYGLLPLSSSAVDLPEIVVAAPLQPPTLSWSLYLPLL KVFEYLPRGSPSEACLMRIFVATVEAILRRTFPSETSEQSRKPRSQSKNLAVAELRTM IHSLFVESCASMDLASRLLFVVLTVCVSHQALPGGSKRPTGSDNHSSEEVTNDSRLTN GRNRCKKRQGPVATFDSYVLAAVCALSCELQLFPFISKNGNHSNLKDSIKIVIPGKTT GISNELHNSISSAILHTRRILGILEALFSLKPSSVGTSWSYSSNEIVAAAMVAAHVSE LFRRSRPCLNALSALKQCKWDAEISTRASSLYHLIDLHGKTVTSIVNKAEPLEAHLTL TPVKKDEPPIEEKNINSSDGGALEKKDASRSHRKNGFARPLLKCAEDVILNGDVASTS GKAIASLQVEASDLANFLTMDRNGGYRGSQTLLRSVLSEKQELCFSVVSLLWQKLIAS PEMQMSAESTSAHQGWRKVVDALCDIVSASPTKASAAIVLQAEKDLQPWIARDDEQGQ KMWRVNQRIVKLIAELMRNHDSPEALVILASASDLLLRATDGMLVDGEACTLPQLELL EVTARAVHLIVEWGDSGVSVADGLSNLLKCRLSTTIRCLSHPSAHVRALSMSVLRDIL NSGQINSSKLIQGEHRNGIQSPTYQCLAASIINWQADVERCIEWEAHSRRATGLTLAF LTAAAKELGCPLTC</aaseq>|<br />
DNA = <dnaseqindica>8415..8720#8230..8322#7894..8112#7625..7693#6024..7520#5016..5225#4732..4882#4414..4654#3489..3547#2792..3066#2490..2632#2357..2411#2187..2264#2008..2094#ccgagccgtttctctccctctctctctcctcctccttcccctgcgcctcctccaccgcccgtgtcgccgccgcgcgcgtctatcccctcgccgccgccgccgtctacgccgcgtgaggtccggattggttgaaaggctgatcgaggtaatgcttttttatttttcccgtgcggttttagtttttctgtgctgtgcttggtggttttttctttaaaactatttttaggctggttttgatgatggcctttctaaggttgtcgtgtatggggggcgaggtggttcgggtggatttgtggtgttatgttgaggggatttttgtttggggggagagagttttggtggtagatgcggttgggattttgaagacgtgaaagggaaattagttttgaaatttgacggcgaggttgagctgattagtattgtaataattcgtggcatcggatagatcaacgcgagatggatgcttattactgcgaatttcaggaggtttttgtggtttttttaggcgagtagcgttgattctagctggattttgctgcttcgtggaggcgtttagatgtggtgcatacggctgtggatgtaatttgtgcggggataattactgaaaattaattcgcagtaagctagctctgctttaatattttttcccctgaaacagtaacaagggatcccttcctcatttcgttaacagtggaagtgaatacagacaccaaccgatgagcagagacacctgtaattaaacaccaaaagaagaacctgttttgttcctatatattttccatttctatgctttaggtaatatactttctggttcggtgtcatagtacccgaaatagaatctttgcttgccaaaaaaagaaagaataacaagatagaacaacatcattaaattctgtcttagacaatacatttataatcgaccaacaaaaacagaccgatgagtgacgtggaataacaaatgaagtataaatgtcacgaaaaacacatctattgattttgtgtgataagccataggagcctacgaggaaccaagtgaatggttctgtacaccacatccattagcattaccataaatttgcctatttctatgatgccttccttaacatatttatcgacatgtgagcttgcaccataactaaatgttatggtaaactttaccttataaatgattgcaacatgtaccttttcatttcgataattaaaaacgagaaagcttccacctttttcctagatatattttttcgtttgattatgaatttagttgtacttgacaataaaataatgcgcttgtaagattatgcaattttcgttgcattacctccctctcttttcatttgtcagtttttttctactccctccgtttcatattttaagtcgtttgacttttttcctagtcagactttgttaagtttgacaaagtttatagtaaaatttagcaatatctaaaatattaaattggtttcatcaaatctaacattgaatatattttgataatatgttttttttgtgttgaaaatacttctatatttttctataaacttagtcaaacttggaagaaatttcagtaagaaaaaagtcaaacggcttataatatgaaacggagggagtaaatttttttttgcattaattaaaattcatgtattttcctccttttttggtttttgttgtgcgatatattacagtgtagagactctgaagctcatatggttggcattggttgatatgataacgtgtcacgttgtgagatgcagacccccgttatgggggttgattgaatgaagtggatagcactcttataactttcttgctcgaagtttcttgataccaaggcaagtagcttataagaagatttggactttggtggcataaatggatgaactaactggcggcttcaacgtggtgttgattacaggtgcttttattgctaactggagcagtttgtgtagtttctttctttctctttgagaattctctcagtttcttgggcaggctcttgagctaccaagtatgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggtaacattattcctatttgattcttgttagaaattctatatgttgaacatacttgcctcaatgggaaacttggttgcgtctcctctctgaaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcaggtactgtatctctaagcatgtttagtatcttctgtgataaaatctttctccagtgtgttaacttttgcattctttttcttgcatgcttgcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattaggtaatttgatccagtgatcaatacatgttccacaaacaattgaagaaagagtttgacatatttttctcatatttccagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgaggtaaggcccctactctaatggttagttccaatttattggttcatttctctgtgtttgataagattttgcctctgctgtgcatttcaatattatatctaaactgtgctgcttatagcttatcaaactactcttgctcataaagatgcaacttttatgcagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgagtcataataaatgataacatattattgttctgtggactgaaaatctgaaatcctcctttcttgagaaatacaacctcaattttatggtgatattgacaagaaggaagagcgcatataatttaaagttaaaacacggaaaataatagaagtatgtgagcagaacgcaggagcaaggggtggaactcataacaagttcaaacttcatgcatgttcataattccccaatgcaagtatggcaagaaaaatagacaacttttaaaagtatgctaatggaactgcatgcattgtgtaaacttgcttttgtcaagcatgttggaacatactatatatttgctgctgatgatgaaaaatttgcttgagtactcttctattgctaatggacgcttaccattccctgttttgtcttcaaatgcatcaggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactggttggtaaaatcagcaatggaattttctatcgtgaattttttataattattgtaacattaattactcccatgtctttttattctgttgagataccatgtgacaatgtctacaattgcatattttgcattctttggtagatcagttcccttcttaagatccccaaagggaaactggagtatcaataacttactccctcaaataaaaaaaaagtgatactttaggcataaacacattcttcaaaacaaaactttgaatagcagtttcttttgtaatatattaataaagcatggaaaaattataatgtgaagaatgtacattcagggacaaatatactcatcatttttgcatggttgatcttaatgtccatagagagattgatcagaccaaaagtttatttttgacaggagtgagtatggattataataggagtgaatggcaatatgctgtgaaagtgagcattttatatttgtttactactagcatttggtgacaaagcaaatgatgtatattgtaaatcaaagtaaacaagtagaacatgtcctgtggcgacaagacataacctacaaattaaagttagcctgattgtttattttactgggagcacattttttctagattctagagtacatattattaaaaaaggtgtgatatgtttgcaaattactttttaatggaagtttgtaaattactttgaaacttgcattcagtttttttgcttcttaaaaaacatagtgcatatgggctttatgagctcatactatgcatgtttttctgcattggagtgggacctttatctttttcttgcaactcggatgctaatgcttattataaacacaagttactaacatgtgcttgggtcattcagcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcataggtaagctcttatgatcctgtccactgttgtaccatgtgtactttccaaatgattgctaacatgctatttttcgtcagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcgggtatatcatacatctctctggctgttgtttcagtgctatcaatcaagatgacacatcatgcaatattatcataacagcacattatgtgttttgaatattttattgattgtcatagctttgttttatgttttagcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcaggtatgcagtgtttcctgcacctacttgctacctattctgtatttgatgtattcaatttcctttttatcagttttctcgaaatatatgctaataggcttgtcactaaacggctacattatccagactctgttctgaatttttttttcattaagtactgttagctgccagattacatcatcatgactaaagtagtttagctacaatattgcaaagctagatttgtctagcctctattgttttgaatagaaaaggcaggtatgcagtgtttcctgcacctacttgctacctattctgtatttgatgtattcaatttcctttttatcagttttctcgaaatatatgctaataggcttgtcactaaacggctacattatccagactctgttctgaatttttttttcattaagtactgttagctgccagattacatcatcatgactaaagtagtttagctacaatattgcaaagctagatttgtctagcctctattgttttgaatagaaaattaaatcttttggtcatttttttgtaaccttttaggagtctctgtatgcttgtcgtcaacatgctatgataagtttatatcagttcttttggagctatgatgtattcttgtatattttatgactgctaggatattatgttaaaactgttttaaaggtatcatcaaacaagttaatatagatattgctacctaccaaagttggaatttgattatgtcgtaaaatatatgcaccctacatgttttagcttcatattgacttatattttcatatgtgctccttcaactgcacaaagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggtatggtcattcctgcttataagataccagaataatgctatgaagaccgtccttggttgtcagcattcttatacacaatgataatctttgtttttcatgggtaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggtcaagatattgttattgttatatttacatacttgttatcatatttaaaagcttcatattgtatgtacctttgaaaaaaaaagttccatactgtagcttcttttgttctcatttcttggtttcaaagcatcagcatcattaccgatataaaatatacccatttcctcagtcatcttccccgactcttgtatctttcacaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagaggtaacaccttcattcatgaggtccttaggtgcaaatcccagtgaagaataatcagttctactttgtatggttcatctcgcttactgaccggttcatctcgggatcaattacgagcagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaaggcatgtcctcttctctgatctcactcacgttcatgccaaacttactcatcatcatccatgatccatcatcctgactgaatcttgatatgcagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctgacaaggccacaccatgactgtcactgcaagcagctgctgtaggatcagcgagtaggaaaggatgacttgccggccagtttcctgagatgtgatttaagactgatattagctgatgttcccaagcatgatacggggcttgctcccaagatgtgatttttactttacttttggttaatgtgctaccgctgacattagatgttcaagcatatgaaaactgcttgtgctgtaacttgtatgtctgtaacgtaaaaatctttccatatggtaaccggggaagcagaggtatatccttttctgctgttgtatctatttccagtttcctgctgaacttgcaaatgaatattcagcaatttccgttttgctc</dnaseqindica>|<br />
Link = [http://www.ncbi.nlm.nih.gov/nuccore/NM_001048755.1 RefSeq:Os01g0182600]|<br />
}}<br />
[[Category:Genes]]<br />
[[Category:Japonica mRNA]]<br />
[[Category:Oryza Sativa Japonica Group]]<br />
[[Category:Japonica Genes]]<br />
[[Category:Japonica Chromosome 1]]<br />
[[Category:Chromosome 1]]</div>Mikahttps://ngdc.cncb.ac.cn/ricewiki/index.php?title=Os01g0182600&diff=183688Os01g01826002014-06-10T14:29:01Z<p>Mika: </p>
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<div>The gene of ''Gigantea(OsGI)''is solo ortholog of ''GI'' in the Arabidopsis circadian system and related to photoperiodic control of flowering.<br />
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==Annotated Information==<br />
===Function===<br />
*''OsGI'',a ''GI'' ortholog, regulates flowering time in response to photoperiodic conditions. Mutations in this gene delay flowering in inductive SDs (short-days),whereas flowering is marginally induced under LDs(long-days).In comparison, over expression of ''OsGI'' in transgenic rice causes late flowering in both SDs and LDs.In addition, ''OsGI'' has function to regulate ''Hd1'', a gene that also shows circadian rhythm<ref name="ref1" />. ''Os-GI'' affects gene expression of approximately 75% of genes among 27,201 genes and is required for strong amplitudes and proper phase-setting of global gene expression under natural field conditions<ref name="ref2" />. <br />
*The cDNA of ''OsGI'' firstly found in ''se5'' mutant that is completely photoperiod-insensitive, the expression patterns of rice and Arabidopsis ''GI'' gene are both photoperiodically regulated and are very similar under LD and SD conditions(FIG1)<ref name="ref3" />. ''GI'' is required for phase-dependent ''CO''transcription through the interaction with FKF1, which leads to cycling DNA binding with one finger (CDF) degradation and consequent derepression of CO. In addition, it has been known that GI protein interacts with ZEITLUPE (ZTL) F-box protein to control the diurnal rhythm of TOC1 protein in regulation of the Arabidopsis circadian clock<ref name="ref4" />. <br />
*''Os-GI'' also has function to maintain bioactive GA level through the regulation of the GA-deactivating enzyme genes in rice, ''Os-GI'' represses major ''OsGA2ox'' genes in order to maintain a proper bioactive GA level, ''osgi-1'' plants showed a semi-dwarf phenotype with elevated ''OsGA2ox'' gene expression(FIG2) <ref name="ref4" />.<br />
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===Expression===<br />
*''GI'' encodes a nuclear protein that defines the proper amplitude and period length of circadian rhythms. ''GI'' expression is regulated by the circadian clock, with a peak in transcript levels at 8 to 10 h after dawn. The ''GI'' protein acts between the circadian oscillator and ''CO'' to promote flowering by increasing ''CO'' and ''FT'' mRNA abundance<ref name="ref1" />. The expression of the ''OsGI'' messenger RNA was circadian-controlled and that its temporal expression pattern was very similar to that of ''GI'' under both SD and LD conditions. Decreased ''OsGI'' expression led to early flowering under LD conditions and late flowering under SD conditions relative to the wild type, and that this early flowering phenotype under LD conditions was the opposite of that of the Arabidopsis ''gi'' mutants, confirming the opposite role of the ''GI'' gene in flowering under LD conditions in rice and Arabidopsis<ref name="ref5" />.<br />
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===Mutation===<br />
*The single ''osgi-1'' mutation extended the time to flowering only under SD conditions. Under LD conditions the single ''hd1'' mutation induced early flowering, but ''osgi-1'' plants exhibited no phenotypic change<ref name="ref4" /><br />
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===Evolution===<br />
* There is some evolutionary conservation between rice circadian clock–related genes and those associated with Arabidopsis circadian clocks, where the circadian clock consists of interlocked subloops, it is also possible that ''Os-GI'' is a member of a subloop in rice circadian clocks<ref name="ref4" />.But there is a divergence in the photoperiodic pathways of these two species arises from the molecular mechanisms controlling the expression of ''FT/Hd3a'' by ''CO/Hd1(Se1)''<ref name="ref5" />(figure 3)<br />
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==Labs working on this gene==<br />
Please input related labs here.<br />
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==Knowledge Extension==<br />
*The rice orthologues of the two key regulators of flowering time in Arabidopsis, ''CO'' and ''FT'', were recently shown to be important in the photoperiodic control of flowering in rice. The rice ''Hd1(Se1)''gene, which is an orthologue of the Arabidopsis ''CO'' gene, was required for the suppression of flowering under LD conditions and for the promotion of flowering under SD conditions. Furthermore, the rice ''Hd3a'' gene, an orthologue of the Arabidopsis ''FT'' gene, was shown to be an activator of flowering in rice. It was previously demonstrated that ''CO'' controls the flowering time by integrating signals from the circadian clock and light to regulate ''FT'' expression in Arabidopsis<ref name="ref5" />.Unlike Arabidopsis ''FT'', expression of ''Hd3a'' mRNA is more or less opposite to that of ''Hd1'' under inductive short-day conditions<ref name="ref6" />.<br />
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==References==<br />
Please input cited references here.<br />
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==Structured Information==<br />
{{JaponicaGene|<br />
GeneName = Os01g0182600|<br />
Description = GIGANTEA protein|<br />
Version = NM_001048755.1 GI:115434923 GeneID:4325329|<br />
Length = 9083 bp|<br />
Definition = Oryza sativa Japonica Group Os01g0182600, complete gene.|<br />
Source = Oryza sativa Japonica Group<br />
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ORGANISM Oryza sativa Japonica Group<br />
Eukaryota; Viridiplantae; Streptophyta; Embryophyta; Tracheophyta;<br />
Spermatophyta; Magnoliophyta; Liliopsida; Poales; Poaceae; BEP<br />
clade; Ehrhartoideae; Oryzeae; Oryza.<br />
|<br />
Chromosome = [[:category:Japonica Chromosome 1|Chromosome 1]]|<br />
AP = Chromosome 1:4328362..4337444|<br />
CDS = 4328725..4329030,4329123..4329215,4329333..4329551,4329752..4329820,4329925..4331421<br>,4332220..4332429,4332563..4332713,4332791..4333031,4333898..4333956<br>,4334379..4334653,4334813..4334955,4335034..4335088,4335181..4335258<br>,4335351..4335437|<br />
GCID = <gbrowseImage1><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage1>|<br />
GSID = <gbrowseImage2><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage2>|<br />
CDNA = <cdnaseq>atgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactgcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcatagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcggcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctga</cdnaseq>|<br />
AA = <aaseq>MSASNEKWIDGLQFSSLFWPPPQDSQQKQAQILAYVEYFGQFTA DSEQFPEDIAQLIQSCYPSKEKRLVDEVLATFVLHHPEHGHAVVHPILSRIIDGTLSY DRNGFPFMSFISLFSHTSEKEYSEQWALACGEILRVLTHYNRPIFKVDHQHSEAECSS TSDQASSCESMEKRANGSPRNEPDRKPLRPLSPWITDILLAAPLGIRSDYFRWCGGVM GKYAAGGELKPPTTAYSRGSGKHPQLMPSTPRWAVANGAGVILSVCDEEVARYETANL TAAAVPALLLPPPTTPLDEHLVAGLPPLEPYARLFHRYYAIATPSATQRLLFGLLEAP PSWAPDALDAAVQLVELLRAAEDYDSGMRLPKNWMHLHFLRAIGTAMSMRAGIAADTS AALLFRILSQPTLLFPPLRHAEGVELHHEPLGGYVSSYKRQLEVPASEATIDATAQGI ASMLCAHGPDVEWRICTIWEAAYGLLPLSSSAVDLPEIVVAAPLQPPTLSWSLYLPLL KVFEYLPRGSPSEACLMRIFVATVEAILRRTFPSETSEQSRKPRSQSKNLAVAELRTM IHSLFVESCASMDLASRLLFVVLTVCVSHQALPGGSKRPTGSDNHSSEEVTNDSRLTN GRNRCKKRQGPVATFDSYVLAAVCALSCELQLFPFISKNGNHSNLKDSIKIVIPGKTT GISNELHNSISSAILHTRRILGILEALFSLKPSSVGTSWSYSSNEIVAAAMVAAHVSE LFRRSRPCLNALSALKQCKWDAEISTRASSLYHLIDLHGKTVTSIVNKAEPLEAHLTL TPVKKDEPPIEEKNINSSDGGALEKKDASRSHRKNGFARPLLKCAEDVILNGDVASTS GKAIASLQVEASDLANFLTMDRNGGYRGSQTLLRSVLSEKQELCFSVVSLLWQKLIAS PEMQMSAESTSAHQGWRKVVDALCDIVSASPTKASAAIVLQAEKDLQPWIARDDEQGQ KMWRVNQRIVKLIAELMRNHDSPEALVILASASDLLLRATDGMLVDGEACTLPQLELL EVTARAVHLIVEWGDSGVSVADGLSNLLKCRLSTTIRCLSHPSAHVRALSMSVLRDIL NSGQINSSKLIQGEHRNGIQSPTYQCLAASIINWQADVERCIEWEAHSRRATGLTLAF LTAAAKELGCPLTC</aaseq>|<br />
DNA = <dnaseqindica>8415..8720#8230..8322#7894..8112#7625..7693#6024..7520#5016..5225#4732..4882#4414..4654#3489..3547#2792..3066#2490..2632#2357..2411#2187..2264#2008..2094#ccgagccgtttctctccctctctctctcctcctccttcccctgcgcctcctccaccgcccgtgtcgccgccgcgcgcgtctatcccctcgccgccgccgccgtctacgccgcgtgaggtccggattggttgaaaggctgatcgaggtaatgcttttttatttttcccgtgcggttttagtttttctgtgctgtgcttggtggttttttctttaaaactatttttaggctggttttgatgatggcctttctaaggttgtcgtgtatggggggcgaggtggttcgggtggatttgtggtgttatgttgaggggatttttgtttggggggagagagttttggtggtagatgcggttgggattttgaagacgtgaaagggaaattagttttgaaatttgacggcgaggttgagctgattagtattgtaataattcgtggcatcggatagatcaacgcgagatggatgcttattactgcgaatttcaggaggtttttgtggtttttttaggcgagtagcgttgattctagctggattttgctgcttcgtggaggcgtttagatgtggtgcatacggctgtggatgtaatttgtgcggggataattactgaaaattaattcgcagtaagctagctctgctttaatattttttcccctgaaacagtaacaagggatcccttcctcatttcgttaacagtggaagtgaatacagacaccaaccgatgagcagagacacctgtaattaaacaccaaaagaagaacctgttttgttcctatatattttccatttctatgctttaggtaatatactttctggttcggtgtcatagtacccgaaatagaatctttgcttgccaaaaaaagaaagaataacaagatagaacaacatcattaaattctgtcttagacaatacatttataatcgaccaacaaaaacagaccgatgagtgacgtggaataacaaatgaagtataaatgtcacgaaaaacacatctattgattttgtgtgataagccataggagcctacgaggaaccaagtgaatggttctgtacaccacatccattagcattaccataaatttgcctatttctatgatgccttccttaacatatttatcgacatgtgagcttgcaccataactaaatgttatggtaaactttaccttataaatgattgcaacatgtaccttttcatttcgataattaaaaacgagaaagcttccacctttttcctagatatattttttcgtttgattatgaatttagttgtacttgacaataaaataatgcgcttgtaagattatgcaattttcgttgcattacctccctctcttttcatttgtcagtttttttctactccctccgtttcatattttaagtcgtttgacttttttcctagtcagactttgttaagtttgacaaagtttatagtaaaatttagcaatatctaaaatattaaattggtttcatcaaatctaacattgaatatattttgataatatgttttttttgtgttgaaaatacttctatatttttctataaacttagtcaaacttggaagaaatttcagtaagaaaaaagtcaaacggcttataatatgaaacggagggagtaaatttttttttgcattaattaaaattcatgtattttcctccttttttggtttttgttgtgcgatatattacagtgtagagactctgaagctcatatggttggcattggttgatatgataacgtgtcacgttgtgagatgcagacccccgttatgggggttgattgaatgaagtggatagcactcttataactttcttgctcgaagtttcttgataccaaggcaagtagcttataagaagatttggactttggtggcataaatggatgaactaactggcggcttcaacgtggtgttgattacaggtgcttttattgctaactggagcagtttgtgtagtttctttctttctctttgagaattctctcagtttcttgggcaggctcttgagctaccaagtatgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggtaacattattcctatttgattcttgttagaaattctatatgttgaacatacttgcctcaatgggaaacttggttgcgtctcctctctgaaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcaggtactgtatctctaagcatgtttagtatcttctgtgataaaatctttctccagtgtgttaacttttgcattctttttcttgcatgcttgcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattaggtaatttgatccagtgatcaatacatgttccacaaacaattgaagaaagagtttgacatatttttctcatatttccagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgaggtaaggcccctactctaatggttagttccaatttattggttcatttctctgtgtttgataagattttgcctctgctgtgcatttcaatattatatctaaactgtgctgcttatagcttatcaaactactcttgctcataaagatgcaacttttatgcagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgagtcataataaatgataacatattattgttctgtggactgaaaatctgaaatcctcctttcttgagaaatacaacctcaattttatggtgatattgacaagaaggaagagcgcatataatttaaagttaaaacacggaaaataatagaagtatgtgagcagaacgcaggagcaaggggtggaactcataacaagttcaaacttcatgcatgttcataattccccaatgcaagtatggcaagaaaaatagacaacttttaaaagtatgctaatggaactgcatgcattgtgtaaacttgcttttgtcaagcatgttggaacatactatatatttgctgctgatgatgaaaaatttgcttgagtactcttctattgctaatggacgcttaccattccctgttttgtcttcaaatgcatcaggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactggttggtaaaatcagcaatggaattttctatcgtgaattttttataattattgtaacattaattactcccatgtctttttattctgttgagataccatgtgacaatgtctacaattgcatattttgcattctttggtagatcagttcccttcttaagatccccaaagggaaactggagtatcaataacttactccctcaaataaaaaaaaagtgatactttaggcataaacacattcttcaaaacaaaactttgaatagcagtttcttttgtaatatattaataaagcatggaaaaattataatgtgaagaatgtacattcagggacaaatatactcatcatttttgcatggttgatcttaatgtccatagagagattgatcagaccaaaagtttatttttgacaggagtgagtatggattataataggagtgaatggcaatatgctgtgaaagtgagcattttatatttgtttactactagcatttggtgacaaagcaaatgatgtatattgtaaatcaaagtaaacaagtagaacatgtcctgtggcgacaagacataacctacaaattaaagttagcctgattgtttattttactgggagcacattttttctagattctagagtacatattattaaaaaaggtgtgatatgtttgcaaattactttttaatggaagtttgtaaattactttgaaacttgcattcagtttttttgcttcttaaaaaacatagtgcatatgggctttatgagctcatactatgcatgtttttctgcattggagtgggacctttatctttttcttgcaactcggatgctaatgcttattataaacacaagttactaacatgtgcttgggtcattcagcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcataggtaagctcttatgatcctgtccactgttgtaccatgtgtactttccaaatgattgctaacatgctatttttcgtcagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtac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/>
Link = [http://www.ncbi.nlm.nih.gov/nuccore/NM_001048755.1 RefSeq:Os01g0182600]|<br />
}}<br />
[[Category:Genes]]<br />
[[Category:Japonica mRNA]]<br />
[[Category:Oryza Sativa Japonica Group]]<br />
[[Category:Japonica Genes]]<br />
[[Category:Japonica Chromosome 1]]<br />
[[Category:Chromosome 1]]</div>Mikahttps://ngdc.cncb.ac.cn/ricewiki/index.php?title=Os01g0182600&diff=183679Os01g01826002014-06-10T14:18:14Z<p>Mika: </p>
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<div>Please input one-sentence summary here.<br />
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==Annotated Information==<br />
===Function===<br />
*''OsGI'',a ''GI'' ortholog, regulates flowering time in response to photoperiodic conditions. Mutations in this gene delay flowering in inductive SDs (short-days),whereas flowering is marginally induced under LDs(long-days).In comparison, over expression of ''OsGI'' in transgenic rice causes late flowering in both SDs and LDs.In addition, ''OsGI'' has function to regulate ''Hd1'', a gene that also shows circadian rhythm<ref name="ref1" />. ''Os-GI'' affects gene expression of approximately 75% of genes among 27,201 genes and is required for strong amplitudes and proper phase-setting of global gene expression under natural field conditions<ref name="ref2" />. <br />
*The cDNA of ''OsGI'' firstly found in ''se5'' mutant that is completely photoperiod-insensitive, the expression patterns of rice and Arabidopsis ''GI'' gene are both photoperiodically regulated and are very similar under LD and SD conditions(FIG1)<ref name="ref3" />. ''GI'' is required for phase-dependent ''CO''transcription through the interaction with FKF1, which leads to cycling DNA binding with one finger (CDF) degradation and consequent derepression of CO. In addition, it has been known that GI protein interacts with ZEITLUPE (ZTL) F-box protein to control the diurnal rhythm of TOC1 protein in regulation of the Arabidopsis circadian clock<ref name="ref4" />. <br />
*''Os-GI'' also has function to maintain bioactive GA level through the regulation of the GA-deactivating enzyme genes in rice, ''Os-GI'' represses major ''OsGA2ox'' genes in order to maintain a proper bioactive GA level, ''osgi-1'' plants showed a semi-dwarf phenotype with elevated ''OsGA2ox'' gene expression(FIG2) <ref name="ref4" />.<br />
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===Expression===<br />
*''GI'' encodes a nuclear protein that defines the proper amplitude and period length of circadian rhythms. ''GI'' expression is regulated by the circadian clock, with a peak in transcript levels at 8 to 10 h after dawn. The ''GI'' protein acts between the circadian oscillator and ''CO'' to promote flowering by increasing ''CO'' and ''FT'' mRNA abundance<ref name="ref1" />. The expression of the ''OsGI'' messenger RNA was circadian-controlled and that its temporal expression pattern was very similar to that of ''GI'' under both SD and LD conditions. Decreased ''OsGI'' expression led to early flowering under LD conditions and late flowering under SD conditions relative to the wild type, and that this early flowering phenotype under LD conditions was the opposite of that of the Arabidopsis ''gi'' mutants, confirming the opposite role of the ''GI'' gene in flowering under LD conditions in rice and Arabidopsis<ref name="ref5" />.<br />
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===Mutation===<br />
*The single ''osgi-1'' mutation extended the time to flowering only under SD conditions. Under LD conditions the single ''hd1'' mutation induced early flowering, but ''osgi-1'' plants exhibited no phenotypic change<ref name="ref4" /><br />
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===Evolution===<br />
* There is some evolutionary conservation between rice circadian clock–related genes and those associated with Arabidopsis circadian clocks, where the circadian clock consists of interlocked subloops, it is also possible that ''Os-GI'' is a member of a subloop in rice circadian clocks<ref name="ref4" />.But there is a divergence in the photoperiodic pathways of these two species arises from the molecular mechanisms controlling the expression of ''FT/Hd3a'' by ''CO/Hd1(Se1)''<ref name="ref5" />(figure 3)<br />
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==Labs working on this gene==<br />
Please input related labs here.<br />
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==Knowledge Extension==<br />
*The rice orthologues of the two key regulators of flowering time in Arabidopsis, ''CO'' and ''FT'', were recently shown to be important in the photoperiodic control of flowering in rice. The rice ''Hd1(Se1)''gene, which is an orthologue of the Arabidopsis ''CO'' gene, was required for the suppression of flowering under LD conditions and for the promotion of flowering under SD conditions. Furthermore, the rice ''Hd3a'' gene, an orthologue of the Arabidopsis ''FT'' gene, was shown to be an activator of flowering in rice. It was previously demonstrated that ''CO'' controls the flowering time by integrating signals from the circadian clock and light to regulate ''FT'' expression in Arabidopsis<ref name="ref5" />.Unlike Arabidopsis ''FT'', expression of ''Hd3a'' mRNA is more or less opposite to that of ''Hd1'' under inductive short-day conditions<ref name="ref6" />.<br />
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==References==<br />
Please input cited references here.<br />
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==Structured Information==<br />
{{JaponicaGene|<br />
GeneName = Os01g0182600|<br />
Description = GIGANTEA protein|<br />
Version = NM_001048755.1 GI:115434923 GeneID:4325329|<br />
Length = 9083 bp|<br />
Definition = Oryza sativa Japonica Group Os01g0182600, complete gene.|<br />
Source = Oryza sativa Japonica Group<br />
<br />
ORGANISM Oryza sativa Japonica Group<br />
Eukaryota; Viridiplantae; Streptophyta; Embryophyta; Tracheophyta;<br />
Spermatophyta; Magnoliophyta; Liliopsida; Poales; Poaceae; BEP<br />
clade; Ehrhartoideae; Oryzeae; Oryza.<br />
|<br />
Chromosome = [[:category:Japonica Chromosome 1|Chromosome 1]]|<br />
AP = Chromosome 1:4328362..4337444|<br />
CDS = 4328725..4329030,4329123..4329215,4329333..4329551,4329752..4329820,4329925..4331421<br>,4332220..4332429,4332563..4332713,4332791..4333031,4333898..4333956<br>,4334379..4334653,4334813..4334955,4335034..4335088,4335181..4335258<br>,4335351..4335437|<br />
GCID = <gbrowseImage1><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage1>|<br />
GSID = <gbrowseImage2><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage2>|<br />
CDNA = <cdnaseq>atgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactgcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcatagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcggcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctga</cdnaseq>|<br />
AA = <aaseq>MSASNEKWIDGLQFSSLFWPPPQDSQQKQAQILAYVEYFGQFTA DSEQFPEDIAQLIQSCYPSKEKRLVDEVLATFVLHHPEHGHAVVHPILSRIIDGTLSY DRNGFPFMSFISLFSHTSEKEYSEQWALACGEILRVLTHYNRPIFKVDHQHSEAECSS TSDQASSCESMEKRANGSPRNEPDRKPLRPLSPWITDILLAAPLGIRSDYFRWCGGVM GKYAAGGELKPPTTAYSRGSGKHPQLMPSTPRWAVANGAGVILSVCDEEVARYETANL TAAAVPALLLPPPTTPLDEHLVAGLPPLEPYARLFHRYYAIATPSATQRLLFGLLEAP PSWAPDALDAAVQLVELLRAAEDYDSGMRLPKNWMHLHFLRAIGTAMSMRAGIAADTS AALLFRILSQPTLLFPPLRHAEGVELHHEPLGGYVSSYKRQLEVPASEATIDATAQGI ASMLCAHGPDVEWRICTIWEAAYGLLPLSSSAVDLPEIVVAAPLQPPTLSWSLYLPLL KVFEYLPRGSPSEACLMRIFVATVEAILRRTFPSETSEQSRKPRSQSKNLAVAELRTM IHSLFVESCASMDLASRLLFVVLTVCVSHQALPGGSKRPTGSDNHSSEEVTNDSRLTN GRNRCKKRQGPVATFDSYVLAAVCALSCELQLFPFISKNGNHSNLKDSIKIVIPGKTT GISNELHNSISSAILHTRRILGILEALFSLKPSSVGTSWSYSSNEIVAAAMVAAHVSE LFRRSRPCLNALSALKQCKWDAEISTRASSLYHLIDLHGKTVTSIVNKAEPLEAHLTL TPVKKDEPPIEEKNINSSDGGALEKKDASRSHRKNGFARPLLKCAEDVILNGDVASTS GKAIASLQVEASDLANFLTMDRNGGYRGSQTLLRSVLSEKQELCFSVVSLLWQKLIAS PEMQMSAESTSAHQGWRKVVDALCDIVSASPTKASAAIVLQAEKDLQPWIARDDEQGQ KMWRVNQRIVKLIAELMRNHDSPEALVILASASDLLLRATDGMLVDGEACTLPQLELL EVTARAVHLIVEWGDSGVSVADGLSNLLKCRLSTTIRCLSHPSAHVRALSMSVLRDIL NSGQINSSKLIQGEHRNGIQSPTYQCLAASIINWQADVERCIEWEAHSRRATGLTLAF LTAAAKELGCPLTC</aaseq>|<br />
DNA = <dnaseqindica>8415..8720#8230..8322#7894..8112#7625..7693#6024..7520#5016..5225#4732..4882#4414..4654#3489..3547#2792..3066#2490..2632#2357..2411#2187..2264#2008..2094#ccgagccgtttctctccctctctctctcctcctccttcccctgcgcctcctccaccgcccgtgtcgccgccgcgcgcgtctatcccctcgccgccgccgccgtctacgccgcgtgaggtccggattggttgaaaggctgatcgaggtaatgcttttttatttttcccgtgcggttttagtttttctgtgctgtgcttggtggttttttctttaaaactatttttaggctggttttgatgatggcctttctaaggttgtcgtgtatggggggcgaggtggttcgggtggatttgtggtgttatgttgaggggatttttgtttggggggagagagttttggtggtagatgcggttgggattttgaagacgtgaaagggaaattagttttgaaatttgacggcgaggttgagctgattagtattgtaataattcgtggcatcggatagatcaacgcgagatggatgcttattactgcgaatttcaggaggtttttgtggtttttttaggcgagtagcgttgattctagctggattttgctgcttcgtggaggcgtttagatgtggtgcatacggctgtggatgtaatttgtgcggggataattactgaaaattaattcgcagtaagctagctctgctttaatattttttcccctgaaacagtaacaagggatcccttcctcatttcgttaacagtggaagtgaatacagacaccaaccgatgagcagagacacctgtaattaaacaccaaaagaagaacctgttttgttcctatatattttccatttctatgctttaggtaatatactttctggttcggtgtcatagtacccgaaatagaatctttgcttgccaaaaaaagaaagaataacaagatagaacaacatcattaaattctgtcttagacaatacatttataatcgaccaacaaaaacagaccgatgagtgacgtggaataacaaatgaagtataaatgtcacgaaaaacacatctattgattttgtgtgataagccataggagcctacgaggaaccaagtgaatggttctgtacaccacatccattagcattaccataaatttgcctatttctatgatgccttccttaacatatttatcgacatgtgagcttgcaccataactaaatgttatggtaaactttaccttataaatgattgcaacatgtaccttttcatttcgataattaaaaacgagaaagcttccacctttttcctagatatattttttcgtttgattatgaatttagttgtacttgacaataaaataatgcgcttgtaagattatgcaattttcgttgcattacctccctctcttttcatttgtcagtttttttctactccctccgtttcatattttaagtcgtttgacttttttcctagtcagactttgttaagtttgacaaagtttatagtaaaatttagcaatatctaaaatattaaattggtttcatcaaatctaacattgaatatattttgataatatgttttttttgtgttgaaaatacttctatatttttctataaacttagtcaaacttggaagaaatttcagtaagaaaaaagtcaaacggcttataatatgaaacggagggagtaaatttttttttgcattaattaaaattcatgtattttcctccttttttggtttttgttgtgcgatatattacagtgtagagactctgaagctcatatggttggcattggttgatatgataacgtgtcacgttgtgagatgcagacccccgttatgggggttgattgaatgaagtggatagcactcttataactttcttgctcgaagtttcttgataccaaggcaagtagcttataagaagatttggactttggtggcataaatggatgaactaactggcggcttcaacgtggtgttgattacaggtgcttttattgctaactggagcagtttgtgtagtttctttctttctctttgagaattctctcagtttcttgggcaggctcttgagctaccaagtatgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggtaacattattcctatttgattcttgttagaaattctatatgttgaacatacttgcctcaatgggaaacttggttgcgtctcctctctgaaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcaggtactgtatctctaagcatgtttagtatcttctgtgataaaatctttctccagtgtgttaacttttgcattctttttcttgcatgcttgcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattaggtaatttgatccagtgatcaatacatgttccacaaacaattgaagaaagagtttgacatatttttctcatatttccagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgaggtaaggcccctactctaatggttagttccaatttattggttcatttctctgtgtttgataagattttgcctctgctgtgcatttcaatattatatctaaactgtgctgcttatagcttatcaaactactcttgctcataaagatgcaacttttatgcagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgagtcataataaatgataacatattattgttctgtggactgaaaatctgaaatcctcctttcttgagaaatacaacctcaattttatggtgatattgacaagaaggaagagcgcatataatttaaagttaaaacacggaaaataatagaagtatgtgagcagaacgcaggagcaaggggtggaactcataacaagttcaaacttcatgcatgttcataattccccaatgcaagtatggcaagaaaaatagacaacttttaaaagtatgctaatggaactgcatgcattgtgtaaacttgcttttgtcaagcatgttggaacatactatatatttgctgctgatgatgaaaaatttgcttgagtactcttctattgctaatggacgcttaccattccctgttttgtcttcaaatgcatcaggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactggttggtaaaatcagcaatggaattttctatcgtgaattttttataattattgtaacattaattactcccatgtctttttattctgttgagataccatgtgacaatgtctacaattgcatattttgcattctttggtagatcagttcccttcttaagatccccaaagggaaactggagtatcaataacttactccctcaaataaaaaaaaagtgatactttaggcataaacacattcttcaaaacaaaactttgaatagcagtttcttttgtaatatattaataaagcatggaaaaattataatgtgaagaatgtacattcagggacaaatatactcatcatttttgcatggttgatcttaatgtccatagagagattgatcagaccaaaagtttatttttgacaggagtgagtatggattataataggagtgaatggcaatatgctgtgaaagtgagcattttatatttgtttactactagcatttggtgacaaagcaaatgatgtatattgtaaatcaaagtaaacaagtagaacatgtcctgtggcgacaagacataacctacaaattaaagttagcctgattgtttattttactgggagcacattttttctagattctagagtacatattattaaaaaaggtgtgatatgtttgcaaattactttttaatggaagtttgtaaattactttgaaacttgcattcagtttttttgcttcttaaaaaacatagtgcatatgggctttatgagctcatactatgcatgtttttctgcattggagtgggacctttatctttttcttgcaactcggatgctaatgcttattataaacacaagttactaacatgtgcttgggtcattcagcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcataggtaagctcttatgatcctgtccactgttgtaccatgtgtactttccaaatgattgctaacatgctatttttcgtcagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcgggtatatcatacatctctctggctgttgtttcagtgctatcaatcaagatgacacatcatgcaatattatcataacagcacattatgtgttttgaatattttattgattgtcatagctttgttttatgttttagcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcaggtatgcagtgtttcctgcacctacttgctacctattctgtatttgatgtattcaatttcctttttatcagttttctcgaaatatatgctaataggcttgtcactaaacggctacattatccagactctgttctgaatttttttttcattaagtactgttagctgccagattacatcatcatgactaaagtagtttagctacaatattgcaaagctagatttgtctagcctctattgttttgaatagaaaaggcaggtatgcagtgtttcctgcacctacttgctacctattctgtatttgatgtattcaatttcctttttatcagttttctcgaaatatatgctaataggcttgtcactaaacggctacattatccagactctgttctgaatttttttttcattaagtactgttagctgccagattacatcatcatgactaaagtagtttagctacaatattgcaaagctagatttgtctagcctctattgttttgaatagaaaattaaatcttttggtcatttttttgtaaccttttaggagtctctgtatgcttgtcgtcaacatgctatgataagtttatatcagttcttttggagctatgatgtattcttgtatattttatgactgctaggatattatgttaaaactgttttaaaggtatcatcaaacaagttaatatagatattgctacctaccaaagttggaatttgattatgtcgtaaaatatatgcaccctacatgttttagcttcatattgacttatattttcatatgtgctccttcaactgcacaaagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggtatggtcattcctgcttataagataccagaataatgctatgaagaccgtccttggttgtcagcattcttatacacaatgataatctttgtttttcatgggtaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggtcaagatattgttattgttatatttacatacttgttatcatatttaaaagcttcatattgtatgtacctttgaaaaaaaaagttccatactgtagcttcttttgttctcatttcttggtttcaaagcatcagcatcattaccgatataaaatatacccatttcctcagtcatcttccccgactcttgtatctttcacaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagaggtaacaccttcattcatgaggtccttaggtgcaaatcccagtgaagaataatcagttctactttgtatggttcatctcgcttactgaccggttcatctcgggatcaattacgagcagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaaggcatgtcctcttctctgatctcactcacgttcatgccaaacttactcatcatcatccatgatccatcatcctgactgaatcttgatatgcagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctgacaaggccacaccatgactgtcactgcaagcagctgctgtaggatcagcgagtaggaaaggatgacttgccggccagtttcctgagatgtgatttaagactgatattagctgatgttcccaagcatgatacggggcttgctcccaagatgtgatttttactttacttttggttaatgtgctaccgctgacattagatgttcaagcatatgaaaactgcttgtgctgtaacttgtatgtctgtaacgtaaaaatctttccatatggtaaccggggaagcagaggtatatccttttctgctgttgtatctatttccagtttcctgctgaacttgcaaatgaatattcagcaatttccgttttgctc</dnaseqindica>|<br />
Link = [http://www.ncbi.nlm.nih.gov/nuccore/NM_001048755.1 RefSeq:Os01g0182600]|<br />
}}<br />
[[Category:Genes]]<br />
[[Category:Japonica mRNA]]<br />
[[Category:Oryza Sativa Japonica Group]]<br />
[[Category:Japonica Genes]]<br />
[[Category:Japonica Chromosome 1]]<br />
[[Category:Chromosome 1]]</div>Mikahttps://ngdc.cncb.ac.cn/ricewiki/index.php?title=Os01g0182600&diff=183678Os01g01826002014-06-10T14:16:41Z<p>Mika: </p>
<hr />
<div>Please input one-sentence summary here.<br />
<br />
==Annotated Information==<br />
===Function===<br />
*''OsGI'',a ''GI'' ortholog, regulates flowering time in response to photoperiodic conditions. Mutations in this gene delay flowering in inductive SDs (short-days),whereas flowering is marginally induced under LDs(long-days).In comparison, over expression of ''OsGI'' in transgenic rice causes late flowering in both SDs and LDs.In addition, ''OsGI'' has function to regulate ''Hd1'', a gene that also shows circadian rhythm<ref name="ref1" />. ''Os-GI'' affects gene expression of approximately 75% of genes among 27,201 genes and is required for strong amplitudes and proper phase-setting of global gene expression under natural field conditions<ref name="ref2" />. <br />
*The cDNA of ''OsGI'' firstly found in ''se5'' mutant that is completely photoperiod-insensitive, the expression patterns of rice and Arabidopsis ''GI'' gene are both photoperiodically regulated and are very similar under LD and SD conditions(FIG1)<ref name="ref3" />. ''GI'' is required for phase-dependent ''CO''transcription through the interaction with FKF1, which leads to cycling DNA binding with one finger (CDF) degradation and consequent derepression of CO. In addition, it has been known that GI protein interacts with ZEITLUPE (ZTL) F-box protein to control the diurnal rhythm of TOC1 protein in regulation of the Arabidopsis circadian clock<ref name="ref4" />. <br />
*''Os-GI'' also has function to maintain bioactive GA level through the regulation of the GA-deactivating enzyme genes in rice, ''Os-GI'' represses major ''OsGA2ox'' genes in order to maintain a proper bioactive GA level, ''osgi-1'' plants showed a semi-dwarf phenotype with elevated ''OsGA2ox'' gene expression(FIG2) <ref name="ref4" />.<br />
<br />
===Expression===<br />
*''GI'' encodes a nuclear protein that defines the proper amplitude and period length of circadian rhythms. ''GI'' expression is regulated by the circadian clock, with a peak in transcript levels at 8 to 10 h after dawn. The ''GI'' protein acts between the circadian oscillator and ''CO'' to promote flowering by increasing ''CO'' and ''FT'' mRNA abundance<ref name="ref1" />. The expression of the ''OsGI'' messenger RNA was circadian-controlled and that its temporal expression pattern was very similar to that of ''GI'' under both SD and LD conditions. Decreased ''OsGI'' expression led to early flowering under LD conditions and late flowering under SD conditions relative to the wild type, and that this early flowering phenotype under LD conditions was the opposite of that of the Arabidopsis ''gi'' mutants, confirming the opposite role of the ''GI'' gene in flowering under LD conditions in rice and Arabidopsis<ref name="ref5" />.<br />
<br />
='''=Mutation'''==<br />
*The single ''osgi-1'' mutation extended the time to flowering only under SD conditions. Under LD conditions the single ''hd1'' mutation induced early flowering, but ''osgi-1'' plants exhibited no phenotypic change<ref name="ref4" /><br />
<br />
===Evolution===<br />
* There is some evolutionary conservation between rice circadian clock–related genes and those associated with Arabidopsis circadian clocks, where the circadian clock consists of interlocked subloops, it is also possible that ''Os-GI'' is a member of a subloop in rice circadian clocks<ref name="ref4" />.But there is a divergence in the photoperiodic pathways of these two species arises from the molecular mechanisms controlling the expression of ''FT/Hd3a'' by ''CO/Hd1(Se1)''<ref name="ref5" />(figure 3)<br />
<br />
==Labs working on this gene==<br />
Please input related labs here.<br />
<br />
==Knowledge Extension==<br />
*The rice orthologues of the two key regulators of flowering time in Arabidopsis, ''CO'' and ''FT'', were recently shown to be important in the photoperiodic control of flowering in rice. The rice ''Hd1(Se1)''gene, which is an orthologue of the Arabidopsis ''CO'' gene, was required for the suppression of flowering under LD conditions and for the promotion of flowering under SD conditions. Furthermore, the rice ''Hd3a'' gene, an orthologue of the Arabidopsis ''FT'' gene, was shown to be an activator of flowering in rice. It was previously demonstrated that ''CO'' controls the flowering time by integrating signals from the circadian clock and light to regulate ''FT'' expression in Arabidopsis<ref name="ref5" />.Unlike Arabidopsis ''FT'', expression of ''Hd3a'' mRNA is more or less opposite to that of ''Hd1'' under inductive short-day conditions<ref name="ref6" />.<br />
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==References==<br />
Please input cited references here.<br />
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==Structured Information==<br />
{{JaponicaGene|<br />
GeneName = Os01g0182600|<br />
Description = GIGANTEA protein|<br />
Version = NM_001048755.1 GI:115434923 GeneID:4325329|<br />
Length = 9083 bp|<br />
Definition = Oryza sativa Japonica Group Os01g0182600, complete gene.|<br />
Source = Oryza sativa Japonica Group<br />
<br />
ORGANISM Oryza sativa Japonica Group<br />
Eukaryota; Viridiplantae; Streptophyta; Embryophyta; Tracheophyta;<br />
Spermatophyta; Magnoliophyta; Liliopsida; Poales; Poaceae; BEP<br />
clade; Ehrhartoideae; Oryzeae; Oryza.<br />
|<br />
Chromosome = [[:category:Japonica Chromosome 1|Chromosome 1]]|<br />
AP = Chromosome 1:4328362..4337444|<br />
CDS = 4328725..4329030,4329123..4329215,4329333..4329551,4329752..4329820,4329925..4331421<br>,4332220..4332429,4332563..4332713,4332791..4333031,4333898..4333956<br>,4334379..4334653,4334813..4334955,4335034..4335088,4335181..4335258<br>,4335351..4335437|<br />
GCID = <gbrowseImage1><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage1>|<br />
GSID = <gbrowseImage2><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage2>|<br />
CDNA = <cdnaseq>atgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactgcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcatagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcggcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctga</cdnaseq>|<br />
AA = <aaseq>MSASNEKWIDGLQFSSLFWPPPQDSQQKQAQILAYVEYFGQFTA DSEQFPEDIAQLIQSCYPSKEKRLVDEVLATFVLHHPEHGHAVVHPILSRIIDGTLSY DRNGFPFMSFISLFSHTSEKEYSEQWALACGEILRVLTHYNRPIFKVDHQHSEAECSS TSDQASSCESMEKRANGSPRNEPDRKPLRPLSPWITDILLAAPLGIRSDYFRWCGGVM GKYAAGGELKPPTTAYSRGSGKHPQLMPSTPRWAVANGAGVILSVCDEEVARYETANL TAAAVPALLLPPPTTPLDEHLVAGLPPLEPYARLFHRYYAIATPSATQRLLFGLLEAP PSWAPDALDAAVQLVELLRAAEDYDSGMRLPKNWMHLHFLRAIGTAMSMRAGIAADTS AALLFRILSQPTLLFPPLRHAEGVELHHEPLGGYVSSYKRQLEVPASEATIDATAQGI ASMLCAHGPDVEWRICTIWEAAYGLLPLSSSAVDLPEIVVAAPLQPPTLSWSLYLPLL KVFEYLPRGSPSEACLMRIFVATVEAILRRTFPSETSEQSRKPRSQSKNLAVAELRTM IHSLFVESCASMDLASRLLFVVLTVCVSHQALPGGSKRPTGSDNHSSEEVTNDSRLTN GRNRCKKRQGPVATFDSYVLAAVCALSCELQLFPFISKNGNHSNLKDSIKIVIPGKTT GISNELHNSISSAILHTRRILGILEALFSLKPSSVGTSWSYSSNEIVAAAMVAAHVSE LFRRSRPCLNALSALKQCKWDAEISTRASSLYHLIDLHGKTVTSIVNKAEPLEAHLTL TPVKKDEPPIEEKNINSSDGGALEKKDASRSHRKNGFARPLLKCAEDVILNGDVASTS GKAIASLQVEASDLANFLTMDRNGGYRGSQTLLRSVLSEKQELCFSVVSLLWQKLIAS PEMQMSAESTSAHQGWRKVVDALCDIVSASPTKASAAIVLQAEKDLQPWIARDDEQGQ KMWRVNQRIVKLIAELMRNHDSPEALVILASASDLLLRATDGMLVDGEACTLPQLELL EVTARAVHLIVEWGDSGVSVADGLSNLLKCRLSTTIRCLSHPSAHVRALSMSVLRDIL NSGQINSSKLIQGEHRNGIQSPTYQCLAASIINWQADVERCIEWEAHSRRATGLTLAF LTAAAKELGCPLTC</aaseq>|<br />
DNA = <dnaseqindica>8415..8720#8230..8322#7894..8112#7625..7693#6024..7520#5016..5225#4732..4882#4414..4654#3489..3547#2792..3066#2490..2632#2357..2411#2187..2264#2008..2094#ccgagccgtttctctccctctctctctcctcctccttcccctgcgcctcctccaccgcccgtgtcgccgccgcgcgcgtctatcccctcgccgccgccgccgtctacgccgcgtgaggtccggattggttgaaaggctgatcgaggtaatgcttttttatttttcccgtgcggttttagtttttctgtgctgtgcttggtggttttttctttaaaactatttttaggctggttttgatgatggcctttctaaggttgtcgtgtatggggggcgaggtggttcgggtggatttgtggtgttatgttgaggggatttttgtttggggggagagagttttggtggtagatgcggttgggattttgaagacgtgaaagggaaattagttttgaaatttgacggcgaggttgagctgattagtattgtaataattcgtggcatcggatagatcaacgcgagatggatgcttattactgcgaatttcaggaggtttttgtggtttttttaggcgagtagcgttgattctagctggattttgctgcttcgtggaggcgtttagatgtggtgcatacggctgtggatgtaatttgtgcggggataattactgaaaattaattcgcagtaagctagctctgctttaatattttttcccctgaaacagtaacaagggatcccttcctcatttcgttaacagtggaagtgaatacagacaccaaccgatgagcagagacacctgtaattaaacaccaaaagaagaacctgttttgttcctatatattttccatttctatgctttaggtaatatactttctggttcggtgtcatagtacccgaaatagaatctttgcttgccaaaaaaagaaagaataacaagatagaacaacatcattaaattctgtcttagacaatacatttataatcgaccaacaaaaacagaccgatgagtgacgtggaataacaaatgaagtataaatgtcacgaaaaacacatctattgattttgtgtgataagccataggagcctacgaggaaccaagtgaatggttctgtacaccacatccattagcattaccataaatttgcctatttctatgatgccttccttaacatatttatcgacatgtgagcttgcaccataactaaatgttatggtaaactttaccttataaatgattgcaacatgtaccttttcatttcgataattaaaaacgagaaagcttccacctttttcctagatatattttttcgtttgattatgaatttagttgtacttgacaataaaataatgcgcttgtaagattatgcaattttcgttgcattacctccctctcttttcatttgtcagtttttttctactccctccgtttcatattttaagtcgtttgacttttttcctagtcagactttgttaagtttgacaaagtttatagtaaaatttagcaatatctaaaatattaaattggtttcatcaaatctaacattgaatatattttgataatatgttttttttgtgttgaaaatacttctatatttttctataaacttagtcaaacttggaagaaatttcagtaagaaaaaagtcaaacggcttataatatgaaacggagggagtaaatttttttttgcattaattaaaattcatgtattttcctccttttttggtttttgttgtgcgatatattacagtgtagagactctgaagctcatatggttggcattggttgatatgataacgtgtcacgttgtgagatgcagacccccgttatgggggttgattgaatgaagtggatagcactcttataactttcttgctcgaagtttcttgataccaaggcaagtagcttataagaagatttggactttggtggcataaatggatgaactaactggcggcttcaacgtggtgttgattacaggtgcttttattgctaactggagcagtttgtgtagtttctttctttctctttgagaattctctcagtttcttgggcaggctcttgagctaccaagtatgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggtaacattattcctatttgattcttgttagaaattctatatgttgaacatacttgcctcaatgggaaacttggttgcgtctcctctctgaaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcaggtactgtatctctaagcatgtttagtatcttctgtgataaaatctttctccagtgtgttaacttttgcattctttttcttgcatgcttgcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattaggtaatttgatccagtgatcaatacatgttccacaaacaattgaagaaagagtttgacatatttttctcatatttccagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgaggtaaggcccctactctaatggttagttccaatttattggttcatttctctgtgtttgataagattttgcctctgctgtgcatttcaatattatatctaaactgtgctgcttatagcttatcaaactactcttgctcataaagatgcaacttttatgcagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgagtcataataaatgataacatattattgttctgtggactgaaaatctgaaatcctcctttcttgagaaatacaacctcaattttatggtgatattgacaagaaggaagagcgcatataatttaaagttaaaacacggaaaataatagaagtatgtgagcagaacgcaggagcaaggggtggaactcataacaagttcaaacttcatgcatgttcataattccccaatgcaagtatggcaagaaaaatagacaacttttaaaagtatgctaatggaactgcatgcattgtgtaaacttgcttttgtcaagcatgttggaacatactatatatttgctgctgatgatgaaaaatttgcttgagtactcttctattgctaatggacgcttaccattccctgttttgtcttcaaatgcatcaggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactggttggtaaaatcagcaatggaattttctatcgtgaattttttataattattgtaacattaattactcccatgtctttttattctgttgagataccatgtgacaatgtctacaattgcatattttgcattctttggtagatcagttcccttcttaagatccccaaagggaaactggagtatcaataacttactccctcaaataaaaaaaaagtgatactttaggcataaacacattcttcaaaacaaaactttgaatagcagtttcttttgtaatatattaataaagcatggaaaaattataatgtgaagaatgtacattcagggacaaatatactcatcatttttgcatggttgatcttaatgtccatagagagattgatcagaccaaaagtttatttttgacaggagtgagtatggattataataggagtgaatggcaatatgctgtgaaagtgagcattttatatttgtttactactagcatttggtgacaaagcaaatgatgtatattgtaaatcaaagtaaacaagtagaacatgtcctgtggcgacaagacataacctacaaattaaagttagcctgattgtttattttactgggagcacattttttctagattctagagtacatattattaaaaaaggtgtgatatgtttgcaaattactttttaatggaagtttgtaaattactttgaaacttgcattcagtttttttgcttcttaaaaaacatagtgcatatgggctttatgagctcatactatgcatgtttttctgcattggagtgggacctttatctttttcttgcaactcggatgctaatgcttattataaacacaagttactaacatgtgcttgggtcattcagcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcataggtaagctcttatgatcctgtccactgttgtaccatgtgtactttccaaatgattgctaacatgctatttttcgtcagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcgggtatatcatacatctctctggctgttgtttcagtgctatcaatcaagatgacacatcatgcaatattatcataacagcacattatgtgttttgaatattttattgattgtcatagctttgttttatgttttagcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcaggtatgcagtgtttcctgcacctacttgctacctattctgtatttgatgtattcaatttcctttttatcagttttctcgaaatatatgctaataggcttgtcactaaacggctacattatccagactctgttctgaatttttttttcattaagtactgttagctgccagattacatcatcatgactaaagtagtttagctacaatattgcaaagctagatttgtctagcctctattgttttgaatagaaaaggcaggtatgcagtgtttcctgcacctacttgctacctattctgtatttgatgtattcaatttcctttttatcagttttctcgaaatatatgctaataggcttgtcactaaacggctacattatccagactctgttctgaatttttttttcattaagtactgttagctgccagattacatcatcatgactaaagtagtttagctacaatattgcaaagctagatttgtctagcctctattgttttgaatagaaaattaaatcttttggtcatttttttgtaaccttttaggagtctctgtatgcttgtcgtcaacatgctatgataagtttatatcagttcttttggagctatgatgtattcttgtatattttatgactgctaggatattatgttaaaactgttttaaaggtatcatcaaacaagttaatatagatattgctacctaccaaagttggaatttgattatgtcgtaaaatatatgcaccctacatgttttagcttcatattgacttatattttcatatgtgctccttcaactgcacaaagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggtatggtcattcctgcttataagataccagaataatgctatgaagaccgtccttggttgtcagcattcttatacacaatgataatctttgtttttcatgggtaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggtcaagatattgttattgttatatttacatacttgttatcatatttaaaagcttcatattgtatgtacctttgaaaaaaaaagttccatactgtagcttcttttgttctcatttcttggtttcaaagcatcagcatcattaccgatataaaatatacccatttcctcagtcatcttccccgactcttgtatctttcacaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagaggtaacaccttcattcatgaggtccttaggtgcaaatcccagtgaagaataatcagttctactttgtatggttcatctcgcttactgaccggttcatctcgggatcaattacgagcagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaaggcatgtcctcttctctgatctcactcacgttcatgccaaacttactcatcatcatccatgatccatcatcctgactgaatcttgatatgcagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctgacaaggccacaccatgactgtcactgcaagcagctgctgtaggatcagcgagtaggaaaggatgacttgccggccagtttcctgagatgtgatttaagactgatattagctgatgttcccaagcatgatacggggcttgctcccaagatgtgatttttactttacttttggttaatgtgctaccgctgacattagatgttcaagcatatgaaaactgcttgtgctgtaacttgtatgtctgtaacgtaaaaatctttccatatggtaaccggggaagcagaggtatatccttttctgctgttgtatctatttccagtttcctgctgaacttgcaaatgaatattcagcaatttccgttttgctc</dnaseqindica>|<br />
Link = [http://www.ncbi.nlm.nih.gov/nuccore/NM_001048755.1 RefSeq:Os01g0182600]|<br />
}}<br />
[[Category:Genes]]<br />
[[Category:Japonica mRNA]]<br />
[[Category:Oryza Sativa Japonica Group]]<br />
[[Category:Japonica Genes]]<br />
[[Category:Japonica Chromosome 1]]<br />
[[Category:Chromosome 1]]</div>Mikahttps://ngdc.cncb.ac.cn/ricewiki/index.php?title=Os01g0182600&diff=183677Os01g01826002014-06-10T14:16:12Z<p>Mika: </p>
<hr />
<div>Please input one-sentence summary here.<br />
<br />
==Annotated Information==<br />
===Function===<br />
*''OsGI'',a ''GI'' ortholog, regulates flowering time in response to photoperiodic conditions. Mutations in this gene delay flowering in inductive SDs (short-days),whereas flowering is marginally induced under LDs(long-days).In comparison, over expression of ''OsGI'' in transgenic rice causes late flowering in both SDs and LDs.In addition, ''OsGI'' has function to regulate ''Hd1'', a gene that also shows circadian rhythm<ref name="ref1" />. ''Os-GI'' affects gene expression of approximately 75% of genes among 27,201 genes and is required for strong amplitudes and proper phase-setting of global gene expression under natural field conditions<ref name="ref2" />. <br />
*The cDNA of ''OsGI'' firstly found in ''se5'' mutant that is completely photoperiod-insensitive, the expression patterns of rice and Arabidopsis ''GI'' gene are both photoperiodically regulated and are very similar under LD and SD conditions(FIG1)<ref name="ref3" />. ''GI'' is required for phase-dependent ''CO''transcription through the interaction with FKF1, which leads to cycling DNA binding with one finger (CDF) degradation and consequent derepression of CO. In addition, it has been known that GI protein interacts with ZEITLUPE (ZTL) F-box protein to control the diurnal rhythm of TOC1 protein in regulation of the Arabidopsis circadian clock<ref name="ref4" />. <br />
*''Os-GI'' also has function to maintain bioactive GA level through the regulation of the GA-deactivating enzyme genes in rice, ''Os-GI'' represses major ''OsGA2ox'' genes in order to maintain a proper bioactive GA level, ''osgi-1'' plants showed a semi-dwarf phenotype with elevated ''OsGA2ox'' gene expression(FIG2) <ref name="ref4" />.<br />
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===Expression===<br />
*''GI'' encodes a nuclear protein that defines the proper amplitude and period length of circadian rhythms. ''GI'' expression is regulated by the circadian clock, with a peak in transcript levels at 8 to 10 h after dawn. The ''GI'' protein acts between the circadian oscillator and ''CO'' to promote flowering by increasing ''CO'' and ''FT'' mRNA abundance<ref name="ref1" />. The expression of the ''OsGI'' messenger RNA was circadian-controlled and that its temporal expression pattern was very similar to that of ''GI'' under both SD and LD conditions. Decreased ''OsGI'' expression led to early flowering under LD conditions and late flowering under SD conditions relative to the wild type, and that this early flowering phenotype under LD conditions was the opposite of that of the Arabidopsis ''gi'' mutants, confirming the opposite role of the ''GI'' gene in flowering under LD conditions in rice and Arabidopsis<ref name="ref5" />.<br />
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==Mutation==<br />
*The single ''osgi-1'' mutation extended the time to flowering only under SD conditions. Under LD conditions the single ''hd1'' mutation induced early flowering, but ''osgi-1'' plants exhibited no phenotypic change<ref name="ref4" /><br />
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===Evolution===<br />
* There is some evolutionary conservation between rice circadian clock–related genes and those associated with Arabidopsis circadian clocks, where the circadian clock consists of interlocked subloops, it is also possible that ''Os-GI'' is a member of a subloop in rice circadian clocks<ref name="ref4" />.But there is a divergence in the photoperiodic pathways of these two species arises from the molecular mechanisms controlling the expression of ''FT/Hd3a'' by ''CO/Hd1(Se1)''<ref name="ref5" />(figure 3)<br />
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==Labs working on this gene==<br />
Please input related labs here.<br />
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==Knowledge Extension==<br />
*The rice orthologues of the two key regulators of flowering time in Arabidopsis, ''CO'' and ''FT'', were recently shown to be important in the photoperiodic control of flowering in rice. The rice ''Hd1(Se1)''gene, which is an orthologue of the Arabidopsis ''CO'' gene, was required for the suppression of flowering under LD conditions and for the promotion of flowering under SD conditions. Furthermore, the rice ''Hd3a'' gene, an orthologue of the Arabidopsis ''FT'' gene, was shown to be an activator of flowering in rice. It was previously demonstrated that ''CO'' controls the flowering time by integrating signals from the circadian clock and light to regulate ''FT'' expression in Arabidopsis<ref name="ref5" />.Unlike Arabidopsis ''FT'', expression of ''Hd3a'' mRNA is more or less opposite to that of ''Hd1'' under inductive short-day conditions<ref name="ref6" />.<br />
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==References==<br />
Please input cited references here.<br />
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==Structured Information==<br />
{{JaponicaGene|<br />
GeneName = Os01g0182600|<br />
Description = GIGANTEA protein|<br />
Version = NM_001048755.1 GI:115434923 GeneID:4325329|<br />
Length = 9083 bp|<br />
Definition = Oryza sativa Japonica Group Os01g0182600, complete gene.|<br />
Source = Oryza sativa Japonica Group<br />
<br />
ORGANISM Oryza sativa Japonica Group<br />
Eukaryota; Viridiplantae; Streptophyta; Embryophyta; Tracheophyta;<br />
Spermatophyta; Magnoliophyta; Liliopsida; Poales; Poaceae; BEP<br />
clade; Ehrhartoideae; Oryzeae; Oryza.<br />
|<br />
Chromosome = [[:category:Japonica Chromosome 1|Chromosome 1]]|<br />
AP = Chromosome 1:4328362..4337444|<br />
CDS = 4328725..4329030,4329123..4329215,4329333..4329551,4329752..4329820,4329925..4331421<br>,4332220..4332429,4332563..4332713,4332791..4333031,4333898..4333956<br>,4334379..4334653,4334813..4334955,4335034..4335088,4335181..4335258<br>,4335351..4335437|<br />
GCID = <gbrowseImage1><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage1>|<br />
GSID = <gbrowseImage2><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage2>|<br />
CDNA = <cdnaseq>atgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactgcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcatagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcggcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctga</cdnaseq>|<br />
AA = <aaseq>MSASNEKWIDGLQFSSLFWPPPQDSQQKQAQILAYVEYFGQFTA DSEQFPEDIAQLIQSCYPSKEKRLVDEVLATFVLHHPEHGHAVVHPILSRIIDGTLSY DRNGFPFMSFISLFSHTSEKEYSEQWALACGEILRVLTHYNRPIFKVDHQHSEAECSS TSDQASSCESMEKRANGSPRNEPDRKPLRPLSPWITDILLAAPLGIRSDYFRWCGGVM GKYAAGGELKPPTTAYSRGSGKHPQLMPSTPRWAVANGAGVILSVCDEEVARYETANL TAAAVPALLLPPPTTPLDEHLVAGLPPLEPYARLFHRYYAIATPSATQRLLFGLLEAP PSWAPDALDAAVQLVELLRAAEDYDSGMRLPKNWMHLHFLRAIGTAMSMRAGIAADTS AALLFRILSQPTLLFPPLRHAEGVELHHEPLGGYVSSYKRQLEVPASEATIDATAQGI ASMLCAHGPDVEWRICTIWEAAYGLLPLSSSAVDLPEIVVAAPLQPPTLSWSLYLPLL KVFEYLPRGSPSEACLMRIFVATVEAILRRTFPSETSEQSRKPRSQSKNLAVAELRTM IHSLFVESCASMDLASRLLFVVLTVCVSHQALPGGSKRPTGSDNHSSEEVTNDSRLTN GRNRCKKRQGPVATFDSYVLAAVCALSCELQLFPFISKNGNHSNLKDSIKIVIPGKTT GISNELHNSISSAILHTRRILGILEALFSLKPSSVGTSWSYSSNEIVAAAMVAAHVSE LFRRSRPCLNALSALKQCKWDAEISTRASSLYHLIDLHGKTVTSIVNKAEPLEAHLTL TPVKKDEPPIEEKNINSSDGGALEKKDASRSHRKNGFARPLLKCAEDVILNGDVASTS GKAIASLQVEASDLANFLTMDRNGGYRGSQTLLRSVLSEKQELCFSVVSLLWQKLIAS PEMQMSAESTSAHQGWRKVVDALCDIVSASPTKASAAIVLQAEKDLQPWIARDDEQGQ KMWRVNQRIVKLIAELMRNHDSPEALVILASASDLLLRATDGMLVDGEACTLPQLELL EVTARAVHLIVEWGDSGVSVADGLSNLLKCRLSTTIRCLSHPSAHVRALSMSVLRDIL NSGQINSSKLIQGEHRNGIQSPTYQCLAASIINWQADVERCIEWEAHSRRATGLTLAF LTAAAKELGCPLTC</aaseq>|<br />
DNA = <dnaseqindica>8415..8720#8230..8322#7894..8112#7625..7693#6024..7520#5016..5225#4732..4882#4414..4654#3489..3547#2792..3066#2490..2632#2357..2411#2187..2264#2008..2094#ccgagccgtttctctccctctctctctcctcctccttcccctgcgcctcctccaccgcccgtgtcgccgccgcgcgcgtctatcccctcgccgccgccgccgtctacgccgcgtgaggtccggattggttgaaaggctgatcgaggtaatgcttttttatttttcccgtgcggttttagtttttctgtgctgtgcttggtggttttttctttaaaactatttttaggctggttttgatgatggcctttctaaggttgtcgtgtatggggggcgaggtggttcgggtggatttgtggtgttatgttgaggggatttttgtttggggggagagagttttggtggtagatgcggttgggattttgaagacgtgaaagggaaattagttttgaaatttgacggcgaggttgagctgattagtattgtaataattcgtggcatcggatagatcaacgcgagatggatgcttattactgcgaatttcaggaggtttttgtggtttttttaggcgagtagcgttgattctagctggattttgctgcttcgtggaggcgtttagatgtggtgcatacggctgtggatgtaatttgtgcggggataattactgaaaattaattcgcagtaagctagctctgctttaatattttttcccctgaaacagtaacaagggatcccttcctcatttcgttaacagtggaagtgaatacagacaccaaccgatgagcagagacacctgtaattaaacaccaaaagaagaacctgttttgttcctatatattttccatttctatgctttaggtaatatactttctggttcggtgtcatagtacccgaaatagaatctttgcttgccaaaaaaagaaagaataacaagatagaacaacatcattaaattctgtcttagacaatacatttataatcgaccaacaaaaacagaccgatgagtgacgtggaataacaaatgaagtataaatgtcacgaaaaacacatctattgattttgtgtgataagccataggagcctacgaggaaccaagtgaatggttctgtacaccacatccattagcattaccataaatttgcctatttctatgatgccttccttaacatatttatcgacatgtgagcttgcaccataactaaatgttatggtaaactttaccttataaatgattgcaacatgtaccttttcatttcgataattaaaaacgagaaagcttccacctttttcctagatatattttttcgtttgattatgaatttagttgtacttgacaataaaataatgcgcttgtaagattatgcaattttcgttgcattacctccctctcttttcatttgtcagtttttttctactccctccgtttcatattttaagtcgtttgacttttttcctagtcagactttgttaagtttgacaaagtttatagtaaaatttagcaatatctaaaatattaaattggtttcatcaaatctaacattgaatatattttgataatatgttttttttgtgttgaaaatacttctatatttttctataaacttagtcaaacttggaagaaatttcagtaagaaaaaagtcaaacggcttataatatgaaacggagggagtaaatttttttttgcattaattaaaattcatgtattttcctccttttttggtttttgttgtgcgatatattacagtgtagagactctgaagctcatatggttggcattggttgatatgataacgtgtcacgttgtgagatgcagacccccgttatgggggttgattgaatgaagtggatagcactcttataactttcttgctcgaagtttcttgataccaaggcaagtagcttataagaagatttggactttggtggcataaatggatgaactaactggcggcttcaacgtggtgttgattacaggtgcttttattgctaactggagcagtttgtgtagtttctttctttctctttgagaattctctcagtttcttgggcaggctcttgagctaccaagtatgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggtaacattattcctatttgattcttgttagaaattctatatgttgaacatacttgcctcaatgggaaacttggttgcgtctcctctctgaaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcaggtactgtatctctaagcatgtttagtatcttctgtgataaaatctttctccagtgtgttaacttttgcattctttttcttgcatgcttgcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattaggtaatttgatccagtgatcaatacatgttccacaaacaattgaagaaagagtttgacatatttttctcatatttccagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgaggtaaggcccctactctaatggttagttccaatttattggttcatttctctgtgtttgataagattttgcctctgctgtgcatttcaatattatatctaaactgtgctgcttatagcttatcaaactactcttgctcataaagatgcaacttttatgcagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgagtcataataaatgataacatattattgttctgtggactgaaaatctgaaatcctcctttcttgagaaatacaacctcaattttatggtgatattgacaagaaggaagagcgcatataatttaaagttaaaacacggaaaataatagaagtatgtgagcagaacgcaggagcaaggggtggaactcataacaagttcaaacttcatgcatgttcataattccccaatgcaagtatggcaagaaaaatagacaacttttaaaagtatgctaatggaactgcatgcattgtgtaaacttgcttttgtcaagcatgttggaacatactatatatttgctgctgatgatgaaaaatttgcttgagtactcttctattgctaatggacgcttaccattccctgttttgtcttcaaatgcatcaggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactggttggtaaaatcagcaatggaattttctatcgtgaattttttataattattgtaacattaattactcccatgtctttttattctgttgagataccatgtgacaatgtctacaattgcatattttgcattctttggtagatcagttcccttcttaagatccccaaagggaaactggagtatcaataacttactccctcaaataaaaaaaaagtgatactttaggcataaacacattcttcaaaacaaaactttgaatagcagtttcttttgtaatatattaataaagcatggaaaaattataatgtgaagaatgtacattcagggacaaatatactcatcatttttgcatggttgatcttaatgtccatagagagattgatcagaccaaaagtttatttttgacaggagtgagtatggattataataggagtgaatggcaatatgctgtgaaagtgagcattttatatttgtttactactagcatttggtgacaaagcaaatgatgtatattgtaaatcaaagtaaacaagtagaacatgtcctgtggcgacaagacataacctacaaattaaagttagcctgattgtttattttactgggagcacattttttctagattctagagtacatattattaaaaaaggtgtgatatgtttgcaaattactttttaatggaagtttgtaaattactttgaaacttgcattcagtttttttgcttcttaaaaaacatagtgcatatgggctttatgagctcatactatgcatgtttttctgcattggagtgggacctttatctttttcttgcaactcggatgctaatgcttattataaacacaagttactaacatgtgcttgggtcattcagcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcataggtaagctcttatgatcctgtccactgttgtaccatgtgtactttccaaatgattgctaacatgctatttttcgtcagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtac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/>
Link = [http://www.ncbi.nlm.nih.gov/nuccore/NM_001048755.1 RefSeq:Os01g0182600]|<br />
}}<br />
[[Category:Genes]]<br />
[[Category:Japonica mRNA]]<br />
[[Category:Oryza Sativa Japonica Group]]<br />
[[Category:Japonica Genes]]<br />
[[Category:Japonica Chromosome 1]]<br />
[[Category:Chromosome 1]]</div>Mikahttps://ngdc.cncb.ac.cn/ricewiki/index.php?title=Os01g0182600&diff=183675Os01g01826002014-06-10T14:13:42Z<p>Mika: </p>
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<div>Please input one-sentence summary here.<br />
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==Annotated Information==<br />
===Function===<br />
*''OsGI'',a ''GI'' ortholog, regulates flowering time in response to photoperiodic conditions. Mutations in this gene delay flowering in inductive SDs (short-days),whereas flowering is marginally induced under LDs(long-days).In comparison, over expression of ''OsGI'' in transgenic rice causes late flowering in both SDs and LDs.In addition, ''OsGI'' has function to regulate ''Hd1'', a gene that also shows circadian rhythm<ref name="ref1" />. ''Os-GI'' affects gene expression of approximately 75% of genes among 27,201 genes and is required for strong amplitudes and proper phase-setting of global gene expression under natural field conditions<ref name="ref2" />. <br />
*The cDNA of ''OsGI'' firstly found in ''se5'' mutant that is completely photoperiod-insensitive, the expression patterns of rice and Arabidopsis ''GI'' gene are both photoperiodically regulated and are very similar under LD and SD conditions(FIG1)<ref name="ref3" />. ''GI'' is required for phase-dependent ''CO''transcription through the interaction with FKF1, which leads to cycling DNA binding with one finger (CDF) degradation and consequent derepression of CO. In addition, it has been known that GI protein interacts with ZEITLUPE (ZTL) F-box protein to control the diurnal rhythm of TOC1 protein in regulation of the Arabidopsis circadian clock<ref name="ref4" />. <br />
*''Os-GI'' also has function to maintain bioactive GA level through the regulation of the GA-deactivating enzyme genes in rice, ''Os-GI'' represses major ''OsGA2ox'' genes in order to maintain a proper bioactive GA level, ''osgi-1'' plants showed a semi-dwarf phenotype with elevated ''OsGA2ox'' gene expression(FIG2) <ref name="ref4" />.<br />
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===Expression===<br />
*''GI'' encodes a nuclear protein that defines the proper amplitude and period length of circadian rhythms. ''GI'' expression is regulated by the circadian clock, with a peak in transcript levels at 8 to 10 h after dawn. The ''GI'' protein acts between the circadian oscillator and ''CO'' to promote flowering by increasing ''CO'' and ''FT'' mRNA abundance<ref name="ref1" />. The expression of the ''OsGI'' messenger RNA was circadian-controlled and that its temporal expression pattern was very similar to that of ''GI'' under both SD and LD conditions. Decreased ''OsGI'' expression led to early flowering under LD conditions and late flowering under SD conditions relative to the wild type, and that this early flowering phenotype under LD conditions was the opposite of that of the Arabidopsis ''gi'' mutants, confirming the opposite role of the ''GI'' gene in flowering under LD conditions in rice and Arabidopsis<ref name="ref5" />.<br />
==Mutation==<br />
*The single ''osgi-1'' mutation extended the time to flowering only under SD conditions. Under LD conditions the single ''hd1'' mutation induced early flowering, but ''osgi-1'' plants exhibited no phenotypic change<ref name="ref4" /><br />
===Evolution===<br />
* There is some evolutionary conservation between rice circadian clock–related genes and those associated with Arabidopsis circadian clocks, where the circadian clock consists of interlocked subloops, it is also possible that ''Os-GI'' is a member of a subloop in rice circadian clocks<ref name="ref4" />.But there is a divergence in the photoperiodic pathways of these two species arises from the molecular mechanisms controlling the expression of ''FT/Hd3a'' by ''CO/Hd1(Se1)''<ref name="ref5" />(figure 3)<br />
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==Labs working on this gene==<br />
Please input related labs here.<br />
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==Knowledge Extension==<br />
*The rice orthologues of the two key regulators of flowering time in Arabidopsis, ''CO'' and ''FT'', were recently shown to be important in the photoperiodic control of flowering in rice. The rice ''Hd1(Se1)''gene, which is an orthologue of the Arabidopsis ''CO'' gene, was required for the suppression of flowering under LD conditions and for the promotion of flowering under SD conditions. Furthermore, the rice ''Hd3a'' gene, an orthologue of the Arabidopsis ''FT'' gene, was shown to be an activator of flowering in rice. It was previously demonstrated that ''CO'' controls the flowering time by integrating signals from the circadian clock and light to regulate ''FT'' expression in Arabidopsis<ref name="ref5" />.Unlike Arabidopsis ''FT'', expression of ''Hd3a'' mRNA is more or less opposite to that of ''Hd1'' under inductive short-day conditions<ref name="ref6" />.<br />
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==References==<br />
Please input cited references here.<br />
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==Structured Information==<br />
{{JaponicaGene|<br />
GeneName = Os01g0182600|<br />
Description = GIGANTEA protein|<br />
Version = NM_001048755.1 GI:115434923 GeneID:4325329|<br />
Length = 9083 bp|<br />
Definition = Oryza sativa Japonica Group Os01g0182600, complete gene.|<br />
Source = Oryza sativa Japonica Group<br />
<br />
ORGANISM Oryza sativa Japonica Group<br />
Eukaryota; Viridiplantae; Streptophyta; Embryophyta; Tracheophyta;<br />
Spermatophyta; Magnoliophyta; Liliopsida; Poales; Poaceae; BEP<br />
clade; Ehrhartoideae; Oryzeae; Oryza.<br />
|<br />
Chromosome = [[:category:Japonica Chromosome 1|Chromosome 1]]|<br />
AP = Chromosome 1:4328362..4337444|<br />
CDS = 4328725..4329030,4329123..4329215,4329333..4329551,4329752..4329820,4329925..4331421<br>,4332220..4332429,4332563..4332713,4332791..4333031,4333898..4333956<br>,4334379..4334653,4334813..4334955,4335034..4335088,4335181..4335258<br>,4335351..4335437|<br />
GCID = <gbrowseImage1><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage1>|<br />
GSID = <gbrowseImage2><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage2>|<br />
CDNA = <cdnaseq>atgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactgcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcatagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcggcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctga</cdnaseq>|<br />
AA = <aaseq>MSASNEKWIDGLQFSSLFWPPPQDSQQKQAQILAYVEYFGQFTA DSEQFPEDIAQLIQSCYPSKEKRLVDEVLATFVLHHPEHGHAVVHPILSRIIDGTLSY DRNGFPFMSFISLFSHTSEKEYSEQWALACGEILRVLTHYNRPIFKVDHQHSEAECSS TSDQASSCESMEKRANGSPRNEPDRKPLRPLSPWITDILLAAPLGIRSDYFRWCGGVM GKYAAGGELKPPTTAYSRGSGKHPQLMPSTPRWAVANGAGVILSVCDEEVARYETANL TAAAVPALLLPPPTTPLDEHLVAGLPPLEPYARLFHRYYAIATPSATQRLLFGLLEAP PSWAPDALDAAVQLVELLRAAEDYDSGMRLPKNWMHLHFLRAIGTAMSMRAGIAADTS AALLFRILSQPTLLFPPLRHAEGVELHHEPLGGYVSSYKRQLEVPASEATIDATAQGI ASMLCAHGPDVEWRICTIWEAAYGLLPLSSSAVDLPEIVVAAPLQPPTLSWSLYLPLL KVFEYLPRGSPSEACLMRIFVATVEAILRRTFPSETSEQSRKPRSQSKNLAVAELRTM IHSLFVESCASMDLASRLLFVVLTVCVSHQALPGGSKRPTGSDNHSSEEVTNDSRLTN GRNRCKKRQGPVATFDSYVLAAVCALSCELQLFPFISKNGNHSNLKDSIKIVIPGKTT GISNELHNSISSAILHTRRILGILEALFSLKPSSVGTSWSYSSNEIVAAAMVAAHVSE LFRRSRPCLNALSALKQCKWDAEISTRASSLYHLIDLHGKTVTSIVNKAEPLEAHLTL TPVKKDEPPIEEKNINSSDGGALEKKDASRSHRKNGFARPLLKCAEDVILNGDVASTS GKAIASLQVEASDLANFLTMDRNGGYRGSQTLLRSVLSEKQELCFSVVSLLWQKLIAS PEMQMSAESTSAHQGWRKVVDALCDIVSASPTKASAAIVLQAEKDLQPWIARDDEQGQ KMWRVNQRIVKLIAELMRNHDSPEALVILASASDLLLRATDGMLVDGEACTLPQLELL EVTARAVHLIVEWGDSGVSVADGLSNLLKCRLSTTIRCLSHPSAHVRALSMSVLRDIL NSGQINSSKLIQGEHRNGIQSPTYQCLAASIINWQADVERCIEWEAHSRRATGLTLAF LTAAAKELGCPLTC</aaseq>|<br />
DNA = <dnaseqindica>8415..8720#8230..8322#7894..8112#7625..7693#6024..7520#5016..5225#4732..4882#4414..4654#3489..3547#2792..3066#2490..2632#2357..2411#2187..2264#2008..2094#ccgagccgtttctctccctctctctctcctcctccttcccctgcgcctcctccaccgcccgtgtcgccgccgcgcgcgtctatcccctcgccgccgccgccgtctacgccgcgtgaggtccggattggttgaaaggctgatcgaggtaatgcttttttatttttcccgtgcggttttagtttttctgtgctgtgcttggtggttttttctttaaaactatttttaggctggttttgatgatggcctttctaaggttgtcgtgtatggggggcgaggtggttcgggtggatttgtggtgttatgttgaggggatttttgtttggggggagagagttttggtggtagatgcggttgggattttgaagacgtgaaagggaaattagttttgaaatttgacggcgaggttgagctgattagtattgtaataattcgtggcatcggatagatcaacgcgagatggatgcttattactgcgaatttcaggaggtttttgtggtttttttaggcgagtagcgttgattctagctggattttgctgcttcgtggaggcgtttagatgtggtgcatacggctgtggatgtaatttgtgcggggataattactgaaaattaattcgcagtaagctagctctgctttaatattttttcccctgaaacagtaacaagggatcccttcctcatttcgttaacagtggaagtgaatacagacaccaaccgatgagcagagacacctgtaattaaacaccaaaagaagaacctgttttgttcctatatattttccatttctatgctttaggtaatatactttctggttcggtgtcatagtacccgaaatagaatctttgcttgccaaaaaaagaaagaataacaagatagaacaacatcattaaattctgtcttagacaatacatttataatcgaccaacaaaaacagaccgatgagtgacgtggaataacaaatgaagtataaatgtcacgaaaaacacatctattgattttgtgtgataagccataggagcctacgaggaaccaagtgaatggttctgtacaccacatccattagcattaccataaatttgcctatttctatgatgccttccttaacatatttatcgacatgtgagcttgcaccataactaaatgttatggtaaactttaccttataaatgattgcaacatgtaccttttcatttcgataattaaaaacgagaaagcttccacctttttcctagatatattttttcgtttgattatgaatttagttgtacttgacaataaaataatgcgcttgtaagattatgcaattttcgttgcattacctccctctcttttcatttgtcagtttttttctactccctccgtttcatattttaagtcgtttgacttttttcctagtcagactttgttaagtttgacaaagtttatagtaaaatttagcaatatctaaaatattaaattggtttcatcaaatctaacattgaatatattttgataatatgttttttttgtgttgaaaatacttctatatttttctataaacttagtcaaacttggaagaaatttcagtaagaaaaaagtcaaacggcttataatatgaaacggagggagtaaatttttttttgcattaattaaaattcatgtattttcctccttttttggtttttgttgtgcgatatattacagtgtagagactctgaagctcatatggttggcattggttgatatgataacgtgtcacgttgtgagatgcagacccccgttatgggggttgattgaatgaagtggatagcactcttataactttcttgctcgaagtttcttgataccaaggcaagtagcttataagaagatttggactttggtggcataaatggatgaactaactggcggcttcaacgtggtgttgattacaggtgcttttattgctaactggagcagtttgtgtagtttctttctttctctttgagaattctctcagtttcttgggcaggctcttgagctaccaagtatgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggtaacattattcctatttgattcttgttagaaattctatatgttgaacatacttgcctcaatgggaaacttggttgcgtctcctctctgaaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcaggtactgtatctctaagcatgtttagtatcttctgtgataaaatctttctccagtgtgttaacttttgcattctttttcttgcatgcttgcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattaggtaatttgatccagtgatcaatacatgttccacaaacaattgaagaaagagtttgacatatttttctcatatttccagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgaggtaaggcccctactctaatggttagttccaatttattggttcatttctctgtgtttgataagattttgcctctgctgtgcatttcaatattatatctaaactgtgctgcttatagcttatcaaactactcttgctcataaagatgcaacttttatgcagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgagtcataataaatgataacatattattgttctgtggactgaaaatctgaaatcctcctttcttgagaaatacaacctcaattttatggtgatattgacaagaaggaagagcgcatataatttaaagttaaaacacggaaaataatagaagtatgtgagcagaacgcaggagcaaggggtggaactcataacaagttcaaacttcatgcatgttcataattccccaatgcaagtatggcaagaaaaatagacaacttttaaaagtatgctaatggaactgcatgcattgtgtaaacttgcttttgtcaagcatgttggaacatactatatatttgctgctgatgatgaaaaatttgcttgagtactcttctattgctaatggacgcttaccattccctgttttgtcttcaaatgcatcaggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactggttggtaaaatcagcaatggaattttctatcgtgaattttttataattattgtaacattaattactcccatgtctttttattctgttgagataccatgtgacaatgtctacaattgcatattttgcattctttggtagatcagttcccttcttaagatccccaaagggaaactggagtatcaataacttactccctcaaataaaaaaaaagtgatactttaggcataaacacattcttcaaaacaaaactttgaatagcagtttcttttgtaatatattaataaagcatggaaaaattataatgtgaagaatgtacattcagggacaaatatactcatcatttttgcatggttgatcttaatgtccatagagagattgatcagaccaaaagtttatttttgacaggagtgagtatggattataataggagtgaatggcaatatgctgtgaaagtgagcattttatatttgtttactactagcatttggtgacaaagcaaatgatgtatattgtaaatcaaagtaaacaagtagaacatgtcctgtggcgacaagacataacctacaaattaaagttagcctgattgtttattttactgggagcacattttttctagattctagagtacatattattaaaaaaggtgtgatatgtttgcaaattactttttaatggaagtttgtaaattactttgaaacttgcattcagtttttttgcttcttaaaaaacatagtgcatatgggctttatgagctcatactatgcatgtttttctgcattggagtgggacctttatctttttcttgcaactcggatgctaatgcttattataaacacaagttactaacatgtgcttgggtcattcagcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcataggtaagctcttatgatcctgtccactgttgtaccatgtgtactttccaaatgattgctaacatgctatttttcgtcagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcgggtatatcatacatctctctggctgttgtttcagtgctatcaatcaagatgacacatcatgcaatattatcataacagcacattatgtgttttgaatattttattgattgtcatagctttgttttatgttttagcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcaggtatgcagtgtttcctgcacctacttgctacctattctgtatttgatgtattcaatttcctttttatcagttttctcgaaatatatgctaataggcttgtcactaaacggctacattatccagactctgttctgaatttttttttcattaagtactgttagctgccagattacatcatcatgactaaagtagtttagctacaatattgcaaagctagatttgtctagcctctattgttttgaatagaaaaggcaggtatgcagtgtttcctgcacctacttgctacctattctgtatttgatgtattcaatttcctttttatcagttttctcgaaatatatgctaataggcttgtcactaaacggctacattatccagactctgttctgaatttttttttcattaagtactgttagctgccagattacatcatcatgactaaagtagtttagctacaatattgcaaagctagatttgtctagcctctattgttttgaatagaaaattaaatcttttggtcatttttttgtaaccttttaggagtctctgtatgcttgtcgtcaacatgctatgataagtttatatcagttcttttggagctatgatgtattcttgtatattttatgactgctaggatattatgttaaaactgttttaaaggtatcatcaaacaagttaatatagatattgctacctaccaaagttggaatttgattatgtcgtaaaatatatgcaccctacatgttttagcttcatattgacttatattttcatatgtgctccttcaactgcacaaagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggtatggtcattcctgcttataagataccagaataatgctatgaagaccgtccttggttgtcagcattcttatacacaatgataatctttgtttttcatgggtaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggtcaagatattgttattgttatatttacatacttgttatcatatttaaaagcttcatattgtatgtacctttgaaaaaaaaagttccatactgtagcttcttttgttctcatttcttggtttcaaagcatcagcatcattaccgatataaaatatacccatttcctcagtcatcttccccgactcttgtatctttcacaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagaggtaacaccttcattcatgaggtccttaggtgcaaatcccagtgaagaataatcagttctactttgtatggttcatctcgcttactgaccggttcatctcgggatcaattacgagcagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaaggcatgtcctcttctctgatctcactcacgttcatgccaaacttactcatcatcatccatgatccatcatcctgactgaatcttgatatgcagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctgacaaggccacaccatgactgtcactgcaagcagctgctgtaggatcagcgagtaggaaaggatgacttgccggccagtttcctgagatgtgatttaagactgatattagctgatgttcccaagcatgatacggggcttgctcccaagatgtgatttttactttacttttggttaatgtgctaccgctgacattagatgttcaagcatatgaaaactgcttgtgctgtaacttgtatgtctgtaacgtaaaaatctttccatatggtaaccggggaagcagaggtatatccttttctgctgttgtatctatttccagtttcctgctgaacttgcaaatgaatattcagcaatttccgttttgctc</dnaseqindica>|<br />
Link = [http://www.ncbi.nlm.nih.gov/nuccore/NM_001048755.1 RefSeq:Os01g0182600]|<br />
}}<br />
[[Category:Genes]]<br />
[[Category:Japonica mRNA]]<br />
[[Category:Oryza Sativa Japonica Group]]<br />
[[Category:Japonica Genes]]<br />
[[Category:Japonica Chromosome 1]]<br />
[[Category:Chromosome 1]]</div>Mikahttps://ngdc.cncb.ac.cn/ricewiki/index.php?title=Os01g0182600&diff=183674Os01g01826002014-06-10T14:10:59Z<p>Mika: /* Knowledge Extension */</p>
<hr />
<div>Please input one-sentence summary here.<br />
<br />
==Annotated Information==<br />
===Function===<br />
*''OsGI'',a ''GI'' ortholog, regulates flowering time in response to photoperiodic conditions. Mutations in this gene delay flowering in inductive SDs (short-days),whereas flowering is marginally induced under LDs(long-days).In comparison, over expression of ''OsGI'' in transgenic rice causes late flowering in both SDs and LDs.In addition, ''OsGI'' has function to regulate ''Hd1'', a gene that also shows circadian rhythm<ref name="ref1" />. ''Os-GI'' affects gene expression of approximately 75% of genes among 27,201 genes and is required for strong amplitudes and proper phase-setting of global gene expression under natural field conditions<ref name="ref2" />. <br />
*The cDNA of ''OsGI'' firstly found in ''se5'' mutant that is completely photoperiod-insensitive, the expression patterns of rice and Arabidopsis ''GI'' gene are both photoperiodically regulated and are very similar under LD and SD conditions(FIG1)<ref name="ref3" />. ''GI'' is required for phase-dependent ''CO''transcription through the interaction with FKF1, which leads to cycling DNA binding with one finger (CDF) degradation and consequent derepression of CO. In addition, it has been known that GI protein interacts with ZEITLUPE (ZTL) F-box protein to control the diurnal rhythm of TOC1 protein in regulation of the Arabidopsis circadian clock<ref name="ref4" />. <br />
*''Os-GI'' also has function to maintain bioactive GA level through the regulation of the GA-deactivating enzyme genes in rice, ''Os-GI'' represses major ''OsGA2ox'' genes in order to maintain a proper bioactive GA level, ''osgi-1'' plants showed a semi-dwarf phenotype with elevated ''OsGA2ox'' gene expression(FIG2) <ref name="ref4" />.<br />
<br />
===Expression===<br />
*''GI'' encodes a nuclear protein that defines the proper amplitude and period length of circadian rhythms. ''GI'' expression is regulated by the circadian clock, with a peak in transcript levels at 8 to 10 h after dawn. The ''GI'' protein acts between the circadian oscillator and ''CO'' to promote flowering by increasing ''CO'' and ''FT'' mRNA abundance<ref name="ref1" />. The expression of the ''OsGI'' messenger RNA was circadian-controlled and that its temporal expression pattern was very similar to that of ''GI'' under both SD and LD conditions. Decreased ''OsGI'' expression led to early flowering under LD conditions and late flowering under SD conditions relative to the wild type, and that this early flowering phenotype under LD conditions was the opposite of that of the Arabidopsis ''gi'' mutants, confirming the opposite role of the ''GI'' gene in flowering under LD conditions in rice and Arabidopsis<ref name="ref5" />.<br />
<br />
===Evolution===<br />
* There is some evolutionary conservation between rice circadian clock–related genes and those associated with Arabidopsis circadian clocks, where the circadian clock consists of interlocked subloops, it is also possible that ''Os-GI'' is a member of a subloop in rice circadian clocks<ref name="ref4" />.But there is a divergence in the photoperiodic pathways of these two species arises from the molecular mechanisms controlling the expression of ''FT/Hd3a'' by ''CO/Hd1(Se1)''<ref name="ref5" />(figure 3)<br />
<br />
==Labs working on this gene==<br />
Please input related labs here.<br />
<br />
==Knowledge Extension==<br />
*The rice orthologues of the two key regulators of flowering time in Arabidopsis, ''CO'' and ''FT'', were recently shown to be important in the photoperiodic control of flowering in rice. The rice ''Hd1(Se1)''gene, which is an orthologue of the Arabidopsis ''CO'' gene, was required for the suppression of flowering under LD conditions and for the promotion of flowering under SD conditions. Furthermore, the rice ''Hd3a'' gene, an orthologue of the Arabidopsis ''FT'' gene, was shown to be an activator of flowering in rice. It was previously demonstrated that ''CO'' controls the flowering time by integrating signals from the circadian clock and light to regulate ''FT'' expression in Arabidopsis<ref name="ref5" />.Unlike Arabidopsis ''FT'', expression of ''Hd3a'' mRNA is more or less opposite to that of ''Hd1'' under inductive short-day conditions<ref name="ref6" />.<br />
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==References==<br />
Please input cited references here.<br />
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==Structured Information==<br />
{{JaponicaGene|<br />
GeneName = Os01g0182600|<br />
Description = GIGANTEA protein|<br />
Version = NM_001048755.1 GI:115434923 GeneID:4325329|<br />
Length = 9083 bp|<br />
Definition = Oryza sativa Japonica Group Os01g0182600, complete gene.|<br />
Source = Oryza sativa Japonica Group<br />
<br />
ORGANISM Oryza sativa Japonica Group<br />
Eukaryota; Viridiplantae; Streptophyta; Embryophyta; Tracheophyta;<br />
Spermatophyta; Magnoliophyta; Liliopsida; Poales; Poaceae; BEP<br />
clade; Ehrhartoideae; Oryzeae; Oryza.<br />
|<br />
Chromosome = [[:category:Japonica Chromosome 1|Chromosome 1]]|<br />
AP = Chromosome 1:4328362..4337444|<br />
CDS = 4328725..4329030,4329123..4329215,4329333..4329551,4329752..4329820,4329925..4331421<br>,4332220..4332429,4332563..4332713,4332791..4333031,4333898..4333956<br>,4334379..4334653,4334813..4334955,4335034..4335088,4335181..4335258<br>,4335351..4335437|<br />
GCID = <gbrowseImage1><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage1>|<br />
GSID = <gbrowseImage2><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage2>|<br />
CDNA = <cdnaseq>atgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactgcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcatagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcggcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctga</cdnaseq>|<br />
AA = <aaseq>MSASNEKWIDGLQFSSLFWPPPQDSQQKQAQILAYVEYFGQFTA DSEQFPEDIAQLIQSCYPSKEKRLVDEVLATFVLHHPEHGHAVVHPILSRIIDGTLSY DRNGFPFMSFISLFSHTSEKEYSEQWALACGEILRVLTHYNRPIFKVDHQHSEAECSS TSDQASSCESMEKRANGSPRNEPDRKPLRPLSPWITDILLAAPLGIRSDYFRWCGGVM GKYAAGGELKPPTTAYSRGSGKHPQLMPSTPRWAVANGAGVILSVCDEEVARYETANL TAAAVPALLLPPPTTPLDEHLVAGLPPLEPYARLFHRYYAIATPSATQRLLFGLLEAP PSWAPDALDAAVQLVELLRAAEDYDSGMRLPKNWMHLHFLRAIGTAMSMRAGIAADTS AALLFRILSQPTLLFPPLRHAEGVELHHEPLGGYVSSYKRQLEVPASEATIDATAQGI ASMLCAHGPDVEWRICTIWEAAYGLLPLSSSAVDLPEIVVAAPLQPPTLSWSLYLPLL KVFEYLPRGSPSEACLMRIFVATVEAILRRTFPSETSEQSRKPRSQSKNLAVAELRTM IHSLFVESCASMDLASRLLFVVLTVCVSHQALPGGSKRPTGSDNHSSEEVTNDSRLTN GRNRCKKRQGPVATFDSYVLAAVCALSCELQLFPFISKNGNHSNLKDSIKIVIPGKTT GISNELHNSISSAILHTRRILGILEALFSLKPSSVGTSWSYSSNEIVAAAMVAAHVSE LFRRSRPCLNALSALKQCKWDAEISTRASSLYHLIDLHGKTVTSIVNKAEPLEAHLTL TPVKKDEPPIEEKNINSSDGGALEKKDASRSHRKNGFARPLLKCAEDVILNGDVASTS GKAIASLQVEASDLANFLTMDRNGGYRGSQTLLRSVLSEKQELCFSVVSLLWQKLIAS PEMQMSAESTSAHQGWRKVVDALCDIVSASPTKASAAIVLQAEKDLQPWIARDDEQGQ KMWRVNQRIVKLIAELMRNHDSPEALVILASASDLLLRATDGMLVDGEACTLPQLELL EVTARAVHLIVEWGDSGVSVADGLSNLLKCRLSTTIRCLSHPSAHVRALSMSVLRDIL NSGQINSSKLIQGEHRNGIQSPTYQCLAASIINWQADVERCIEWEAHSRRATGLTLAF LTAAAKELGCPLTC</aaseq>|<br />
DNA = <dnaseqindica>8415..8720#8230..8322#7894..8112#7625..7693#6024..7520#5016..5225#4732..4882#4414..4654#3489..3547#2792..3066#2490..2632#2357..2411#2187..2264#2008..2094#ccgagccgtttctctccctctctctctcctcctccttcccctgcgcctcctccaccgcccgtgtcgccgccgcgcgcgtctatcccctcgccgccgccgccgtctacgccgcgtgaggtccggattggttgaaaggctgatcgaggtaatgcttttttatttttcccgtgcggttttagtttttctgtgctgtgcttggtggttttttctttaaaactatttttaggctggttttgatgatggcctttctaaggttgtcgtgtatggggggcgaggtggttcgggtggatttgtggtgttatgttgaggggatttttgtttggggggagagagttttggtggtagatgcggttgggattttgaagacgtgaaagggaaattagttttgaaatttgacggcgaggttgagctgattagtattgtaataattcgtggcatcggatagatcaacgcgagatggatgcttattactgcgaatttcaggaggtttttgtggtttttttaggcgagtagcgttgattctagctggattttgctgcttcgtggaggcgtttagatgtggtgcatacggctgtggatgtaatttgtgcggggataattactgaaaattaattcgcagtaagctagctctgctttaatattttttcccctgaaacagtaacaagggatcccttcctcatttcgttaacagtggaagtgaatacagacaccaaccgatgagcagagacacctgtaattaaacaccaaaagaagaacctgttttgttcctatatattttccatttctatgctttaggtaatatactttctggttcggtgtcatagtacccgaaatagaatctttgcttgccaaaaaaagaaagaataacaagatagaacaacatcattaaattctgtcttagacaatacatttataatcgaccaacaaaaacagaccgatgagtgacgtggaataacaaatgaagtataaatgtcacgaaaaacacatctattgattttgtgtgataagccataggagcctacgaggaaccaagtgaatggttctgtacaccacatccattagcattaccataaatttgcctatttctatgatgccttccttaacatatttatcgacatgtgagcttgcaccataactaaatgttatggtaaactttaccttataaatgattgcaacatgtaccttttcatttcgataattaaaaacgagaaagcttccacctttttcctagatatattttttcgtttgattatgaatttagttgtacttgacaataaaataatgcgcttgtaagattatgcaattttcgttgcattacctccctctcttttcatttgtcagtttttttctactccctccgtttcatattttaagtcgtttgacttttttcctagtcagactttgttaagtttgacaaagtttatagtaaaatttagcaatatctaaaatattaaattggtttcatcaaatctaacattgaatatattttgataatatgttttttttgtgttgaaaatacttctatatttttctataaacttagtcaaacttggaagaaatttcagtaagaaaaaagtcaaacggcttataatatgaaacggagggagtaaatttttttttgcattaattaaaattcatgtattttcctccttttttggtttttgttgtgcgatatattacagtgtagagactctgaagctcatatggttggcattggttgatatgataacgtgtcacgttgtgagatgcagacccccgttatgggggttgattgaatgaagtggatagcactcttataactttcttgctcgaagtttcttgataccaaggcaagtagcttataagaagatttggactttggtggcataaatggatgaactaactggcggcttcaacgtggtgttgattacaggtgcttttattgctaactggagcagtttgtgtagtttctttctttctctttgagaattctctcagtttcttgggcaggctcttgagctaccaagtatgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggtaacattattcctatttgattcttgttagaaattctatatgttgaacatacttgcctcaatgggaaacttggttgcgtctcctctctgaaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcaggtactgtatctctaagcatgtttagtatcttctgtgataaaatctttctccagtgtgttaacttttgcattctttttcttgcatgcttgcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattaggtaatttgatccagtgatcaatacatgttccacaaacaattgaagaaagagtttgacatatttttctcatatttccagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgaggtaaggcccctactctaatggttagttccaatttattggttcatttctctgtgtttgataagattttgcctctgctgtgcatttcaatattatatctaaactgtgctgcttatagcttatcaaactactcttgctcataaagatgcaacttttatgcagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgagtcataataaatgataacatattattgttctgtggactgaaaatctgaaatcctcctttcttgagaaatacaacctcaattttatggtgatattgacaagaaggaagagcgcatataatttaaagttaaaacacggaaaataatagaagtatgtgagcagaacgcaggagcaaggggtggaactcataacaagttcaaacttcatgcatgttcataattccccaatgcaagtatggcaagaaaaatagacaacttttaaaagtatgctaatggaactgcatgcattgtgtaaacttgcttttgtcaagcatgttggaacatactatatatttgctgctgatgatgaaaaatttgcttgagtactcttctattgctaatggacgcttaccattccctgttttgtcttcaaatgcatcaggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactggttggtaaaatcagcaatggaattttctatcgtgaattttttataattattgtaacattaattactcccatgtctttttattctgttgagataccatgtgacaatgtctacaattgcatattttgcattctttggtagatcagttcccttcttaagatccccaaagggaaactggagtatcaataacttactccctcaaataaaaaaaaagtgatactttaggcataaacacattcttcaaaacaaaactttgaatagcagtttcttttgtaatatattaataaagcatggaaaaattataatgtgaagaatgtacattcagggacaaatatactcatcatttttgcatggttgatcttaatgtccatagagagattgatcagaccaaaagtttatttttgacaggagtgagtatggattataataggagtgaatggcaatatgctgtgaaagtgagcattttatatttgtttactactagcatttggtgacaaagcaaatgatgtatattgtaaatcaaagtaaacaagtagaacatgtcctgtggcgacaagacataacctacaaattaaagttagcctgattgtttattttactgggagcacattttttctagattctagagtacatattattaaaaaaggtgtgatatgtttgcaaattactttttaatggaagtttgtaaattactttgaaacttgcattcagtttttttgcttcttaaaaaacatagtgcatatgggctttatgagctcatactatgcatgtttttctgcattggagtgggacctttatctttttcttgcaactcggatgctaatgcttattataaacacaagttactaacatgtgcttgggtcattcagcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcataggtaagctcttatgatcctgtccactgttgtaccatgtgtactttccaaatgattgctaacatgctatttttcgtcagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcgggtatatcatacatctctctggctgttgtttcagtgctatcaatcaagatgacacatcatgcaatattatcataacagcacattatgtgttttgaatattttattgattgtcatagctttgttttatgttttagcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcaggtatgcagtgtttcctgcacctacttgctacctattctgtatttgatgtattcaatttcctttttatcagttttctcgaaatatatgctaataggcttgtcactaaacggctacattatccagactctgttctgaatttttttttcattaagtactgttagctgccagattacatcatcatgactaaagtagtttagctacaatattgcaaagctagatttgtctagcctctattgttttgaatagaaaaggcaggtatgcagtgtttcctgcacctacttgctacctattctgtatttgatgtattcaatttcctttttatcagttttctcgaaatatatgctaataggcttgtcactaaacggctacattatccagactctgttctgaatttttttttcattaagtactgttagctgccagattacatcatcatgactaaagtagtttagctacaatattgcaaagctagatttgtctagcctctattgttttgaatagaaaattaaatcttttggtcatttttttgtaaccttttaggagtctctgtatgcttgtcgtcaacatgctatgataagtttatatcagttcttttggagctatgatgtattcttgtatattttatgactgctaggatattatgttaaaactgttttaaaggtatcatcaaacaagttaatatagatattgctacctaccaaagttggaatttgattatgtcgtaaaatatatgcaccctacatgttttagcttcatattgacttatattttcatatgtgctccttcaactgcacaaagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggtatggtcattcctgcttataagataccagaataatgctatgaagaccgtccttggttgtcagcattcttatacacaatgataatctttgtttttcatgggtaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggtcaagatattgttattgttatatttacatacttgttatcatatttaaaagcttcatattgtatgtacctttgaaaaaaaaagttccatactgtagcttcttttgttctcatttcttggtttcaaagcatcagcatcattaccgatataaaatatacccatttcctcagtcatcttccccgactcttgtatctttcacaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagaggtaacaccttcattcatgaggtccttaggtgcaaatcccagtgaagaataatcagttctactttgtatggttcatctcgcttactgaccggttcatctcgggatcaattacgagcagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaaggcatgtcctcttctctgatctcactcacgttcatgccaaacttactcatcatcatccatgatccatcatcctgactgaatcttgatatgcagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctgacaaggccacaccatgactgtcactgcaagcagctgctgtaggatcagcgagtaggaaaggatgacttgccggccagtttcctgagatgtgatttaagactgatattagctgatgttcccaagcatgatacggggcttgctcccaagatgtgatttttactttacttttggttaatgtgctaccgctgacattagatgttcaagcatatgaaaactgcttgtgctgtaacttgtatgtctgtaacgtaaaaatctttccatatggtaaccggggaagcagaggtatatccttttctgctgttgtatctatttccagtttcctgctgaacttgcaaatgaatattcagcaatttccgttttgctc</dnaseqindica>|<br />
Link = [http://www.ncbi.nlm.nih.gov/nuccore/NM_001048755.1 RefSeq:Os01g0182600]|<br />
}}<br />
[[Category:Genes]]<br />
[[Category:Japonica mRNA]]<br />
[[Category:Oryza Sativa Japonica Group]]<br />
[[Category:Japonica Genes]]<br />
[[Category:Japonica Chromosome 1]]<br />
[[Category:Chromosome 1]]</div>Mikahttps://ngdc.cncb.ac.cn/ricewiki/index.php?title=Os01g0182600&diff=183673Os01g01826002014-06-10T14:07:12Z<p>Mika: /* Expression */</p>
<hr />
<div>Please input one-sentence summary here.<br />
<br />
==Annotated Information==<br />
===Function===<br />
*''OsGI'',a ''GI'' ortholog, regulates flowering time in response to photoperiodic conditions. Mutations in this gene delay flowering in inductive SDs (short-days),whereas flowering is marginally induced under LDs(long-days).In comparison, over expression of ''OsGI'' in transgenic rice causes late flowering in both SDs and LDs.In addition, ''OsGI'' has function to regulate ''Hd1'', a gene that also shows circadian rhythm<ref name="ref1" />. ''Os-GI'' affects gene expression of approximately 75% of genes among 27,201 genes and is required for strong amplitudes and proper phase-setting of global gene expression under natural field conditions<ref name="ref2" />. <br />
*The cDNA of ''OsGI'' firstly found in ''se5'' mutant that is completely photoperiod-insensitive, the expression patterns of rice and Arabidopsis ''GI'' gene are both photoperiodically regulated and are very similar under LD and SD conditions(FIG1)<ref name="ref3" />. ''GI'' is required for phase-dependent ''CO''transcription through the interaction with FKF1, which leads to cycling DNA binding with one finger (CDF) degradation and consequent derepression of CO. In addition, it has been known that GI protein interacts with ZEITLUPE (ZTL) F-box protein to control the diurnal rhythm of TOC1 protein in regulation of the Arabidopsis circadian clock<ref name="ref4" />. <br />
*''Os-GI'' also has function to maintain bioactive GA level through the regulation of the GA-deactivating enzyme genes in rice, ''Os-GI'' represses major ''OsGA2ox'' genes in order to maintain a proper bioactive GA level, ''osgi-1'' plants showed a semi-dwarf phenotype with elevated ''OsGA2ox'' gene expression(FIG2) <ref name="ref4" />.<br />
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===Expression===<br />
*''GI'' encodes a nuclear protein that defines the proper amplitude and period length of circadian rhythms. ''GI'' expression is regulated by the circadian clock, with a peak in transcript levels at 8 to 10 h after dawn. The ''GI'' protein acts between the circadian oscillator and ''CO'' to promote flowering by increasing ''CO'' and ''FT'' mRNA abundance<ref name="ref1" />. The expression of the ''OsGI'' messenger RNA was circadian-controlled and that its temporal expression pattern was very similar to that of ''GI'' under both SD and LD conditions. Decreased ''OsGI'' expression led to early flowering under LD conditions and late flowering under SD conditions relative to the wild type, and that this early flowering phenotype under LD conditions was the opposite of that of the Arabidopsis ''gi'' mutants, confirming the opposite role of the ''GI'' gene in flowering under LD conditions in rice and Arabidopsis<ref name="ref5" />.<br />
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===Evolution===<br />
* There is some evolutionary conservation between rice circadian clock–related genes and those associated with Arabidopsis circadian clocks, where the circadian clock consists of interlocked subloops, it is also possible that ''Os-GI'' is a member of a subloop in rice circadian clocks<ref name="ref4" />.But there is a divergence in the photoperiodic pathways of these two species arises from the molecular mechanisms controlling the expression of ''FT/Hd3a'' by ''CO/Hd1(Se1)''<ref name="ref5" />(figure 3)<br />
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==Labs working on this gene==<br />
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==Knowledge Extension==<br />
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==References==<br />
Please input cited references here.<br />
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==Structured Information==<br />
{{JaponicaGene|<br />
GeneName = Os01g0182600|<br />
Description = GIGANTEA protein|<br />
Version = NM_001048755.1 GI:115434923 GeneID:4325329|<br />
Length = 9083 bp|<br />
Definition = Oryza sativa Japonica Group Os01g0182600, complete gene.|<br />
Source = Oryza sativa Japonica Group<br />
<br />
ORGANISM Oryza sativa Japonica Group<br />
Eukaryota; Viridiplantae; Streptophyta; Embryophyta; Tracheophyta;<br />
Spermatophyta; Magnoliophyta; Liliopsida; Poales; Poaceae; BEP<br />
clade; Ehrhartoideae; Oryzeae; Oryza.<br />
|<br />
Chromosome = [[:category:Japonica Chromosome 1|Chromosome 1]]|<br />
AP = Chromosome 1:4328362..4337444|<br />
CDS = 4328725..4329030,4329123..4329215,4329333..4329551,4329752..4329820,4329925..4331421<br>,4332220..4332429,4332563..4332713,4332791..4333031,4333898..4333956<br>,4334379..4334653,4334813..4334955,4335034..4335088,4335181..4335258<br>,4335351..4335437|<br />
GCID = <gbrowseImage1><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage1>|<br />
GSID = <gbrowseImage2><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage2>|<br />
CDNA = <cdnaseq>atgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactgcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcatagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcggcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctga</cdnaseq>|<br />
AA = <aaseq>MSASNEKWIDGLQFSSLFWPPPQDSQQKQAQILAYVEYFGQFTA DSEQFPEDIAQLIQSCYPSKEKRLVDEVLATFVLHHPEHGHAVVHPILSRIIDGTLSY DRNGFPFMSFISLFSHTSEKEYSEQWALACGEILRVLTHYNRPIFKVDHQHSEAECSS TSDQASSCESMEKRANGSPRNEPDRKPLRPLSPWITDILLAAPLGIRSDYFRWCGGVM GKYAAGGELKPPTTAYSRGSGKHPQLMPSTPRWAVANGAGVILSVCDEEVARYETANL TAAAVPALLLPPPTTPLDEHLVAGLPPLEPYARLFHRYYAIATPSATQRLLFGLLEAP PSWAPDALDAAVQLVELLRAAEDYDSGMRLPKNWMHLHFLRAIGTAMSMRAGIAADTS AALLFRILSQPTLLFPPLRHAEGVELHHEPLGGYVSSYKRQLEVPASEATIDATAQGI ASMLCAHGPDVEWRICTIWEAAYGLLPLSSSAVDLPEIVVAAPLQPPTLSWSLYLPLL KVFEYLPRGSPSEACLMRIFVATVEAILRRTFPSETSEQSRKPRSQSKNLAVAELRTM IHSLFVESCASMDLASRLLFVVLTVCVSHQALPGGSKRPTGSDNHSSEEVTNDSRLTN GRNRCKKRQGPVATFDSYVLAAVCALSCELQLFPFISKNGNHSNLKDSIKIVIPGKTT GISNELHNSISSAILHTRRILGILEALFSLKPSSVGTSWSYSSNEIVAAAMVAAHVSE LFRRSRPCLNALSALKQCKWDAEISTRASSLYHLIDLHGKTVTSIVNKAEPLEAHLTL TPVKKDEPPIEEKNINSSDGGALEKKDASRSHRKNGFARPLLKCAEDVILNGDVASTS GKAIASLQVEASDLANFLTMDRNGGYRGSQTLLRSVLSEKQELCFSVVSLLWQKLIAS PEMQMSAESTSAHQGWRKVVDALCDIVSASPTKASAAIVLQAEKDLQPWIARDDEQGQ KMWRVNQRIVKLIAELMRNHDSPEALVILASASDLLLRATDGMLVDGEACTLPQLELL EVTARAVHLIVEWGDSGVSVADGLSNLLKCRLSTTIRCLSHPSAHVRALSMSVLRDIL NSGQINSSKLIQGEHRNGIQSPTYQCLAASIINWQADVERCIEWEAHSRRATGLTLAF LTAAAKELGCPLTC</aaseq>|<br />
DNA = <dnaseqindica>8415..8720#8230..8322#7894..8112#7625..7693#6024..7520#5016..5225#4732..4882#4414..4654#3489..3547#2792..3066#2490..2632#2357..2411#2187..2264#2008..2094#ccgagccgtttctctccctctctctctcctcctccttcccctgcgcctcctccaccgcccgtgtcgccgccgcgcgcgtctatcccctcgccgccgccgccgtctacgccgcgtgaggtccggattggttgaaaggctgatcgaggtaatgcttttttatttttcccgtgcggttttagtttttctgtgctgtgcttggtggttttttctttaaaactatttttaggctggttttgatgatggcctttctaaggttgtcgtgtatggggggcgaggtggttcgggtggatttgtggtgttatgttgaggggatttttgtttggggggagagagttttggtggtagatgcggttgggattttgaagacgtgaaagggaaattagttttgaaatttgacggcgaggttgagctgattagtattgtaataattcgtggcatcggatagatcaacgcgagatggatgcttattactgcgaatttcaggaggtttttgtggtttttttaggcgagtagcgttgattctagctggattttgctgcttcgtggaggcgtttagatgtggtgcatacggctgtggatgtaatttgtgcggggataattactgaaaattaattcgcagtaagctagctctgctttaatattttttcccctgaaacagtaacaagggatcccttcctcatttcgttaacagtggaagtgaatacagacaccaaccgatgagcagagacacctgtaattaaacaccaaaagaagaacctgttttgttcctatatattttccatttctatgctttaggtaatatactttctggttcggtgtcatagtacccgaaatagaatctttgcttgccaaaaaaagaaagaataacaagatagaacaacatcattaaattctgtcttagacaatacatttataatcgaccaacaaaaacagaccgatgagtgacgtggaataacaaatgaagtataaatgtcacgaaaaacacatctattgattttgtgtgataagccataggagcctacgaggaaccaagtgaatggttctgtacaccacatccattagcattaccataaatttgcctatttctatgatgccttccttaacatatttatcgacatgtgagcttgcaccataactaaatgttatggtaaactttaccttataaatgattgcaacatgtaccttttcatttcgataattaaaaacgagaaagcttccacctttttcctagatatattttttcgtttgattatgaatttagttgtacttgacaataaaataatgcgcttgtaagattatgcaattttcgttgcattacctccctctcttttcatttgtcagtttttttctactccctccgtttcatattttaagtcgtttgacttttttcctagtcagactttgttaagtttgacaaagtttatagtaaaatttagcaatatctaaaatattaaattggtttcatcaaatctaacattgaatatattttgataatatgttttttttgtgttgaaaatacttctatatttttctataaacttagtcaaacttggaagaaatttcagtaagaaaaaagtcaaacggcttataatatgaaacggagggagtaaatttttttttgcattaattaaaattcatgtattttcctccttttttggtttttgttgtgcgatatattacagtgtagagactctgaagctcatatggttggcattggttgatatgataacgtgtcacgttgtgagatgcagacccccgttatgggggttgattgaatgaagtggatagcactcttataactttcttgctcgaagtttcttgataccaaggcaagtagcttataagaagatttggactttggtggcataaatggatgaactaactggcggcttcaacgtggtgttgattacaggtgcttttattgctaactggagcagtttgtgtagtttctttctttctctttgagaattctctcagtttcttgggcaggctcttgagctaccaagtatgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggtaacattattcctatttgattcttgttagaaattctatatgttgaacatacttgcctcaatgggaaacttggttgcgtctcctctctgaaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcaggtactgtatctctaagcatgtttagtatcttctgtgataaaatctttctccagtgtgttaacttttgcattctttttcttgcatgcttgcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattaggtaatttgatccagtgatcaatacatgttccacaaacaattgaagaaagagtttgacatatttttctcatatttccagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgaggtaaggcccctactctaatggttagttccaatttattggttcatttctctgtgtttgataagattttgcctctgctgtgcatttcaatattatatctaaactgtgctgcttatagcttatcaaactactcttgctcataaagatgcaacttttatgcagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgagtcataataaatgataacatattattgttctgtggactgaaaatctgaaatcctcctttcttgagaaatacaacctcaattttatggtgatattgacaagaaggaagagcgcatataatttaaagttaaaacacggaaaataatagaagtatgtgagcagaacgcaggagcaaggggtggaactcataacaagttcaaacttcatgcatgttcataattccccaatgcaagtatggcaagaaaaatagacaacttttaaaagtatgctaatggaactgcatgcattgtgtaaacttgcttttgtcaagcatgttggaacatactatatatttgctgctgatgatgaaaaatttgcttgagtactcttctattgctaatggacgcttaccattccctgttttgtcttcaaatgcatcaggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactggttggtaaaatcagcaatggaattttctatcgtgaattttttataattattgtaacattaattactcccatgtctttttattctgttgagataccatgtgacaatgtctacaattgcatattttgcattctttggtagatcagttcccttcttaagatccccaaagggaaactggagtatcaataacttactccctcaaataaaaaaaaagtgatactttaggcataaacacattcttcaaaacaaaactttgaatagcagtttcttttgtaatatattaataaagcatggaaaaattataatgtgaagaatgtacattcagggacaaatatactcatcatttttgcatggttgatcttaatgtccatagagagattgatcagaccaaaagtttatttttgacaggagtgagtatggattataataggagtgaatggcaatatgctgtgaaagtgagcattttatatttgtttactactagcatttggtgacaaagcaaatgatgtatattgtaaatcaaagtaaacaagtagaacatgtcctgtggcgacaagacataacctacaaattaaagttagcctgattgtttattttactgggagcacattttttctagattctagagtacatattattaaaaaaggtgtgatatgtttgcaaattactttttaatggaagtttgtaaattactttgaaacttgcattcagtttttttgcttcttaaaaaacatagtgcatatgggctttatgagctcatactatgcatgtttttctgcattggagtgggacctttatctttttcttgcaactcggatgctaatgcttattataaacacaagttactaacatgtgcttgggtcattcagcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcataggtaagctcttatgatcctgtccactgttgtaccatgtgtactttccaaatgattgctaacatgctatttttcgtcagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcgggtatatcatacatctctctggctgttgtttcagtgctatcaatcaagatgacacatcatgcaatattatcataacagcacattatgtgttttgaatattttattgattgtcatagctttgttttatgttttagcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcaggtatgcagtgtttcctgcacctacttgctacctattctgtatttgatgtattcaatttcctttttatcagttttctcgaaatatatgctaataggcttgtcactaaacggctacattatccagactctgttctgaatttttttttcattaagtactgttagctgccagattacatcatcatgactaaagtagtttagctacaatattgcaaagctagatttgtctagcctctattgttttgaatagaaaaggcaggtatgcagtgtttcctgcacctacttgctacctattctgtatttgatgtattcaatttcctttttatcagttttctcgaaatatatgctaataggcttgtcactaaacggctacattatccagactctgttctgaatttttttttcattaagtactgttagctgccagattacatcatcatgactaaagtagtttagctacaatattgcaaagctagatttgtctagcctctattgttttgaatagaaaattaaatcttttggtcatttttttgtaaccttttaggagtctctgtatgcttgtcgtcaacatgctatgataagtttatatcagttcttttggagctatgatgtattcttgtatattttatgactgctaggatattatgttaaaactgttttaaaggtatcatcaaacaagttaatatagatattgctacctaccaaagttggaatttgattatgtcgtaaaatatatgcaccctacatgttttagcttcatattgacttatattttcatatgtgctccttcaactgcacaaagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggtatggtcattcctgcttataagataccagaataatgctatgaagaccgtccttggttgtcagcattcttatacacaatgataatctttgtttttcatgggtaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggtcaagatattgttattgttatatttacatacttgttatcatatttaaaagcttcatattgtatgtacctttgaaaaaaaaagttccatactgtagcttcttttgttctcatttcttggtttcaaagcatcagcatcattaccgatataaaatatacccatttcctcagtcatcttccccgactcttgtatctttcacaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagaggtaacaccttcattcatgaggtccttaggtgcaaatcccagtgaagaataatcagttctactttgtatggttcatctcgcttactgaccggttcatctcgggatcaattacgagcagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaaggcatgtcctcttctctgatctcactcacgttcatgccaaacttactcatcatcatccatgatccatcatcctgactgaatcttgatatgcagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctgacaaggccacaccatgactgtcactgcaagcagctgctgtaggatcagcgagtaggaaaggatgacttgccggccagtttcctgagatgtgatttaagactgatattagctgatgttcccaagcatgatacggggcttgctcccaagatgtgatttttactttacttttggttaatgtgctaccgctgacattagatgttcaagcatatgaaaactgcttgtgctgtaacttgtatgtctgtaacgtaaaaatctttccatatggtaaccggggaagcagaggtatatccttttctgctgttgtatctatttccagtttcctgctgaacttgcaaatgaatattcagcaatttccgttttgctc</dnaseqindica>|<br />
Link = [http://www.ncbi.nlm.nih.gov/nuccore/NM_001048755.1 RefSeq:Os01g0182600]|<br />
}}<br />
[[Category:Genes]]<br />
[[Category:Japonica mRNA]]<br />
[[Category:Oryza Sativa Japonica Group]]<br />
[[Category:Japonica Genes]]<br />
[[Category:Japonica Chromosome 1]]<br />
[[Category:Chromosome 1]]</div>Mikahttps://ngdc.cncb.ac.cn/ricewiki/index.php?title=Os01g0182600&diff=183672Os01g01826002014-06-10T14:03:24Z<p>Mika: /* Evolution */</p>
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<div>Please input one-sentence summary here.<br />
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==Annotated Information==<br />
===Function===<br />
*''OsGI'',a ''GI'' ortholog, regulates flowering time in response to photoperiodic conditions. Mutations in this gene delay flowering in inductive SDs (short-days),whereas flowering is marginally induced under LDs(long-days).In comparison, over expression of ''OsGI'' in transgenic rice causes late flowering in both SDs and LDs.In addition, ''OsGI'' has function to regulate ''Hd1'', a gene that also shows circadian rhythm<ref name="ref1" />. ''Os-GI'' affects gene expression of approximately 75% of genes among 27,201 genes and is required for strong amplitudes and proper phase-setting of global gene expression under natural field conditions<ref name="ref2" />. <br />
*The cDNA of ''OsGI'' firstly found in ''se5'' mutant that is completely photoperiod-insensitive, the expression patterns of rice and Arabidopsis ''GI'' gene are both photoperiodically regulated and are very similar under LD and SD conditions(FIG1)<ref name="ref3" />. ''GI'' is required for phase-dependent ''CO''transcription through the interaction with FKF1, which leads to cycling DNA binding with one finger (CDF) degradation and consequent derepression of CO. In addition, it has been known that GI protein interacts with ZEITLUPE (ZTL) F-box protein to control the diurnal rhythm of TOC1 protein in regulation of the Arabidopsis circadian clock<ref name="ref4" />. <br />
*''Os-GI'' also has function to maintain bioactive GA level through the regulation of the GA-deactivating enzyme genes in rice, ''Os-GI'' represses major ''OsGA2ox'' genes in order to maintain a proper bioactive GA level, ''osgi-1'' plants showed a semi-dwarf phenotype with elevated ''OsGA2ox'' gene expression(FIG2) <ref name="ref4" />.<br />
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===Expression===<br />
Please input expression information here.<br />
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===Evolution===<br />
* There is some evolutionary conservation between rice circadian clock–related genes and those associated with Arabidopsis circadian clocks, where the circadian clock consists of interlocked subloops, it is also possible that ''Os-GI'' is a member of a subloop in rice circadian clocks<ref name="ref4" />.But there is a divergence in the photoperiodic pathways of these two species arises from the molecular mechanisms controlling the expression of ''FT/Hd3a'' by ''CO/Hd1(Se1)''<ref name="ref5" />(figure 3)<br />
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==Labs working on this gene==<br />
Please input related labs here.<br />
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==Knowledge Extension==<br />
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==References==<br />
Please input cited references here.<br />
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==Structured Information==<br />
{{JaponicaGene|<br />
GeneName = Os01g0182600|<br />
Description = GIGANTEA protein|<br />
Version = NM_001048755.1 GI:115434923 GeneID:4325329|<br />
Length = 9083 bp|<br />
Definition = Oryza sativa Japonica Group Os01g0182600, complete gene.|<br />
Source = Oryza sativa Japonica Group<br />
<br />
ORGANISM Oryza sativa Japonica Group<br />
Eukaryota; Viridiplantae; Streptophyta; Embryophyta; Tracheophyta;<br />
Spermatophyta; Magnoliophyta; Liliopsida; Poales; Poaceae; BEP<br />
clade; Ehrhartoideae; Oryzeae; Oryza.<br />
|<br />
Chromosome = [[:category:Japonica Chromosome 1|Chromosome 1]]|<br />
AP = Chromosome 1:4328362..4337444|<br />
CDS = 4328725..4329030,4329123..4329215,4329333..4329551,4329752..4329820,4329925..4331421<br>,4332220..4332429,4332563..4332713,4332791..4333031,4333898..4333956<br>,4334379..4334653,4334813..4334955,4335034..4335088,4335181..4335258<br>,4335351..4335437|<br />
GCID = <gbrowseImage1><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage1>|<br />
GSID = <gbrowseImage2><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage2>|<br />
CDNA = <cdnaseq>atgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactgcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcatagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcggcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctga</cdnaseq>|<br />
AA = <aaseq>MSASNEKWIDGLQFSSLFWPPPQDSQQKQAQILAYVEYFGQFTA DSEQFPEDIAQLIQSCYPSKEKRLVDEVLATFVLHHPEHGHAVVHPILSRIIDGTLSY DRNGFPFMSFISLFSHTSEKEYSEQWALACGEILRVLTHYNRPIFKVDHQHSEAECSS TSDQASSCESMEKRANGSPRNEPDRKPLRPLSPWITDILLAAPLGIRSDYFRWCGGVM GKYAAGGELKPPTTAYSRGSGKHPQLMPSTPRWAVANGAGVILSVCDEEVARYETANL TAAAVPALLLPPPTTPLDEHLVAGLPPLEPYARLFHRYYAIATPSATQRLLFGLLEAP PSWAPDALDAAVQLVELLRAAEDYDSGMRLPKNWMHLHFLRAIGTAMSMRAGIAADTS AALLFRILSQPTLLFPPLRHAEGVELHHEPLGGYVSSYKRQLEVPASEATIDATAQGI ASMLCAHGPDVEWRICTIWEAAYGLLPLSSSAVDLPEIVVAAPLQPPTLSWSLYLPLL KVFEYLPRGSPSEACLMRIFVATVEAILRRTFPSETSEQSRKPRSQSKNLAVAELRTM IHSLFVESCASMDLASRLLFVVLTVCVSHQALPGGSKRPTGSDNHSSEEVTNDSRLTN GRNRCKKRQGPVATFDSYVLAAVCALSCELQLFPFISKNGNHSNLKDSIKIVIPGKTT GISNELHNSISSAILHTRRILGILEALFSLKPSSVGTSWSYSSNEIVAAAMVAAHVSE LFRRSRPCLNALSALKQCKWDAEISTRASSLYHLIDLHGKTVTSIVNKAEPLEAHLTL TPVKKDEPPIEEKNINSSDGGALEKKDASRSHRKNGFARPLLKCAEDVILNGDVASTS GKAIASLQVEASDLANFLTMDRNGGYRGSQTLLRSVLSEKQELCFSVVSLLWQKLIAS PEMQMSAESTSAHQGWRKVVDALCDIVSASPTKASAAIVLQAEKDLQPWIARDDEQGQ KMWRVNQRIVKLIAELMRNHDSPEALVILASASDLLLRATDGMLVDGEACTLPQLELL EVTARAVHLIVEWGDSGVSVADGLSNLLKCRLSTTIRCLSHPSAHVRALSMSVLRDIL NSGQINSSKLIQGEHRNGIQSPTYQCLAASIINWQADVERCIEWEAHSRRATGLTLAF LTAAAKELGCPLTC</aaseq>|<br />
DNA = <dnaseqindica>8415..8720#8230..8322#7894..8112#7625..7693#6024..7520#5016..5225#4732..4882#4414..4654#3489..3547#2792..3066#2490..2632#2357..2411#2187..2264#2008..2094#ccgagccgtttctctccctctctctctcctcctccttcccctgcgcctcctccaccgcccgtgtcgccgccgcgcgcgtctatcccctcgccgccgccgccgtctacgccgcgtgaggtccggattggttgaaaggctgatcgaggtaatgcttttttatttttcccgtgcggttttagtttttctgtgctgtgcttggtggttttttctttaaaactatttttaggctggttttgatgatggcctttctaaggttgtcgtgtatggggggcgaggtggttcgggtggatttgtggtgttatgttgaggggatttttgtttggggggagagagttttggtggtagatgcggttgggattttgaagacgtgaaagggaaattagttttgaaatttgacggcgaggttgagctgattagtattgtaataattcgtggcatcggatagatcaacgcgagatggatgcttattactgcgaatttcaggaggtttttgtggtttttttaggcgagtagcgttgattctagctggattttgctgcttcgtggaggcgtttagatgtggtgcatacggctgtggatgtaatttgtgcggggataattactgaaaattaattcgcagtaagctagctctgctttaatattttttcccctgaaacagtaacaagggatcccttcctcatttcgttaacagtggaagtgaatacagacaccaaccgatgagcagagacacctgtaattaaacaccaaaagaagaacctgttttgttcctatatattttccatttctatgctttaggtaatatactttctggttcggtgtcatagtacccgaaatagaatctttgcttgccaaaaaaagaaagaataacaagatagaacaacatcattaaattctgtcttagacaatacatttataatcgaccaacaaaaacagaccgatgagtgacgtggaataacaaatgaagtataaatgtcacgaaaaacacatctattgattttgtgtgataagccataggagcctacgaggaaccaagtgaatggttctgtacaccacatccattagcattaccataaatttgcctatttctatgatgccttccttaacatatttatcgacatgtgagcttgcaccataactaaatgttatggtaaactttaccttataaatgattgcaacatgtaccttttcatttcgataattaaaaacgagaaagcttccacctttttcctagatatattttttcgtttgattatgaatttagttgtacttgacaataaaataatgcgcttgtaagattatgcaattttcgttgcattacctccctctcttttcatttgtcagtttttttctactccctccgtttcatattttaagtcgtttgacttttttcctagtcagactttgttaagtttgacaaagtttatagtaaaatttagcaatatctaaaatattaaattggtttcatcaaatctaacattgaatatattttgataatatgttttttttgtgttgaaaatacttctatatttttctataaacttagtcaaacttggaagaaatttcagtaagaaaaaagtcaaacggcttataatatgaaacggagggagtaaatttttttttgcattaattaaaattcatgtattttcctccttttttggtttttgttgtgcgatatattacagtgtagagactctgaagctcatatggttggcattggttgatatgataacgtgtcacgttgtgagatgcagacccccgttatgggggttgattgaatgaagtggatagcactcttataactttcttgctcgaagtttcttgataccaaggcaagtagcttataagaagatttggactttggtggcataaatggatgaactaactggcggcttcaacgtggtgttgattacaggtgcttttattgctaactggagcagtttgtgtagtttctttctttctctttgagaattctctcagtttcttgggcaggctcttgagctaccaagtatgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggtaacattattcctatttgattcttgttagaaattctatatgttgaacatacttgcctcaatgggaaacttggttgcgtctcctctctgaaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcaggtactgtatctctaagcatgtttagtatcttctgtgataaaatctttctccagtgtgttaacttttgcattctttttcttgcatgcttgcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattaggtaatttgatccagtgatcaatacatgttccacaaacaattgaagaaagagtttgacatatttttctcatatttccagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgaggtaaggcccctactctaatggttagttccaatttattggttcatttctctgtgtttgataagattttgcctctgctgtgcatttcaatattatatctaaactgtgctgcttatagcttatcaaactactcttgctcataaagatgcaacttttatgcagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgagtcataataaatgataacatattattgttctgtggactgaaaatctgaaatcctcctttcttgagaaatacaacctcaattttatggtgatattgacaagaaggaagagcgcatataatttaaagttaaaacacggaaaataatagaagtatgtgagcagaacgcaggagcaaggggtggaactcataacaagttcaaacttcatgcatgttcataattccccaatgcaagtatggcaagaaaaatagacaacttttaaaagtatgctaatggaactgcatgcattgtgtaaacttgcttttgtcaagcatgttggaacatactatatatttgctgctgatgatgaaaaatttgcttgagtactcttctattgctaatggacgcttaccattccctgttttgtcttcaaatgcatcaggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactggttggtaaaatcagcaatggaattttctatcgtgaattttttataattattgtaacattaattactcccatgtctttttattctgttgagataccatgtgacaatgtctacaattgcatattttgcattctttggtagatcagttcccttcttaagatccccaaagggaaactggagtatcaataacttactccctcaaataaaaaaaaagtgatactttaggcataaacacattcttcaaaacaaaactttgaatagcagtttcttttgtaatatattaataaagcatggaaaaattataatgtgaagaatgtacattcagggacaaatatactcatcatttttgcatggttgatcttaatgtccatagagagattgatcagaccaaaagtttatttttgacaggagtgagtatggattataataggagtgaatggcaatatgctgtgaaagtgagcattttatatttgtttactactagcatttggtgacaaagcaaatgatgtatattgtaaatcaaagtaaacaagtagaacatgtcctgtggcgacaagacataacctacaaattaaagttagcctgattgtttattttactgggagcacattttttctagattctagagtacatattattaaaaaaggtgtgatatgtttgcaaattactttttaatggaagtttgtaaattactttgaaacttgcattcagtttttttgcttcttaaaaaacatagtgcatatgggctttatgagctcatactatgcatgtttttctgcattggagtgggacctttatctttttcttgcaactcggatgctaatgcttattataaacacaagttactaacatgtgcttgggtcattcagcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcataggtaagctcttatgatcctgtccactgttgtaccatgtgtactttccaaatgattgctaacatgctatttttcgtcagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcgggtatatcatacatctctctggctgttgtttcagtgctatcaatcaagatgacacatcatgcaatattatcataacagcacattatgtgttttgaatattttattgattgtcatagctttgttttatgttttagcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcaggtatgcagtgtttcctgcacctacttgctacctattctgtatttgatgtattcaatttcctttttatcagttttctcgaaatatatgctaataggcttgtcactaaacggctacattatccagactctgttctgaatttttttttcattaagtactgttagctgccagattacatcatcatgactaaagtagtttagctacaatattgcaaagctagatttgtctagcctctattgttttgaatagaaaaggcaggtatgcagtgtttcctgcacctacttgctacctattctgtatttgatgtattcaatttcctttttatcagttttctcgaaatatatgctaataggcttgtcactaaacggctacattatccagactctgttctgaatttttttttcattaagtactgttagctgccagattacatcatcatgactaaagtagtttagctacaatattgcaaagctagatttgtctagcctctattgttttgaatagaaaattaaatcttttggtcatttttttgtaaccttttaggagtctctgtatgcttgtcgtcaacatgctatgataagtttatatcagttcttttggagctatgatgtattcttgtatattttatgactgctaggatattatgttaaaactgttttaaaggtatcatcaaacaagttaatatagatattgctacctaccaaagttggaatttgattatgtcgtaaaatatatgcaccctacatgttttagcttcatattgacttatattttcatatgtgctccttcaactgcacaaagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggtatggtcattcctgcttataagataccagaataatgctatgaagaccgtccttggttgtcagcattcttatacacaatgataatctttgtttttcatgggtaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggtcaagatattgttattgttatatttacatacttgttatcatatttaaaagcttcatattgtatgtacctttgaaaaaaaaagttccatactgtagcttcttttgttctcatttcttggtttcaaagcatcagcatcattaccgatataaaatatacccatttcctcagtcatcttccccgactcttgtatctttcacaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagaggtaacaccttcattcatgaggtccttaggtgcaaatcccagtgaagaataatcagttctactttgtatggttcatctcgcttactgaccggttcatctcgggatcaattacgagcagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaaggcatgtcctcttctctgatctcactcacgttcatgccaaacttactcatcatcatccatgatccatcatcctgactgaatcttgatatgcagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctgacaaggccacaccatgactgtcactgcaagcagctgctgtaggatcagcgagtaggaaaggatgacttgccggccagtttcctgagatgtgatttaagactgatattagctgatgttcccaagcatgatacggggcttgctcccaagatgtgatttttactttacttttggttaatgtgctaccgctgacattagatgttcaagcatatgaaaactgcttgtgctgtaacttgtatgtctgtaacgtaaaaatctttccatatggtaaccggggaagcagaggtatatccttttctgctgttgtatctatttccagtttcctgctgaacttgcaaatgaatattcagcaatttccgttttgctc</dnaseqindica>|<br />
Link = [http://www.ncbi.nlm.nih.gov/nuccore/NM_001048755.1 RefSeq:Os01g0182600]|<br />
}}<br />
[[Category:Genes]]<br />
[[Category:Japonica mRNA]]<br />
[[Category:Oryza Sativa Japonica Group]]<br />
[[Category:Japonica Genes]]<br />
[[Category:Japonica Chromosome 1]]<br />
[[Category:Chromosome 1]]</div>Mikahttps://ngdc.cncb.ac.cn/ricewiki/index.php?title=Os01g0182600&diff=183670Os01g01826002014-06-10T13:59:15Z<p>Mika: /* Function */</p>
<hr />
<div>Please input one-sentence summary here.<br />
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==Annotated Information==<br />
===Function===<br />
*''OsGI'',a ''GI'' ortholog, regulates flowering time in response to photoperiodic conditions. Mutations in this gene delay flowering in inductive SDs (short-days),whereas flowering is marginally induced under LDs(long-days).In comparison, over expression of ''OsGI'' in transgenic rice causes late flowering in both SDs and LDs.In addition, ''OsGI'' has function to regulate ''Hd1'', a gene that also shows circadian rhythm<ref name="ref1" />. ''Os-GI'' affects gene expression of approximately 75% of genes among 27,201 genes and is required for strong amplitudes and proper phase-setting of global gene expression under natural field conditions<ref name="ref2" />. <br />
*The cDNA of ''OsGI'' firstly found in ''se5'' mutant that is completely photoperiod-insensitive, the expression patterns of rice and Arabidopsis ''GI'' gene are both photoperiodically regulated and are very similar under LD and SD conditions(FIG1)<ref name="ref3" />. ''GI'' is required for phase-dependent ''CO''transcription through the interaction with FKF1, which leads to cycling DNA binding with one finger (CDF) degradation and consequent derepression of CO. In addition, it has been known that GI protein interacts with ZEITLUPE (ZTL) F-box protein to control the diurnal rhythm of TOC1 protein in regulation of the Arabidopsis circadian clock<ref name="ref4" />. <br />
*''Os-GI'' also has function to maintain bioactive GA level through the regulation of the GA-deactivating enzyme genes in rice, ''Os-GI'' represses major ''OsGA2ox'' genes in order to maintain a proper bioactive GA level, ''osgi-1'' plants showed a semi-dwarf phenotype with elevated ''OsGA2ox'' gene expression(FIG2) <ref name="ref4" />.<br />
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===Expression===<br />
Please input expression information here.<br />
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===Evolution===<br />
Please input evolution information here.<br />
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*there is some evolutionary conservation between rice circadian clock–related genes and those associated with Arabidopsis circadian clocks,where the circadian clock consists of interlocked subloops, it is also possible that Os-GI is a member of a subloop in rice circadian clocks<br />
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==Labs working on this gene==<br />
Please input related labs here.<br />
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==Knowledge Extension==<br />
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==References==<br />
Please input cited references here.<br />
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==Structured Information==<br />
{{JaponicaGene|<br />
GeneName = Os01g0182600|<br />
Description = GIGANTEA protein|<br />
Version = NM_001048755.1 GI:115434923 GeneID:4325329|<br />
Length = 9083 bp|<br />
Definition = Oryza sativa Japonica Group Os01g0182600, complete gene.|<br />
Source = Oryza sativa Japonica Group<br />
<br />
ORGANISM Oryza sativa Japonica Group<br />
Eukaryota; Viridiplantae; Streptophyta; Embryophyta; Tracheophyta;<br />
Spermatophyta; Magnoliophyta; Liliopsida; Poales; Poaceae; BEP<br />
clade; Ehrhartoideae; Oryzeae; Oryza.<br />
|<br />
Chromosome = [[:category:Japonica Chromosome 1|Chromosome 1]]|<br />
AP = Chromosome 1:4328362..4337444|<br />
CDS = 4328725..4329030,4329123..4329215,4329333..4329551,4329752..4329820,4329925..4331421<br>,4332220..4332429,4332563..4332713,4332791..4333031,4333898..4333956<br>,4334379..4334653,4334813..4334955,4335034..4335088,4335181..4335258<br>,4335351..4335437|<br />
GCID = <gbrowseImage1><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage1>|<br />
GSID = <gbrowseImage2><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage2>|<br />
CDNA = <cdnaseq>atgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactgcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcatagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcggcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctga</cdnaseq>|<br />
AA = <aaseq>MSASNEKWIDGLQFSSLFWPPPQDSQQKQAQILAYVEYFGQFTA DSEQFPEDIAQLIQSCYPSKEKRLVDEVLATFVLHHPEHGHAVVHPILSRIIDGTLSY DRNGFPFMSFISLFSHTSEKEYSEQWALACGEILRVLTHYNRPIFKVDHQHSEAECSS TSDQASSCESMEKRANGSPRNEPDRKPLRPLSPWITDILLAAPLGIRSDYFRWCGGVM GKYAAGGELKPPTTAYSRGSGKHPQLMPSTPRWAVANGAGVILSVCDEEVARYETANL TAAAVPALLLPPPTTPLDEHLVAGLPPLEPYARLFHRYYAIATPSATQRLLFGLLEAP PSWAPDALDAAVQLVELLRAAEDYDSGMRLPKNWMHLHFLRAIGTAMSMRAGIAADTS AALLFRILSQPTLLFPPLRHAEGVELHHEPLGGYVSSYKRQLEVPASEATIDATAQGI ASMLCAHGPDVEWRICTIWEAAYGLLPLSSSAVDLPEIVVAAPLQPPTLSWSLYLPLL KVFEYLPRGSPSEACLMRIFVATVEAILRRTFPSETSEQSRKPRSQSKNLAVAELRTM IHSLFVESCASMDLASRLLFVVLTVCVSHQALPGGSKRPTGSDNHSSEEVTNDSRLTN GRNRCKKRQGPVATFDSYVLAAVCALSCELQLFPFISKNGNHSNLKDSIKIVIPGKTT GISNELHNSISSAILHTRRILGILEALFSLKPSSVGTSWSYSSNEIVAAAMVAAHVSE LFRRSRPCLNALSALKQCKWDAEISTRASSLYHLIDLHGKTVTSIVNKAEPLEAHLTL TPVKKDEPPIEEKNINSSDGGALEKKDASRSHRKNGFARPLLKCAEDVILNGDVASTS GKAIASLQVEASDLANFLTMDRNGGYRGSQTLLRSVLSEKQELCFSVVSLLWQKLIAS PEMQMSAESTSAHQGWRKVVDALCDIVSASPTKASAAIVLQAEKDLQPWIARDDEQGQ KMWRVNQRIVKLIAELMRNHDSPEALVILASASDLLLRATDGMLVDGEACTLPQLELL EVTARAVHLIVEWGDSGVSVADGLSNLLKCRLSTTIRCLSHPSAHVRALSMSVLRDIL NSGQINSSKLIQGEHRNGIQSPTYQCLAASIINWQADVERCIEWEAHSRRATGLTLAF LTAAAKELGCPLTC</aaseq>|<br />
DNA = <dnaseqindica>8415..8720#8230..8322#7894..8112#7625..7693#6024..7520#5016..5225#4732..4882#4414..4654#3489..3547#2792..3066#2490..2632#2357..2411#2187..2264#2008..2094#ccgagccgtttctctccctctctctctcctcctccttcccctgcgcctcctccaccgcccgtgtcgccgccgcgcgcgtctatcccctcgccgccgccgccgtctacgccgcgtgaggtccggattggttgaaaggctgatcgaggtaatgcttttttatttttcccgtgcggttttagtttttctgtgctgtgcttggtggttttttctttaaaactatttttaggctggttttgatgatggcctttctaaggttgtcgtgtatggggggcgaggtggttcgggtggatttgtggtgttatgttgaggggatttttgtttggggggagagagttttggtggtagatgcggttgggattttgaagacgtgaaagggaaattagttttgaaatttgacggcgaggttgagctgattagtattgtaataattcgtggcatcggatagatcaacgcgagatggatgcttattactgcgaatttcaggaggtttttgtggtttttttaggcgagtagcgttgattctagctggattttgctgcttcgtggaggcgtttagatgtggtgcatacggctgtggatgtaatttgtgcggggataattactgaaaattaattcgcagtaagctagctctgctttaatattttttcccctgaaacagtaacaagggatcccttcctcatttcgttaacagtggaagtgaatacagacaccaaccgatgagcagagacacctgtaattaaacaccaaaagaagaacctgttttgttcctatatattttccatttctatgctttaggtaatatactttctggttcggtgtcatagtacccgaaatagaatctttgcttgccaaaaaaagaaagaataacaagatagaacaacatcattaaattctgtcttagacaatacatttataatcgaccaacaaaaacagaccgatgagtgacgtggaataacaaatgaagtataaatgtcacgaaaaacacatctattgattttgtgtgataagccataggagcctacgaggaaccaagtgaatggttctgtacaccacatccattagcattaccataaatttgcctatttctatgatgccttccttaacatatttatcgacatgtgagcttgcaccataactaaatgttatggtaaactttaccttataaatgattgcaacatgtaccttttcatttcgataattaaaaacgagaaagcttccacctttttcctagatatattttttcgtttgattatgaatttagttgtacttgacaataaaataatgcgcttgtaagattatgcaattttcgttgcattacctccctctcttttcatttgtcagtttttttctactccctccgtttcatattttaagtcgtttgacttttttcctagtcagactttgttaagtttgacaaagtttatagtaaaatttagcaatatctaaaatattaaattggtttcatcaaatctaacattgaatatattttgataatatgttttttttgtgttgaaaatacttctatatttttctataaacttagtcaaacttggaagaaatttcagtaagaaaaaagtcaaacggcttataatatgaaacggagggagtaaatttttttttgcattaattaaaattcatgtattttcctccttttttggtttttgttgtgcgatatattacagtgtagagactctgaagctcatatggttggcattggttgatatgataacgtgtcacgttgtgagatgcagacccccgttatgggggttgattgaatgaagtggatagcactcttataactttcttgctcgaagtttcttgataccaaggcaagtagcttataagaagatttggactttggtggcataaatggatgaactaactggcggcttcaacgtggtgttgattacaggtgcttttattgctaactggagcagtttgtgtagtttctttctttctctttgagaattctctcagtttcttgggcaggctcttgagctaccaagtatgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggtaacattattcctatttgattcttgttagaaattctatatgttgaacatacttgcctcaatgggaaacttggttgcgtctcctctctgaaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcaggtactgtatctctaagcatgtttagtatcttctgtgataaaatctttctccagtgtgttaacttttgcattctttttcttgcatgcttgcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattaggtaatttgatccagtgatcaatacatgttccacaaacaattgaagaaagagtttgacatatttttctcatatttccagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgaggtaaggcccctactctaatggttagttccaatttattggttcatttctctgtgtttgataagattttgcctctgctgtgcatttcaatattatatctaaactgtgctgcttatagcttatcaaactactcttgctcataaagatgcaacttttatgcagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgagtcataataaatgataacatattattgttctgtggactgaaaatctgaaatcctcctttcttgagaaatacaacctcaattttatggtgatattgacaagaaggaagagcgcatataatttaaagttaaaacacggaaaataatagaagtatgtgagcagaacgcaggagcaaggggtggaactcataacaagttcaaacttcatgcatgttcataattccccaatgcaagtatggcaagaaaaatagacaacttttaaaagtatgctaatggaactgcatgcattgtgtaaacttgcttttgtcaagcatgttggaacatactatatatttgctgctgatgatgaaaaatttgcttgagtactcttctattgctaatggacgcttaccattccctgttttgtcttcaaatgcatcaggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactggttggtaaaatcagcaatggaattttctatcgtgaattttttataattattgtaacattaattactcccatgtctttttattctgttgagataccatgtgacaatgtctacaattgcatattttgcattctttggtagatcagttcccttcttaagatccccaaagggaaactggagtatcaataacttactccctcaaataaaaaaaaagtgatactttaggcataaacacattcttcaaaacaaaactttgaatagcagtttcttttgtaatatattaataaagcatggaaaaattataatgtgaagaatgtacattcagggacaaatatactcatcatttttgcatggttgatcttaatgtccatagagagattgatcagaccaaaagtttatttttgacaggagtgagtatggattataataggagtgaatggcaatatgctgtgaaagtgagcattttatatttgtttactactagcatttggtgacaaagcaaatgatgtatattgtaaatcaaagtaaacaagtagaacatgtcctgtggcgacaagacataacctacaaattaaagttagcctgattgtttattttactgggagcacattttttctagattctagagtacatattattaaaaaaggtgtgatatgtttgcaaattactttttaatggaagtttgtaaattactttgaaacttgcattcagtttttttgcttcttaaaaaacatagtgcatatgggctttatgagctcatactatgcatgtttttctgcattggagtgggacctttatctttttcttgcaactcggatgctaatgcttattataaacacaagttactaacatgtgcttgggtcattcagcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcataggtaagctcttatgatcctgtccactgttgtaccatgtgtactttccaaatgattgctaacatgctatttttcgtcagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcgggtatatcatacatctctctggctgttgtttcagtgctatcaatcaagatgacacatcatgcaatattatcataacagcacattatgtgttttgaatattttattgattgtcatagctttgttttatgttttagcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcaggtatgcagtgtttcctgcacctacttgctacctattctgtatttgatgtattcaatttcctttttatcagttttctcgaaatatatgctaataggcttgtcactaaacggctacattatccagactctgttctgaatttttttttcattaagtactgttagctgccagattacatcatcatgactaaagtagtttagctacaatattgcaaagctagatttgtctagcctctattgttttgaatagaaaaggcaggtatgcagtgtttcctgcacctacttgctacctattctgtatttgatgtattcaatttcctttttatcagttttctcgaaatatatgctaataggcttgtcactaaacggctacattatccagactctgttctgaatttttttttcattaagtactgttagctgccagattacatcatcatgactaaagtagtttagctacaatattgcaaagctagatttgtctagcctctattgttttgaatagaaaattaaatcttttggtcatttttttgtaaccttttaggagtctctgtatgcttgtcgtcaacatgctatgataagtttatatcagttcttttggagctatgatgtattcttgtatattttatgactgctaggatattatgttaaaactgttttaaaggtatcatcaaacaagttaatatagatattgctacctaccaaagttggaatttgattatgtcgtaaaatatatgcaccctacatgttttagcttcatattgacttatattttcatatgtgctccttcaactgcacaaagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggtatggtcattcctgcttataagataccagaataatgctatgaagaccgtccttggttgtcagcattcttatacacaatgataatctttgtttttcatgggtaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggtcaagatattgttattgttatatttacatacttgttatcatatttaaaagcttcatattgtatgtacctttgaaaaaaaaagttccatactgtagcttcttttgttctcatttcttggtttcaaagcatcagcatcattaccgatataaaatatacccatttcctcagtcatcttccccgactcttgtatctttcacaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagaggtaacaccttcattcatgaggtccttaggtgcaaatcccagtgaagaataatcagttctactttgtatggttcatctcgcttactgaccggttcatctcgggatcaattacgagcagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaaggcatgtcctcttctctgatctcactcacgttcatgccaaacttactcatcatcatccatgatccatcatcctgactgaatcttgatatgcagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctgacaaggccacaccatgactgtcactgcaagcagctgctgtaggatcagcgagtaggaaaggatgacttgccggccagtttcctgagatgtgatttaagactgatattagctgatgttcccaagcatgatacggggcttgctcccaagatgtgatttttactttacttttggttaatgtgctaccgctgacattagatgttcaagcatatgaaaactgcttgtgctgtaacttgtatgtctgtaacgtaaaaatctttccatatggtaaccggggaagcagaggtatatccttttctgctgttgtatctatttccagtttcctgctgaacttgcaaatgaatattcagcaatttccgttttgctc</dnaseqindica>|<br />
Link = [http://www.ncbi.nlm.nih.gov/nuccore/NM_001048755.1 RefSeq:Os01g0182600]|<br />
}}<br />
[[Category:Genes]]<br />
[[Category:Japonica mRNA]]<br />
[[Category:Oryza Sativa Japonica Group]]<br />
[[Category:Japonica Genes]]<br />
[[Category:Japonica Chromosome 1]]<br />
[[Category:Chromosome 1]]</div>Mikahttps://ngdc.cncb.ac.cn/ricewiki/index.php?title=Os01g0182600&diff=183669Os01g01826002014-06-10T13:58:20Z<p>Mika: /* Function */</p>
<hr />
<div>Please input one-sentence summary here.<br />
<br />
==Annotated Information==<br />
===Function===<br />
*''OsGI'',a ''GI'' ortholog, regulates flowering time in response to photoperiodic conditions. Mutations in this gene delay flowering in inductive SDs (short-days),whereas flowering is marginally induced under LDs(long-days).In comparison, over expression of ''OsGI'' in transgenic rice causes late flowering in both SDs and LDs.In addition, ''OsGI'' has function to regulate ''Hd1'', a gene that also shows circadian rhythm<ref name="ref1" />. ''Os-GI'' affects gene expression of approximately 75% of genes among 27,201 genes and is required for strong amplitudes and proper phase-setting of global gene expression under natural field conditions<ref name="ref2" />. <br />
*The cDNA of ''OsGI'' firstly found in ''se5'' mutant that is completely photoperiod-insensitive, the expression patterns of rice and Arabidopsis ''GI'' gene are both photoperiodically regulated and are very similar under LD and SD conditions(FIG1)<ref name="ref3" />. ''GI'' is required for phase-dependent ''CO'' transcription through the interaction with FKF1, which leads to cycling DNA binding with one finger (CDF) degradation and consequent derepression of CO. In addition, it has been known that GI protein interacts with ZEITLUPE (ZTL) F-box protein to control the diurnal rhythm of TOC1 protein in regulation of the Arabidopsis circadian clock<ref name="ref4" />. <br />
*''Os-GI'' also has function to maintain bioactive GA level through the regulation of the GA-deactivating enzyme genes in rice, ''Os-GI'' represses major <br />
''OsGA2ox'' genes in order to maintain a proper bioactive GA level, ''osgi-1'' plants showed a semi-dwarf phenotype with elevated ''OsGA2ox'' gene expression(FIG2) <ref name="ref4" />.<br />
<br />
===Expression===<br />
Please input expression information here.<br />
<br />
===Evolution===<br />
Please input evolution information here.<br />
<br />
*there is some evolutionary conservation between rice circadian clock–related genes and those associated with Arabidopsis circadian clocks,where the circadian clock consists of interlocked subloops, it is also possible that Os-GI is a member of a subloop in rice circadian clocks<br />
<br />
==Labs working on this gene==<br />
Please input related labs here.<br />
<br />
==Knowledge Extension==<br />
<br />
==References==<br />
Please input cited references here.<br />
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==Structured Information==<br />
{{JaponicaGene|<br />
GeneName = Os01g0182600|<br />
Description = GIGANTEA protein|<br />
Version = NM_001048755.1 GI:115434923 GeneID:4325329|<br />
Length = 9083 bp|<br />
Definition = Oryza sativa Japonica Group Os01g0182600, complete gene.|<br />
Source = Oryza sativa Japonica Group<br />
<br />
ORGANISM Oryza sativa Japonica Group<br />
Eukaryota; Viridiplantae; Streptophyta; Embryophyta; Tracheophyta;<br />
Spermatophyta; Magnoliophyta; Liliopsida; Poales; Poaceae; BEP<br />
clade; Ehrhartoideae; Oryzeae; Oryza.<br />
|<br />
Chromosome = [[:category:Japonica Chromosome 1|Chromosome 1]]|<br />
AP = Chromosome 1:4328362..4337444|<br />
CDS = 4328725..4329030,4329123..4329215,4329333..4329551,4329752..4329820,4329925..4331421<br>,4332220..4332429,4332563..4332713,4332791..4333031,4333898..4333956<br>,4334379..4334653,4334813..4334955,4335034..4335088,4335181..4335258<br>,4335351..4335437|<br />
GCID = <gbrowseImage1><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage1>|<br />
GSID = <gbrowseImage2><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage2>|<br />
CDNA = <cdnaseq>atgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactgcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcatagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcggcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctga</cdnaseq>|<br />
AA = <aaseq>MSASNEKWIDGLQFSSLFWPPPQDSQQKQAQILAYVEYFGQFTA DSEQFPEDIAQLIQSCYPSKEKRLVDEVLATFVLHHPEHGHAVVHPILSRIIDGTLSY DRNGFPFMSFISLFSHTSEKEYSEQWALACGEILRVLTHYNRPIFKVDHQHSEAECSS TSDQASSCESMEKRANGSPRNEPDRKPLRPLSPWITDILLAAPLGIRSDYFRWCGGVM GKYAAGGELKPPTTAYSRGSGKHPQLMPSTPRWAVANGAGVILSVCDEEVARYETANL TAAAVPALLLPPPTTPLDEHLVAGLPPLEPYARLFHRYYAIATPSATQRLLFGLLEAP PSWAPDALDAAVQLVELLRAAEDYDSGMRLPKNWMHLHFLRAIGTAMSMRAGIAADTS AALLFRILSQPTLLFPPLRHAEGVELHHEPLGGYVSSYKRQLEVPASEATIDATAQGI ASMLCAHGPDVEWRICTIWEAAYGLLPLSSSAVDLPEIVVAAPLQPPTLSWSLYLPLL KVFEYLPRGSPSEACLMRIFVATVEAILRRTFPSETSEQSRKPRSQSKNLAVAELRTM IHSLFVESCASMDLASRLLFVVLTVCVSHQALPGGSKRPTGSDNHSSEEVTNDSRLTN GRNRCKKRQGPVATFDSYVLAAVCALSCELQLFPFISKNGNHSNLKDSIKIVIPGKTT GISNELHNSISSAILHTRRILGILEALFSLKPSSVGTSWSYSSNEIVAAAMVAAHVSE LFRRSRPCLNALSALKQCKWDAEISTRASSLYHLIDLHGKTVTSIVNKAEPLEAHLTL TPVKKDEPPIEEKNINSSDGGALEKKDASRSHRKNGFARPLLKCAEDVILNGDVASTS GKAIASLQVEASDLANFLTMDRNGGYRGSQTLLRSVLSEKQELCFSVVSLLWQKLIAS PEMQMSAESTSAHQGWRKVVDALCDIVSASPTKASAAIVLQAEKDLQPWIARDDEQGQ KMWRVNQRIVKLIAELMRNHDSPEALVILASASDLLLRATDGMLVDGEACTLPQLELL EVTARAVHLIVEWGDSGVSVADGLSNLLKCRLSTTIRCLSHPSAHVRALSMSVLRDIL NSGQINSSKLIQGEHRNGIQSPTYQCLAASIINWQADVERCIEWEAHSRRATGLTLAF LTAAAKELGCPLTC</aaseq>|<br />
DNA = <dnaseqindica>8415..8720#8230..8322#7894..8112#7625..7693#6024..7520#5016..5225#4732..4882#4414..4654#3489..3547#2792..3066#2490..2632#2357..2411#2187..2264#2008..2094#ccgagccgtttctctccctctctctctcctcctccttcccctgcgcctcctccaccgcccgtgtcgccgccgcgcgcgtctatcccctcgccgccgccgccgtctacgccgcgtgaggtccggattggttgaaaggctgatcgaggtaatgcttttttatttttcccgtgcggttttagtttttctgtgctgtgcttggtggttttttctttaaaactatttttaggctggttttgatgatggcctttctaaggttgtcgtgtatggggggcgaggtggttcgggtggatttgtggtgttatgttgaggggatttttgtttggggggagagagttttggtggtagatgcggttgggattttgaagacgtgaaagggaaattagttttgaaatttgacggcgaggttgagctgattagtattgtaataattcgtggcatcggatagatcaacgcgagatggatgcttattactgcgaatttcaggaggtttttgtggtttttttaggcgagtagcgttgattctagctggattttgctgcttcgtggaggcgtttagatgtggtgcatacggctgtggatgtaatttgtgcggggataattactgaaaattaattcgcagtaagctagctctgctttaatattttttcccctgaaacagtaacaagggatcccttcctcatttcgttaacagtggaagtgaatacagacaccaaccgatgagcagagacacctgtaattaaacaccaaaagaagaacctgttttgttcctatatattttccatttctatgctttaggtaatatactttctggttcggtgtcatagtacccgaaatagaatctttgcttgccaaaaaaagaaagaataacaagatagaacaacatcattaaattctgtcttagacaatacatttataatcgaccaacaaaaacagaccgatgagtgacgtggaataacaaatgaagtataaatgtcacgaaaaacacatctattgattttgtgtgataagccataggagcctacgaggaaccaagtgaatggttctgtacaccacatccattagcattaccataaatttgcctatttctatgatgccttccttaacatatttatcgacatgtgagcttgcaccataactaaatgttatggtaaactttaccttataaatgattgcaacatgtaccttttcatttcgataattaaaaacgagaaagcttccacctttttcctagatatattttttcgtttgattatgaatttagttgtacttgacaataaaataatgcgcttgtaagattatgcaattttcgttgcattacctccctctcttttcatttgtcagtttttttctactccctccgtttcatattttaagtcgtttgacttttttcctagtcagactttgttaagtttgacaaagtttatagtaaaatttagcaatatctaaaatattaaattggtttcatcaaatctaacattgaatatattttgataatatgttttttttgtgttgaaaatacttctatatttttctataaacttagtcaaacttggaagaaatttcagtaagaaaaaagtcaaacggcttataatatgaaacggagggagtaaatttttttttgcattaattaaaattcatgtattttcctccttttttggtttttgttgtgcgatatattacagtgtagagactctgaagctcatatggttggcattggttgatatgataacgtgtcacgttgtgagatgcagacccccgttatgggggttgattgaatgaagtggatagcactcttataactttcttgctcgaagtttcttgataccaaggcaagtagcttataagaagatttggactttggtggcataaatggatgaactaactggcggcttcaacgtggtgttgattacaggtgcttttattgctaactggagcagtttgtgtagtttctttctttctctttgagaattctctcagtttcttgggcaggctcttgagctaccaagtatgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggtaacattattcctatttgattcttgttagaaattctatatgttgaacatacttgcctcaatgggaaacttggttgcgtctcctctctgaaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcaggtactgtatctctaagcatgtttagtatcttctgtgataaaatctttctccagtgtgttaacttttgcattctttttcttgcatgcttgcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattaggtaatttgatccagtgatcaatacatgttccacaaacaattgaagaaagagtttgacatatttttctcatatttccagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgaggtaaggcccctactctaatggttagttccaatttattggttcatttctctgtgtttgataagattttgcctctgctgtgcatttcaatattatatctaaactgtgctgcttatagcttatcaaactactcttgctcataaagatgcaacttttatgcagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgagtcataataaatgataacatattattgttctgtggactgaaaatctgaaatcctcctttcttgagaaatacaacctcaattttatggtgatattgacaagaaggaagagcgcatataatttaaagttaaaacacggaaaataatagaagtatgtgagcagaacgcaggagcaaggggtggaactcataacaagttcaaacttcatgcatgttcataattccccaatgcaagtatggcaagaaaaatagacaacttttaaaagtatgctaatggaactgcatgcattgtgtaaacttgcttttgtcaagcatgttggaacatactatatatttgctgctgatgatgaaaaatttgcttgagtactcttctattgctaatggacgcttaccattccctgttttgtcttcaaatgcatcaggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactggttggtaaaatcagcaatggaattttctatcgtgaattttttataattattgtaacattaattactcccatgtctttttattctgttgagataccatgtgacaatgtctacaattgcatattttgcattctttggtagatcagttcccttcttaagatccccaaagggaaactggagtatcaataacttactccctcaaataaaaaaaaagtgatactttaggcataaacacattcttcaaaacaaaactttgaatagcagtttcttttgtaatatattaataaagcatggaaaaattataatgtgaagaatgtacattcagggacaaatatactcatcatttttgcatggttgatcttaatgtccatagagagattgatcagaccaaaagtttatttttgacaggagtgagtatggattataataggagtgaatggcaatatgctgtgaaagtgagcattttatatttgtttactactagcatttggtgacaaagcaaatgatgtatattgtaaatcaaagtaaacaagtagaacatgtcctgtggcgacaagacataacctacaaattaaagttagcctgattgtttattttactgggagcacattttttctagattctagagtacatattattaaaaaaggtgtgatatgtttgcaaattactttttaatggaagtttgtaaattactttgaaacttgcattcagtttttttgcttcttaaaaaacatagtgcatatgggctttatgagctcatactatgcatgtttttctgcattggagtgggacctttatctttttcttgcaactcggatgctaatgcttattataaacacaagttactaacatgtgcttgggtcattcagcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcataggtaagctcttatgatcctgtccactgttgtaccatgtgtactttccaaatgattgctaacatgctatttttcgtcagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcgggtatatcatacatctctctggctgttgtttcagtgctatcaatcaagatgacacatcatgcaatattatcataacagcacattatgtgttttgaatattttattgattgtcatagctttgttttatgttttagcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcaggtatgcagtgtttcctgcacctacttgctacctattctgtatttgatgtattcaatttcctttttatcagttttctcgaaatatatgctaataggcttgtcactaaacggctacattatccagactctgttctgaatttttttttcattaagtactgttagctgccagattacatcatcatgactaaagtagtttagctacaatattgcaaagctagatttgtctagcctctattgttttgaatagaaaaggcaggtatgcagtgtttcctgcacctacttgctacctattctgtatttgatgtattcaatttcctttttatcagttttctcgaaatatatgctaataggcttgtcactaaacggctacattatccagactctgttctgaatttttttttcattaagtactgttagctgccagattacatcatcatgactaaagtagtttagctacaatattgcaaagctagatttgtctagcctctattgttttgaatagaaaattaaatcttttggtcatttttttgtaaccttttaggagtctctgtatgcttgtcgtcaacatgctatgataagtttatatcagttcttttggagctatgatgtattcttgtatattttatgactgctaggatattatgttaaaactgttttaaaggtatcatcaaacaagttaatatagatattgctacctaccaaagttggaatttgattatgtcgtaaaatatatgcaccctacatgttttagcttcatattgacttatattttcatatgtgctccttcaactgcacaaagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggtatggtcattcctgcttataagataccagaataatgctatgaagaccgtccttggttgtcagcattcttatacacaatgataatctttgtttttcatgggtaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggtcaagatattgttattgttatatttacatacttgttatcatatttaaaagcttcatattgtatgtacctttgaaaaaaaaagttccatactgtagcttcttttgttctcatttcttggtttcaaagcatcagcatcattaccgatataaaatatacccatttcctcagtcatcttccccgactcttgtatctttcacaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagaggtaacaccttcattcatgaggtccttaggtgcaaatcccagtgaagaataatcagttctactttgtatggttcatctcgcttactgaccggttcatctcgggatcaattacgagcagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaaggcatgtcctcttctctgatctcactcacgttcatgccaaacttactcatcatcatccatgatccatcatcctgactgaatcttgatatgcagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctgacaaggccacaccatgactgtcactgcaagcagctgctgtaggatcagcgagtaggaaaggatgacttgccggccagtttcctgagatgtgatttaagactgatattagctgatgttcccaagcatgatacggggcttgctcccaagatgtgatttttactttacttttggttaatgtgctaccgctgacattagatgttcaagcatatgaaaactgcttgtgctgtaacttgtatgtctgtaacgtaaaaatctttccatatggtaaccggggaagcagaggtatatccttttctgctgttgtatctatttccagtttcctgctgaacttgcaaatgaatattcagcaatttccgttttgctc</dnaseqindica>|<br />
Link = [http://www.ncbi.nlm.nih.gov/nuccore/NM_001048755.1 RefSeq:Os01g0182600]|<br />
}}<br />
[[Category:Genes]]<br />
[[Category:Japonica mRNA]]<br />
[[Category:Oryza Sativa Japonica Group]]<br />
[[Category:Japonica Genes]]<br />
[[Category:Japonica Chromosome 1]]<br />
[[Category:Chromosome 1]]</div>Mikahttps://ngdc.cncb.ac.cn/ricewiki/index.php?title=Os01g0182600&diff=183668Os01g01826002014-06-10T13:57:42Z<p>Mika: /* Function */</p>
<hr />
<div>Please input one-sentence summary here.<br />
<br />
==Annotated Information==<br />
===Function===<br />
*''OsGI'',a ''GI'' ortholog, regulates flowering time in response to photoperiodic conditions. Mutations in this gene delay flowering in inductive SDs (short-days),whereas flowering is marginally induced under LDs(long-days).In comparison, over expression of ''OsGI'' in transgenic rice causes late flowering in both SDs and LDs.In addition, ''OsGI'' has function to regulate ''Hd1'', a gene that also shows circadian rhythm<ref name="ref1" />. ''Os-GI'' affects gene expression of approximately 75% of genes among 27,201 genes and is required for strong amplitudes and proper phase-setting of global gene expression under natural field conditions<ref name="ref2" />. <br />
*The cDNA of ''OsGI'' firstly found in ''se5'' mutant that is completely photoperiod-insensitive, the expression patterns of rice and Arabidopsis ''GI'' gene are both photoperiodically regulated and are very similar under LD and SD conditions(FIG1)<ref name="ref3" />. ''GI'' is required for phase-dependent ''CO'' transcription through the interaction with FKF1, which leads to cycling DNA binding with one finger (CDF) degradation and consequent derepression of CO. In addition, it has been known that GI protein interacts with ZEITLUPE (ZTL) F-box protein to control the diurnal rhythm of TOC1 protein in regulation of the Arabidopsis circadian clock<ref name="ref4" />. <br />
*''Os-GI'' also has function to maintain bioactive GA level through the regulation of the GA-deactivating enzyme genes in rice, ''Os-GI'' represses major <br />
''OsGA2ox'' genes in order to maintain a proper bioactive GA level, ''osgi-1'' plants showed a semi-dwarf phenotype with elevated ''OsGA2ox'' gene expression(FIG2) [4].<br />
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===Expression===<br />
Please input expression information here.<br />
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===Evolution===<br />
Please input evolution information here.<br />
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*there is some evolutionary conservation between rice circadian clock–related genes and those associated with Arabidopsis circadian clocks,where the circadian clock consists of interlocked subloops, it is also possible that Os-GI is a member of a subloop in rice circadian clocks<br />
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==Labs working on this gene==<br />
Please input related labs here.<br />
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==Knowledge Extension==<br />
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==References==<br />
Please input cited references here.<br />
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==Structured Information==<br />
{{JaponicaGene|<br />
GeneName = Os01g0182600|<br />
Description = GIGANTEA protein|<br />
Version = NM_001048755.1 GI:115434923 GeneID:4325329|<br />
Length = 9083 bp|<br />
Definition = Oryza sativa Japonica Group Os01g0182600, complete gene.|<br />
Source = Oryza sativa Japonica Group<br />
<br />
ORGANISM Oryza sativa Japonica Group<br />
Eukaryota; Viridiplantae; Streptophyta; Embryophyta; Tracheophyta;<br />
Spermatophyta; Magnoliophyta; Liliopsida; Poales; Poaceae; BEP<br />
clade; Ehrhartoideae; Oryzeae; Oryza.<br />
|<br />
Chromosome = [[:category:Japonica Chromosome 1|Chromosome 1]]|<br />
AP = Chromosome 1:4328362..4337444|<br />
CDS = 4328725..4329030,4329123..4329215,4329333..4329551,4329752..4329820,4329925..4331421<br>,4332220..4332429,4332563..4332713,4332791..4333031,4333898..4333956<br>,4334379..4334653,4334813..4334955,4335034..4335088,4335181..4335258<br>,4335351..4335437|<br />
GCID = <gbrowseImage1><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage1>|<br />
GSID = <gbrowseImage2><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage2>|<br />
CDNA = <cdnaseq>atgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactgcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcatagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcggcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctga</cdnaseq>|<br />
AA = <aaseq>MSASNEKWIDGLQFSSLFWPPPQDSQQKQAQILAYVEYFGQFTA DSEQFPEDIAQLIQSCYPSKEKRLVDEVLATFVLHHPEHGHAVVHPILSRIIDGTLSY DRNGFPFMSFISLFSHTSEKEYSEQWALACGEILRVLTHYNRPIFKVDHQHSEAECSS TSDQASSCESMEKRANGSPRNEPDRKPLRPLSPWITDILLAAPLGIRSDYFRWCGGVM GKYAAGGELKPPTTAYSRGSGKHPQLMPSTPRWAVANGAGVILSVCDEEVARYETANL TAAAVPALLLPPPTTPLDEHLVAGLPPLEPYARLFHRYYAIATPSATQRLLFGLLEAP PSWAPDALDAAVQLVELLRAAEDYDSGMRLPKNWMHLHFLRAIGTAMSMRAGIAADTS AALLFRILSQPTLLFPPLRHAEGVELHHEPLGGYVSSYKRQLEVPASEATIDATAQGI ASMLCAHGPDVEWRICTIWEAAYGLLPLSSSAVDLPEIVVAAPLQPPTLSWSLYLPLL KVFEYLPRGSPSEACLMRIFVATVEAILRRTFPSETSEQSRKPRSQSKNLAVAELRTM IHSLFVESCASMDLASRLLFVVLTVCVSHQALPGGSKRPTGSDNHSSEEVTNDSRLTN GRNRCKKRQGPVATFDSYVLAAVCALSCELQLFPFISKNGNHSNLKDSIKIVIPGKTT GISNELHNSISSAILHTRRILGILEALFSLKPSSVGTSWSYSSNEIVAAAMVAAHVSE LFRRSRPCLNALSALKQCKWDAEISTRASSLYHLIDLHGKTVTSIVNKAEPLEAHLTL TPVKKDEPPIEEKNINSSDGGALEKKDASRSHRKNGFARPLLKCAEDVILNGDVASTS GKAIASLQVEASDLANFLTMDRNGGYRGSQTLLRSVLSEKQELCFSVVSLLWQKLIAS PEMQMSAESTSAHQGWRKVVDALCDIVSASPTKASAAIVLQAEKDLQPWIARDDEQGQ KMWRVNQRIVKLIAELMRNHDSPEALVILASASDLLLRATDGMLVDGEACTLPQLELL EVTARAVHLIVEWGDSGVSVADGLSNLLKCRLSTTIRCLSHPSAHVRALSMSVLRDIL NSGQINSSKLIQGEHRNGIQSPTYQCLAASIINWQADVERCIEWEAHSRRATGLTLAF LTAAAKELGCPLTC</aaseq>|<br />
DNA = <dnaseqindica>8415..8720#8230..8322#7894..8112#7625..7693#6024..7520#5016..5225#4732..4882#4414..4654#3489..3547#2792..3066#2490..2632#2357..2411#2187..2264#2008..2094#ccgagccgtttctctccctctctctctcctcctccttcccctgcgcctcctccaccgcccgtgtcgccgccgcgcgcgtctatcccctcgccgccgccgccgtctacgccgcgtgaggtccggattggttgaaaggctgatcgaggtaatgcttttttatttttcccgtgcggttttagtttttctgtgctgtgcttggtggttttttctttaaaactatttttaggctggttttgatgatggcctttctaaggttgtcgtgtatggggggcgaggtggttcgggtggatttgtggtgttatgttgaggggatttttgtttggggggagagagttttggtggtagatgcggttgggattttgaagacgtgaaagggaaattagttttgaaatttgacggcgaggttgagctgattagtattgtaataattcgtggcatcggatagatcaacgcgagatggatgcttattactgcgaatttcaggaggtttttgtggtttttttaggcgagtagcgttgattctagctggattttgctgcttcgtggaggcgtttagatgtggtgcatacggctgtggatgtaatttgtgcggggataattactgaaaattaattcgcagtaagctagctctgctttaatattttttcccctgaaacagtaacaagggatcccttcctcatttcgttaacagtggaagtgaatacagacaccaaccgatgagcagagacacctgtaattaaacaccaaaagaagaacctgttttgttcctatatattttccatttctatgctttaggtaatatactttctggttcggtgtcatagtacccgaaatagaatctttgcttgccaaaaaaagaaagaataacaagatagaacaacatcattaaattctgtcttagacaatacatttataatcgaccaacaaaaacagaccgatgagtgacgtggaataacaaatgaagtataaatgtcacgaaaaacacatctattgattttgtgtgataagccataggagcctacgaggaaccaagtgaatggttctgtacaccacatccattagcattaccataaatttgcctatttctatgatgccttccttaacatatttatcgacatgtgagcttgcaccataactaaatgttatggtaaactttaccttataaatgattgcaacatgtaccttttcatttcgataattaaaaacgagaaagcttccacctttttcctagatatattttttcgtttgattatgaatttagttgtacttgacaataaaataatgcgcttgtaagattatgcaattttcgttgcattacctccctctcttttcatttgtcagtttttttctactccctccgtttcatattttaagtcgtttgacttttttcctagtcagactttgttaagtttgacaaagtttatagtaaaatttagcaatatctaaaatattaaattggtttcatcaaatctaacattgaatatattttgataatatgttttttttgtgttgaaaatacttctatatttttctataaacttagtcaaacttggaagaaatttcagtaagaaaaaagtcaaacggcttataatatgaaacggagggagtaaatttttttttgcattaattaaaattcatgtattttcctccttttttggtttttgttgtgcgatatattacagtgtagagactctgaagctcatatggttggcattggttgatatgataacgtgtcacgttgtgagatgcagacccccgttatgggggttgattgaatgaagtggatagcactcttataactttcttgctcgaagtttcttgataccaaggcaagtagcttataagaagatttggactttggtggcataaatggatgaactaactggcggcttcaacgtggtgttgattacaggtgcttttattgctaactggagcagtttgtgtagtttctttctttctctttgagaattctctcagtttcttgggcaggctcttgagctaccaagtatgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggtaacattattcctatttgattcttgttagaaattctatatgttgaacatacttgcctcaatgggaaacttggttgcgtctcctctctgaaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcaggtactgtatctctaagcatgtttagtatcttctgtgataaaatctttctccagtgtgttaacttttgcattctttttcttgcatgcttgcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattaggtaatttgatccagtgatcaatacatgttccacaaacaattgaagaaagagtttgacatatttttctcatatttccagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgaggtaaggcccctactctaatggttagttccaatttattggttcatttctctgtgtttgataagattttgcctctgctgtgcatttcaatattatatctaaactgtgctgcttatagcttatcaaactactcttgctcataaagatgcaacttttatgcagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgagtcataataaatgataacatattattgttctgtggactgaaaatctgaaatcctcctttcttgagaaatacaacctcaattttatggtgatattgacaagaaggaagagcgcatataatttaaagttaaaacacggaaaataatagaagtatgtgagcagaacgcaggagcaaggggtggaactcataacaagttcaaacttcatgcatgttcataattccccaatgcaagtatggcaagaaaaatagacaacttttaaaagtatgctaatggaactgcatgcattgtgtaaacttgcttttgtcaagcatgttggaacatactatatatttgctgctgatgatgaaaaatttgcttgagtactcttctattgctaatggacgcttaccattccctgttttgtcttcaaatgcatcaggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactggttggtaaaatcagcaatggaattttctatcgtgaattttttataattattgtaacattaattactcccatgtctttttattctgttgagataccatgtgacaatgtctacaattgcatattttgcattctttggtagatcagttcccttcttaagatccccaaagggaaactggagtatcaataacttactccctcaaataaaaaaaaagtgatactttaggcataaacacattcttcaaaacaaaactttgaatagcagtttcttttgtaatatattaataaagcatggaaaaattataatgtgaagaatgtacattcagggacaaatatactcatcatttttgcatggttgatcttaatgtccatagagagattgatcagaccaaaagtttatttttgacaggagtgagtatggattataataggagtgaatggcaatatgctgtgaaagtgagcattttatatttgtttactactagcatttggtgacaaagcaaatgatgtatattgtaaatcaaagtaaacaagtagaacatgtcctgtggcgacaagacataacctacaaattaaagttagcctgattgtttattttactgggagcacattttttctagattctagagtacatattattaaaaaaggtgtgatatgtttgcaaattactttttaatggaagtttgtaaattactttgaaacttgcattcagtttttttgcttcttaaaaaacatagtgcatatgggctttatgagctcatactatgcatgtttttctgcattggagtgggacctttatctttttcttgcaactcggatgctaatgcttattataaacacaagttactaacatgtgcttgggtcattcagcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcataggtaagctcttatgatcctgtccactgttgtaccatgtgtactttccaaatgattgctaacatgctatttttcgtcagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcgggtatatcatacatctctctggctgttgtttcagtgctatcaatcaagatgacacatcatgcaatattatcataacagcacattatgtgttttgaatattttattgattgtcatagctttgttttatgttttagcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcaggtatgcagtgtttcctgcacctacttgctacctattctgtatttgatgtattcaatttcctttttatcagttttctcgaaatatatgctaataggcttgtcactaaacggctacattatccagactctgttctgaatttttttttcattaagtactgttagctgccagattacatcatcatgactaaagtagtttagctacaatattgcaaagctagatttgtctagcctctattgttttgaatagaaaaggcaggtatgcagtgtttcctgcacctacttgctacctattctgtatttgatgtattcaatttcctttttatcagttttctcgaaatatatgctaataggcttgtcactaaacggctacattatccagactctgttctgaatttttttttcattaagtactgttagctgccagattacatcatcatgactaaagtagtttagctacaatattgcaaagctagatttgtctagcctctattgttttgaatagaaaattaaatcttttggtcatttttttgtaaccttttaggagtctctgtatgcttgtcgtcaacatgctatgataagtttatatcagttcttttggagctatgatgtattcttgtatattttatgactgctaggatattatgttaaaactgttttaaaggtatcatcaaacaagttaatatagatattgctacctaccaaagttggaatttgattatgtcgtaaaatatatgcaccctacatgttttagcttcatattgacttatattttcatatgtgctccttcaactgcacaaagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggtatggtcattcctgcttataagataccagaataatgctatgaagaccgtccttggttgtcagcattcttatacacaatgataatctttgtttttcatgggtaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggtcaagatattgttattgttatatttacatacttgttatcatatttaaaagcttcatattgtatgtacctttgaaaaaaaaagttccatactgtagcttcttttgttctcatttcttggtttcaaagcatcagcatcattaccgatataaaatatacccatttcctcagtcatcttccccgactcttgtatctttcacaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagaggtaacaccttcattcatgaggtccttaggtgcaaatcccagtgaagaataatcagttctactttgtatggttcatctcgcttactgaccggttcatctcgggatcaattacgagcagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaaggcatgtcctcttctctgatctcactcacgttcatgccaaacttactcatcatcatccatgatccatcatcctgactgaatcttgatatgcagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctgacaaggccacaccatgactgtcactgcaagcagctgctgtaggatcagcgagtaggaaaggatgacttgccggccagtttcctgagatgtgatttaagactgatattagctgatgttcccaagcatgatacggggcttgctcccaagatgtgatttttactttacttttggttaatgtgctaccgctgacattagatgttcaagcatatgaaaactgcttgtgctgtaacttgtatgtctgtaacgtaaaaatctttccatatggtaaccggggaagcagaggtatatccttttctgctgttgtatctatttccagtttcctgctgaacttgcaaatgaatattcagcaatttccgttttgctc</dnaseqindica>|<br />
Link = [http://www.ncbi.nlm.nih.gov/nuccore/NM_001048755.1 RefSeq:Os01g0182600]|<br />
}}<br />
[[Category:Genes]]<br />
[[Category:Japonica mRNA]]<br />
[[Category:Oryza Sativa Japonica Group]]<br />
[[Category:Japonica Genes]]<br />
[[Category:Japonica Chromosome 1]]<br />
[[Category:Chromosome 1]]</div>Mikahttps://ngdc.cncb.ac.cn/ricewiki/index.php?title=Os01g0182600&diff=183662Os01g01826002014-06-10T13:51:28Z<p>Mika: /* Function */</p>
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<div>Please input one-sentence summary here.<br />
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==Annotated Information==<br />
===Function===<br />
*''OsGI'',a GI ortholog, regulates flowering time in response to photoperiodic conditions. Mutations in this gene delay flowering in inductive SDs (short-days),whereas flowering is marginally induced under LDs(long-days).In comparison, over expression of OsGI in transgenic rice causes late flowering in both SDs and LDs.In addition, OsGI has function to regulate Hd1, a gene that also shows circadian rhythm [1]. Os-GI affects gene expression of approximately 75% of genes among 27,201 genes and is required for strong amplitudes and proper phase-setting of global gene expression under natural field conditions[2]. The cDNA of OsGI firstly found in se5 mutant that is completely photoperiod-insensitive, the expression patterns of rice and Arabidopsis GI gene are both photoperiodically regulated and are very similar under LD and SD conditions(FIG1)[3]. GI is required for phase-dependent CO transcription through the interaction with FKF1, which leads to cycling DNA binding with one finger (CDF) degradation and consequent derepression of CO. In addition, it has been known that GI protein interacts with ZEITLUPE (ZTL) F-box protein to control the diurnal rhythm of TOC1 protein in regulation of the Arabidopsis circadian clock[4]. Os-GI also has function to maintain bioactive GA level through the regulation of the GA-deactivating enzyme genes in rice, Os-GI represses major <br />
OsGA2ox genes in order to maintain a proper bioactive GA level, osgi-1 plants showed a semi-dwarf phenotype with elevated OsGA2ox gene expression(FIG2) [4].<br />
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===Expression===<br />
Please input expression information here.<br />
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===Evolution===<br />
Please input evolution information here.<br />
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*there is some evolutionary conservation between rice circadian clock–related genes and those associated with Arabidopsis circadian clocks,where the circadian clock consists of interlocked subloops, it is also possible that Os-GI is a member of a subloop in rice circadian clocks<br />
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==Labs working on this gene==<br />
Please input related labs here.<br />
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==Knowledge Extension==<br />
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==References==<br />
Please input cited references here.<br />
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==Structured Information==<br />
{{JaponicaGene|<br />
GeneName = Os01g0182600|<br />
Description = GIGANTEA protein|<br />
Version = NM_001048755.1 GI:115434923 GeneID:4325329|<br />
Length = 9083 bp|<br />
Definition = Oryza sativa Japonica Group Os01g0182600, complete gene.|<br />
Source = Oryza sativa Japonica Group<br />
<br />
ORGANISM Oryza sativa Japonica Group<br />
Eukaryota; Viridiplantae; Streptophyta; Embryophyta; Tracheophyta;<br />
Spermatophyta; Magnoliophyta; Liliopsida; Poales; Poaceae; BEP<br />
clade; Ehrhartoideae; Oryzeae; Oryza.<br />
|<br />
Chromosome = [[:category:Japonica Chromosome 1|Chromosome 1]]|<br />
AP = Chromosome 1:4328362..4337444|<br />
CDS = 4328725..4329030,4329123..4329215,4329333..4329551,4329752..4329820,4329925..4331421<br>,4332220..4332429,4332563..4332713,4332791..4333031,4333898..4333956<br>,4334379..4334653,4334813..4334955,4335034..4335088,4335181..4335258<br>,4335351..4335437|<br />
GCID = <gbrowseImage1><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage1>|<br />
GSID = <gbrowseImage2><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage2>|<br />
CDNA = <cdnaseq>atgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactgcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcatagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcggcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctga</cdnaseq>|<br />
AA = <aaseq>MSASNEKWIDGLQFSSLFWPPPQDSQQKQAQILAYVEYFGQFTA DSEQFPEDIAQLIQSCYPSKEKRLVDEVLATFVLHHPEHGHAVVHPILSRIIDGTLSY DRNGFPFMSFISLFSHTSEKEYSEQWALACGEILRVLTHYNRPIFKVDHQHSEAECSS TSDQASSCESMEKRANGSPRNEPDRKPLRPLSPWITDILLAAPLGIRSDYFRWCGGVM GKYAAGGELKPPTTAYSRGSGKHPQLMPSTPRWAVANGAGVILSVCDEEVARYETANL TAAAVPALLLPPPTTPLDEHLVAGLPPLEPYARLFHRYYAIATPSATQRLLFGLLEAP PSWAPDALDAAVQLVELLRAAEDYDSGMRLPKNWMHLHFLRAIGTAMSMRAGIAADTS AALLFRILSQPTLLFPPLRHAEGVELHHEPLGGYVSSYKRQLEVPASEATIDATAQGI ASMLCAHGPDVEWRICTIWEAAYGLLPLSSSAVDLPEIVVAAPLQPPTLSWSLYLPLL KVFEYLPRGSPSEACLMRIFVATVEAILRRTFPSETSEQSRKPRSQSKNLAVAELRTM IHSLFVESCASMDLASRLLFVVLTVCVSHQALPGGSKRPTGSDNHSSEEVTNDSRLTN GRNRCKKRQGPVATFDSYVLAAVCALSCELQLFPFISKNGNHSNLKDSIKIVIPGKTT GISNELHNSISSAILHTRRILGILEALFSLKPSSVGTSWSYSSNEIVAAAMVAAHVSE LFRRSRPCLNALSALKQCKWDAEISTRASSLYHLIDLHGKTVTSIVNKAEPLEAHLTL TPVKKDEPPIEEKNINSSDGGALEKKDASRSHRKNGFARPLLKCAEDVILNGDVASTS GKAIASLQVEASDLANFLTMDRNGGYRGSQTLLRSVLSEKQELCFSVVSLLWQKLIAS PEMQMSAESTSAHQGWRKVVDALCDIVSASPTKASAAIVLQAEKDLQPWIARDDEQGQ KMWRVNQRIVKLIAELMRNHDSPEALVILASASDLLLRATDGMLVDGEACTLPQLELL EVTARAVHLIVEWGDSGVSVADGLSNLLKCRLSTTIRCLSHPSAHVRALSMSVLRDIL NSGQINSSKLIQGEHRNGIQSPTYQCLAASIINWQADVERCIEWEAHSRRATGLTLAF LTAAAKELGCPLTC</aaseq>|<br />
DNA = <dnaseqindica>8415..8720#8230..8322#7894..8112#7625..7693#6024..7520#5016..5225#4732..4882#4414..4654#3489..3547#2792..3066#2490..2632#2357..2411#2187..2264#2008..2094#ccgagccgtttctctccctctctctctcctcctccttcccctgcgcctcctccaccgcccgtgtcgccgccgcgcgcgtctatcccctcgccgccgccgccgtctacgccgcgtgaggtccggattggttgaaaggctgatcgaggtaatgcttttttatttttcccgtgcggttttagtttttctgtgctgtgcttggtggttttttctttaaaactatttttaggctggttttgatgatggcctttctaaggttgtcgtgtatggggggcgaggtggttcgggtggatttgtggtgttatgttgaggggatttttgtttggggggagagagttttggtggtagatgcggttgggattttgaagacgtgaaagggaaattagttttgaaatttgacggcgaggttgagctgattagtattgtaataattcgtggcatcggatagatcaacgcgagatggatgcttattactgcgaatttcaggaggtttttgtggtttttttaggcgagtagcgttgattctagctggattttgctgcttcgtggaggcgtttagatgtggtgcatacggctgtggatgtaatttgtgcggggataattactgaaaattaattcgcagtaagctagctctgctttaatattttttcccctgaaacagtaacaagggatcccttcctcatttcgttaacagtggaagtgaatacagacaccaaccgatgagcagagacacctgtaattaaacaccaaaagaagaacctgttttgttcctatatattttccatttctatgctttaggtaatatactttctggttcggtgtcatagtacccgaaatagaatctttgcttgccaaaaaaagaaagaataacaagatagaacaacatcattaaattctgtcttagacaatacatttataatcgaccaacaaaaacagaccgatgagtgacgtggaataacaaatgaagtataaatgtcacgaaaaacacatctattgattttgtgtgataagccataggagcctacgaggaaccaagtgaatggttctgtacaccacatccattagcattaccataaatttgcctatttctatgatgccttccttaacatatttatcgacatgtgagcttgcaccataactaaatgttatggtaaactttaccttataaatgattgcaacatgtaccttttcatttcgataattaaaaacgagaaagcttccacctttttcctagatatattttttcgtttgattatgaatttagttgtacttgacaataaaataatgcgcttgtaagattatgcaattttcgttgcattacctccctctcttttcatttgtcagtttttttctactccctccgtttcatattttaagtcgtttgacttttttcctagtcagactttgttaagtttgacaaagtttatagtaaaatttagcaatatctaaaatattaaattggtttcatcaaatctaacattgaatatattttgataatatgttttttttgtgttgaaaatacttctatatttttctataaacttagtcaaacttggaagaaatttcagtaagaaaaaagtcaaacggcttataatatgaaacggagggagtaaatttttttttgcattaattaaaattcatgtattttcctccttttttggtttttgttgtgcgatatattacagtgtagagactctgaagctcatatggttggcattggttgatatgataacgtgtcacgttgtgagatgcagacccccgttatgggggttgattgaatgaagtggatagcactcttataactttcttgctcgaagtttcttgataccaaggcaagtagcttataagaagatttggactttggtggcataaatggatgaactaactggcggcttcaacgtggtgttgattacaggtgcttttattgctaactggagcagtttgtgtagtttctttctttctctttgagaattctctcagtttcttgggcaggctcttgagctaccaagtatgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggtaacattattcctatttgattcttgttagaaattctatatgttgaacatacttgcctcaatgggaaacttggttgcgtctcctctctgaaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcaggtactgtatctctaagcatgtttagtatcttctgtgataaaatctttctccagtgtgttaacttttgcattctttttcttgcatgcttgcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattaggtaatttgatccagtgatcaatacatgttccacaaacaattgaagaaagagtttgacatatttttctcatatttccagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgaggtaaggcccctactctaatggttagttccaatttattggttcatttctctgtgtttgataagattttgcctctgctgtgcatttcaatattatatctaaactgtgctgcttatagcttatcaaactactcttgctcataaagatgcaacttttatgcagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgagtcataataaatgataacatattattgttctgtggactgaaaatctgaaatcctcctttcttgagaaatacaacctcaattttatggtgatattgacaagaaggaagagcgcatataatttaaagttaaaacacggaaaataatagaagtatgtgagcagaacgcaggagcaaggggtggaactcataacaagttcaaacttcatgcatgttcataattccccaatgcaagtatggcaagaaaaatagacaacttttaaaagtatgctaatggaactgcatgcattgtgtaaacttgcttttgtcaagcatgttggaacatactatatatttgctgctgatgatgaaaaatttgcttgagtactcttctattgctaatggacgcttaccattccctgttttgtcttcaaatgcatcaggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactggttggtaaaatcagcaatggaattttctatcgtgaattttttataattattgtaacattaattactcccatgtctttttattctgttgagataccatgtgacaatgtctacaattgcatattttgcattctttggtagatcagttcccttcttaagatccccaaagggaaactggagtatcaataacttactccctcaaataaaaaaaaagtgatactttaggcataaacacattcttcaaaacaaaactttgaatagcagtttcttttgtaatatattaataaagcatggaaaaattataatgtgaagaatgtacattcagggacaaatatactcatcatttttgcatggttgatcttaatgtccatagagagattgatcagaccaaaagtttatttttgacaggagtgagtatggattataataggagtgaatggcaatatgctgtgaaagtgagcattttatatttgtttactactagcatttggtgacaaagcaaatgatgtatattgtaaatcaaagtaaacaagtagaacatgtcctgtggcgacaagacataacctacaaattaaagttagcctgattgtttattttactgggagcacattttttctagattctagagtacatattattaaaaaaggtgtgatatgtttgcaaattactttttaatggaagtttgtaaattactttgaaacttgcattcagtttttttgcttcttaaaaaacatagtgcatatgggctttatgagctcatactatgcatgtttttctgcattggagtgggacctttatctttttcttgcaactcggatgctaatgcttattataaacacaagttactaacatgtgcttgggtcattcagcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcataggtaagctcttatgatcctgtccactgttgtaccatgtgtactttccaaatgattgctaacatgctatttttcgtcagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcgggtatatcatacatctctctggctgttgtttcagtgctatcaatcaagatgacacatcatgcaatattatcataacagcacattatgtgttttgaatattttattgattgtcatagctttgttttatgttttagcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcaggtatgcagtgtttcctgcacctacttgctacctattctgtatttgatgtattcaatttcctttttatcagttttctcgaaatatatgctaataggcttgtcactaaacggctacattatccagactctgttctgaatttttttttcattaagtactgttagctgccagattacatcatcatgactaaagtagtttagctacaatattgcaaagctagatttgtctagcctctattgttttgaatagaaaaggcaggtatgcagtgtttcctgcacctacttgctacctattctgtatttgatgtattcaatttcctttttatcagttttctcgaaatatatgctaataggcttgtcactaaacggctacattatccagactctgttctgaatttttttttcattaagtactgttagctgccagattacatcatcatgactaaagtagtttagctacaatattgcaaagctagatttgtctagcctctattgttttgaatagaaaattaaatcttttggtcatttttttgtaaccttttaggagtctctgtatgcttgtcgtcaacatgctatgataagtttatatcagttcttttggagctatgatgtattcttgtatattttatgactgctaggatattatgttaaaactgttttaaaggtatcatcaaacaagttaatatagatattgctacctaccaaagttggaatttgattatgtcgtaaaatatatgcaccctacatgttttagcttcatattgacttatattttcatatgtgctccttcaactgcacaaagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggtatggtcattcctgcttataagataccagaataatgctatgaagaccgtccttggttgtcagcattcttatacacaatgataatctttgtttttcatgggtaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggtcaagatattgttattgttatatttacatacttgttatcatatttaaaagcttcatattgtatgtacctttgaaaaaaaaagttccatactgtagcttcttttgttctcatttcttggtttcaaagcatcagcatcattaccgatataaaatatacccatttcctcagtcatcttccccgactcttgtatctttcacaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagaggtaacaccttcattcatgaggtccttaggtgcaaatcccagtgaagaataatcagttctactttgtatggttcatctcgcttactgaccggttcatctcgggatcaattacgagcagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaaggcatgtcctcttctctgatctcactcacgttcatgccaaacttactcatcatcatccatgatccatcatcctgactgaatcttgatatgcagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctgacaaggccacaccatgactgtcactgcaagcagctgctgtaggatcagcgagtaggaaaggatgacttgccggccagtttcctgagatgtgatttaagactgatattagctgatgttcccaagcatgatacggggcttgctcccaagatgtgatttttactttacttttggttaatgtgctaccgctgacattagatgttcaagcatatgaaaactgcttgtgctgtaacttgtatgtctgtaacgtaaaaatctttccatatggtaaccggggaagcagaggtatatccttttctgctgttgtatctatttccagtttcctgctgaacttgcaaatgaatattcagcaatttccgttttgctc</dnaseqindica>|<br />
Link = [http://www.ncbi.nlm.nih.gov/nuccore/NM_001048755.1 RefSeq:Os01g0182600]|<br />
}}<br />
[[Category:Genes]]<br />
[[Category:Japonica mRNA]]<br />
[[Category:Oryza Sativa Japonica Group]]<br />
[[Category:Japonica Genes]]<br />
[[Category:Japonica Chromosome 1]]<br />
[[Category:Chromosome 1]]</div>Mikahttps://ngdc.cncb.ac.cn/ricewiki/index.php?title=Os01g0182600&diff=183660Os01g01826002014-06-10T13:50:42Z<p>Mika: /* Function */</p>
<hr />
<div>Please input one-sentence summary here.<br />
<br />
==Annotated Information==<br />
===Function===<br />
*OsGI,a GI ortholog, regulates flowering time in response to photoperiodic conditions. Mutations in this gene delay flowering in inductive SDs (short-days),whereas flowering is marginally induced under LDs(long-days).In comparison, over expression of OsGI in transgenic rice causes late flowering in both SDs and LDs.In addition, OsGI has function to regulate Hd1, a gene that also shows circadian rhythm [1]. Os-GI affects gene expression of approximately 75% of genes among 27,201 genes and is required for strong amplitudes and proper phase-setting of global gene expression under natural field conditions[2]. The cDNA of OsGI firstly found in se5 mutant that is completely photoperiod-insensitive, the expression patterns of rice and Arabidopsis GI gene are both photoperiodically regulated and are very similar under LD and SD conditions(FIG1)[3]. GI is required for phase-dependent CO transcription through the interaction with FKF1, which leads to cycling DNA binding with one finger (CDF) degradation and consequent derepression of CO. In addition, it has been known that GI protein interacts with ZEITLUPE (ZTL) F-box protein to control the diurnal rhythm of TOC1 protein in regulation of the Arabidopsis circadian clock[4]. Os-GI also has function to maintain bioactive GA level through the regulation of the GA-deactivating enzyme genes in rice, Os-GI represses major <br />
OsGA2ox genes in order to maintain a proper bioactive GA level, osgi-1 plants showed a semi-dwarf phenotype with elevated OsGA2ox gene expression(FIG2) [4].<br />
<br />
===Expression===<br />
Please input expression information here.<br />
<br />
===Evolution===<br />
Please input evolution information here.<br />
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*there is some evolutionary conservation between rice circadian clock–related genes and those associated with Arabidopsis circadian clocks,where the circadian clock consists of interlocked subloops, it is also possible that Os-GI is a member of a subloop in rice circadian clocks<br />
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==Labs working on this gene==<br />
Please input related labs here.<br />
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==Knowledge Extension==<br />
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==References==<br />
Please input cited references here.<br />
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==Structured Information==<br />
{{JaponicaGene|<br />
GeneName = Os01g0182600|<br />
Description = GIGANTEA protein|<br />
Version = NM_001048755.1 GI:115434923 GeneID:4325329|<br />
Length = 9083 bp|<br />
Definition = Oryza sativa Japonica Group Os01g0182600, complete gene.|<br />
Source = Oryza sativa Japonica Group<br />
<br />
ORGANISM Oryza sativa Japonica Group<br />
Eukaryota; Viridiplantae; Streptophyta; Embryophyta; Tracheophyta;<br />
Spermatophyta; Magnoliophyta; Liliopsida; Poales; Poaceae; BEP<br />
clade; Ehrhartoideae; Oryzeae; Oryza.<br />
|<br />
Chromosome = [[:category:Japonica Chromosome 1|Chromosome 1]]|<br />
AP = Chromosome 1:4328362..4337444|<br />
CDS = 4328725..4329030,4329123..4329215,4329333..4329551,4329752..4329820,4329925..4331421<br>,4332220..4332429,4332563..4332713,4332791..4333031,4333898..4333956<br>,4334379..4334653,4334813..4334955,4335034..4335088,4335181..4335258<br>,4335351..4335437|<br />
GCID = <gbrowseImage1><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage1>|<br />
GSID = <gbrowseImage2><br />
name=NC_008394:4328362..4337444<br />
source=RiceChromosome01<br />
preset=GeneLocation<br />
</gbrowseImage2>|<br />
CDNA = <cdnaseq>atgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactgcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcatagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcggcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctga</cdnaseq>|<br />
AA = <aaseq>MSASNEKWIDGLQFSSLFWPPPQDSQQKQAQILAYVEYFGQFTA DSEQFPEDIAQLIQSCYPSKEKRLVDEVLATFVLHHPEHGHAVVHPILSRIIDGTLSY DRNGFPFMSFISLFSHTSEKEYSEQWALACGEILRVLTHYNRPIFKVDHQHSEAECSS TSDQASSCESMEKRANGSPRNEPDRKPLRPLSPWITDILLAAPLGIRSDYFRWCGGVM GKYAAGGELKPPTTAYSRGSGKHPQLMPSTPRWAVANGAGVILSVCDEEVARYETANL TAAAVPALLLPPPTTPLDEHLVAGLPPLEPYARLFHRYYAIATPSATQRLLFGLLEAP PSWAPDALDAAVQLVELLRAAEDYDSGMRLPKNWMHLHFLRAIGTAMSMRAGIAADTS AALLFRILSQPTLLFPPLRHAEGVELHHEPLGGYVSSYKRQLEVPASEATIDATAQGI ASMLCAHGPDVEWRICTIWEAAYGLLPLSSSAVDLPEIVVAAPLQPPTLSWSLYLPLL KVFEYLPRGSPSEACLMRIFVATVEAILRRTFPSETSEQSRKPRSQSKNLAVAELRTM IHSLFVESCASMDLASRLLFVVLTVCVSHQALPGGSKRPTGSDNHSSEEVTNDSRLTN GRNRCKKRQGPVATFDSYVLAAVCALSCELQLFPFISKNGNHSNLKDSIKIVIPGKTT GISNELHNSISSAILHTRRILGILEALFSLKPSSVGTSWSYSSNEIVAAAMVAAHVSE LFRRSRPCLNALSALKQCKWDAEISTRASSLYHLIDLHGKTVTSIVNKAEPLEAHLTL TPVKKDEPPIEEKNINSSDGGALEKKDASRSHRKNGFARPLLKCAEDVILNGDVASTS GKAIASLQVEASDLANFLTMDRNGGYRGSQTLLRSVLSEKQELCFSVVSLLWQKLIAS PEMQMSAESTSAHQGWRKVVDALCDIVSASPTKASAAIVLQAEKDLQPWIARDDEQGQ KMWRVNQRIVKLIAELMRNHDSPEALVILASASDLLLRATDGMLVDGEACTLPQLELL EVTARAVHLIVEWGDSGVSVADGLSNLLKCRLSTTIRCLSHPSAHVRALSMSVLRDIL NSGQINSSKLIQGEHRNGIQSPTYQCLAASIINWQADVERCIEWEAHSRRATGLTLAF LTAAAKELGCPLTC</aaseq>|<br />
DNA = <dnaseqindica>8415..8720#8230..8322#7894..8112#7625..7693#6024..7520#5016..5225#4732..4882#4414..4654#3489..3547#2792..3066#2490..2632#2357..2411#2187..2264#2008..2094#ccgagccgtttctctccctctctctctcctcctccttcccctgcgcctcctccaccgcccgtgtcgccgccgcgcgcgtctatcccctcgccgccgccgccgtctacgccgcgtgaggtccggattggttgaaaggctgatcgaggtaatgcttttttatttttcccgtgcggttttagtttttctgtgctgtgcttggtggttttttctttaaaactatttttaggctggttttgatgatggcctttctaaggttgtcgtgtatggggggcgaggtggttcgggtggatttgtggtgttatgttgaggggatttttgtttggggggagagagttttggtggtagatgcggttgggattttgaagacgtgaaagggaaattagttttgaaatttgacggcgaggttgagctgattagtattgtaataattcgtggcatcggatagatcaacgcgagatggatgcttattactgcgaatttcaggaggtttttgtggtttttttaggcgagtagcgttgattctagctggattttgctgcttcgtggaggcgtttagatgtggtgcatacggctgtggatgtaatttgtgcggggataattactgaaaattaattcgcagtaagctagctctgctttaatattttttcccctgaaacagtaacaagggatcccttcctcatttcgttaacagtggaagtgaatacagacaccaaccgatgagcagagacacctgtaattaaacaccaaaagaagaacctgttttgttcctatatattttccatttctatgctttaggtaatatactttctggttcggtgtcatagtacccgaaatagaatctttgcttgccaaaaaaagaaagaataacaagatagaacaacatcattaaattctgtcttagacaatacatttataatcgaccaacaaaaacagaccgatgagtgacgtggaataacaaatgaagtataaatgtcacgaaaaacacatctattgattttgtgtgataagccataggagcctacgaggaaccaagtgaatggttctgtacaccacatccattagcattaccataaatttgcctatttctatgatgccttccttaacatatttatcgacatgtgagcttgcaccataactaaatgttatggtaaactttaccttataaatgattgcaacatgtaccttttcatttcgataattaaaaacgagaaagcttccacctttttcctagatatattttttcgtttgattatgaatttagttgtacttgacaataaaataatgcgcttgtaagattatgcaattttcgttgcattacctccctctcttttcatttgtcagtttttttctactccctccgtttcatattttaagtcgtttgacttttttcctagtcagactttgttaagtttgacaaagtttatagtaaaatttagcaatatctaaaatattaaattggtttcatcaaatctaacattgaatatattttgataatatgttttttttgtgttgaaaatacttctatatttttctataaacttagtcaaacttggaagaaatttcagtaagaaaaaagtcaaacggcttataatatgaaacggagggagtaaatttttttttgcattaattaaaattcatgtattttcctccttttttggtttttgttgtgcgatatattacagtgtagagactctgaagctcatatggttggcattggttgatatgataacgtgtcacgttgtgagatgcagacccccgttatgggggttgattgaatgaagtggatagcactcttataactttcttgctcgaagtttcttgataccaaggcaagtagcttataagaagatttggactttggtggcataaatggatgaactaactggcggcttcaacgtggtgttgattacaggtgcttttattgctaactggagcagtttgtgtagtttctttctttctctttgagaattctctcagtttcttgggcaggctcttgagctaccaagtatgtcagcttcaaatgagaagtggattgatgggctccagttctcatcactgttctggcccccaccacaggattcacaacaaaaacaggtaacattattcctatttgattcttgttagaaattctatatgttgaacatacttgcctcaatgggaaacttggttgcgtctcctctctgaaggcacaaattctggcctatgttgagtactttggccagtttacagctgacagtgagcaattccctgaagatatagctcaggtactgtatctctaagcatgtttagtatcttctgtgataaaatctttctccagtgtgttaacttttgcattctttttcttgcatgcttgcagctaattcaaagttgctatccatcaaaagaaaagcgccttgtagatgaagtattaggtaatttgatccagtgatcaatacatgttccacaaacaattgaagaaagagtttgacatatttttctcatatttccagcaacttttgttcttcatcatcctgagcatggccatgcagttgtgcatccgattctgtcacgcatcatagatgggacactcagttatgatagaaatggtttcccgttcatgtccttcatctctttatttagccatacttctgaggtaaggcccctactctaatggttagttccaatttattggttcatttctctgtgtttgataagattttgcctctgctgtgcatttcaatattatatctaaactgtgctgcttatagcttatcaaactactcttgctcataaagatgcaacttttatgcagaaagagtactcggagcagtgggccttggcctgtggagaaattcttagagttctaactcactacaacaggccaatcttcaaagttgatcaccaacatagtgaagcggaatgtagcagcacatctgatcaggcctcatcatgtgagtctatggagaaaagggctaacggttctccaagaaatgaacctgaccggaagccattgaggccactatctccttggatcacagacatattgcttgctgcacctctgggtattagaagtgactattttagatggtgagtcataataaatgataacatattattgttctgtggactgaaaatctgaaatcctcctttcttgagaaatacaacctcaattttatggtgatattgacaagaaggaagagcgcatataatttaaagttaaaacacggaaaataatagaagtatgtgagcagaacgcaggagcaaggggtggaactcataacaagttcaaacttcatgcatgttcataattccccaatgcaagtatggcaagaaaaatagacaacttttaaaagtatgctaatggaactgcatgcattgtgtaaacttgcttttgtcaagcatgttggaacatactatatatttgctgctgatgatgaaaaatttgcttgagtactcttctattgctaatggacgcttaccattccctgttttgtcttcaaatgcatcaggtgtggtggagtcatgggaaaatacgcagctggtggagaattgaagcctccaacaactggttggtaaaatcagcaatggaattttctatcgtgaattttttataattattgtaacattaattactcccatgtctttttattctgttgagataccatgtgacaatgtctacaattgcatattttgcattctttggtagatcagttcccttcttaagatccccaaagggaaactggagtatcaataacttactccctcaaataaaaaaaaagtgatactttaggcataaacacattcttcaaaacaaaactttgaatagcagtttcttttgtaatatattaataaagcatggaaaaattataatgtgaagaatgtacattcagggacaaatatactcatcatttttgcatggttgatcttaatgtccatagagagattgatcagaccaaaagtttatttttgacaggagtgagtatggattataataggagtgaatggcaatatgctgtgaaagtgagcattttatatttgtttactactagcatttggtgacaaagcaaatgatgtatattgtaaatcaaagtaaacaagtagaacatgtcctgtggcgacaagacataacctacaaattaaagttagcctgattgtttattttactgggagcacattttttctagattctagagtacatattattaaaaaaggtgtgatatgtttgcaaattactttttaatggaagtttgtaaattactttgaaacttgcattcagtttttttgcttcttaaaaaacatagtgcatatgggctttatgagctcatactatgcatgtttttctgcattggagtgggacctttatctttttcttgcaactcggatgctaatgcttattataaacacaagttactaacatgtgcttgggtcattcagcttacagccgaggatctgggaagcacccacaacttatgccatccacgcccagatgggctgttgccaatggagctggagttatactaagtgtctgtgatgaggaagtagctcgttatgagacagcaaatttgactgcggcagctgttcctgcacttctattacctccaccgaccacaccattggacgaacatttggttgcggggctccctcctcttgaaccatatgctcgcttgtttcataggtaagctcttatgatcctgtccactgttgtaccatgtgtactttccaaatgattgctaacatgctatttttcgtcagatactatgcaattgctactccaagtgctacccaaaggttgctttttggtcttctcgaggcaccaccatcatgggccccagatgcacttgatgcagcagtacagcttgttgaactccttagagcagcggaagattacgattctggcatgcgggtatatcatacatctctctggctgttgtttcagtgctatcaatcaagatgacacatcatgcaatattatcataacagcacattatgtgttttgaatattttattgattgtcatagctttgttttatgttttagcttccaaagaactggatgcatcttcatttcctgcgtgctattggaactgcaatgtcaatgagagctggtatcgctgctgatacgtctgctgctttacttttccgaatactctcccaaccgacattactttttcctccactgagacatgccgaaggagttgaactccatcatgagccactaggtggctatgtatcatcgtacaaaaggcaggtatgcagtgtttcctgcacctacttgctacctattctgtatttgatgtattcaatttcctttttatcagttttctcgaaatatatgctaataggcttgtcactaaacggctacattatccagactctgttctgaatttttttttcattaagtactgttagctgccagattacatcatcatgactaaagtagtttagctacaatattgcaaagctagatttgtctagcctctattgttttgaatagaaaaggcaggtatgcagtgtttcctgcacctacttgctacctattctgtatttgatgtattcaatttcctttttatcagttttctcgaaatatatgctaataggcttgtcactaaacggctacattatccagactctgttctgaatttttttttcattaagtactgttagctgccagattacatcatcatgactaaagtagtttagctacaatattgcaaagctagatttgtctagcctctattgttttgaatagaaaattaaatcttttggtcatttttttgtaaccttttaggagtctctgtatgcttgtcgtcaacatgctatgataagtttatatcagttcttttggagctatgatgtattcttgtatattttatgactgctaggatattatgttaaaactgttttaaaggtatcatcaaacaagttaatatagatattgctacctaccaaagttggaatttgattatgtcgtaaaatatatgcaccctacatgttttagcttcatattgacttatattttcatatgtgctccttcaactgcacaaagctggaagttcctgcatctgaagccactattgatgccactgcgcaaggcattgcttccatgctatgtgctcatggtcccgatgttgagtggagaatatgtaccatctgggaggctgcgtatggtttgctacctctgagttcatcagcagttgatttgcctgaaattgttgtagctgctccacttcagccacctactttgtcatggagcctatacttgccattgttgaaagtatttgagtatttacctcgtgggagtccatctgaagcatgccttatgagaatttttgtggcaacagttgaagctatactgagaagaacttttccatcagaaacctctgaacaatccaggaaaccaagaagtcaatctaagaaccttgctgttgctgaactccgaacaatgatacattcactctttgtggagtcctgtgcttcaatggaccttgcgtccagattactatttgtagtattaactgtttgcgtcagtcatcaagctttgcctgggggaagtaaaaggccaactggtagtgataatcattcctctgaggaggtcacaaatgattcgagattaaccaatggaagaaacagatgtaagaagagacaaggaccagttgctacattcgactcatacgttctagcagccgtttgtgccttatcttgtgagctccagctgttcccttttatttccaagaatgggaaccattcaaatctgaaggactccataaagatagtcatacctggaaaaaccactggtatcagtaacgagctacacaatagcattagctcagcgattcttcatactcgtagaatacttggcatcttggaagctctgttctccttgaagccatcatctgttggtacttcatggagttatagttcaaatgagattgttgcagcagctatggttgctgctcatgtttctgagttatttcgtcgatccaggccatgcttaaatgcactgtctgcgctgaagcaatgcaagtgggatgctgagatttctaccagggcatcatccctttaccatttgattgacttgcatggtaaaacagtgacctccattgtgaacaaagctgagcctctagaagctcacctgacccttacaccagtaaaaaaggatgaacctcccattgaggaaaagaacattaactcatcagatggtggtgcattggaaaaaaaggatgcttcaagatcacacaggaaaaatggttttgcaagaccactcttgaaatgtgcagaagatgttatactaaatggtgatgtcgcaagtacttctgggaaagccattgcaagtttacaggtggaagcttctgatttggcaaacttcctcaccatggaccgaaatgggggttacagaggttctcaaactctcctaagatctgtactgtcagagaagcaggagctatgcttctctgttgtctcattgctctggcagaagctcattgcatctcccgaaatgcagatgtctgcagaaagtacatcagctcatcagggttggagaaaggtatggtcattcctgcttataagataccagaataatgctatgaagaccgtccttggttgtcagcattcttatacacaatgataatctttgtttttcatgggtaggttgtggatgcgctttgtgacattgtttcagcctcaccgaccaaggcttcagctgctatcgttctgcaggtcaagatattgttattgttatatttacatacttgttatcatatttaaaagcttcatattgtatgtacctttgaaaaaaaaagttccatactgtagcttcttttgttctcatttcttggtttcaaagcatcagcatcattaccgatataaaatatacccatttcctcagtcatcttccccgactcttgtatctttcacaggccgagaaggacttgcagccctggattgctcgagatgatgagcaaggtcagaagatgtggagagtcaaccagcgaatagttaagctgatagcagagcttatgaggaaccacgatagcccagaagcattggtgatccttgctagtgcttcagatcttcttcttcgagcaactgatggaatgcttgttgatggtgaagcttgtactttaccacaattagaggtaacaccttcattcatgaggtccttaggtgcaaatcccagtgaagaataatcagttctactttgtatggttcatctcgcttactgaccggttcatctcgggatcaattacgagcagctattggaagtaaccgccagagcagtccatctcatcgtcgaatggggagattcaggtgtatccgtcgctgatggcctctccaatctgctgaaggcatgtcctcttctctgatctcactcacgttcatgccaaacttactcatcatcatccatgatccatcatcctgactgaatcttgatatgcagtgccgtctatcaaccaccatccgctgtctttcgcaccccagcgcgcatgtccgtgcactcagcatgtccgtccttcgcgacatcttgaacagcggacaaataaactccagtaagctcatccaaggggaacaccggaatggcatccagagcccaacctaccagtgcttggcagcaagcatcatcaactggcaagccgatgtggagagatgcatagagtgggaagcccacagccgccgcgccaccgggctgacgctcgccttcctcaccgcggcggcgaaggagctcggctgcccactcacttgctgacaaggccacaccatgactgtcactgcaagcagctgctgtaggatcagcgagtaggaaaggatgacttgccggccagtttcctgagatgtgatttaagactgatattagctgatgttcccaagcatgatacggggcttgctcccaagatgtgatttttactttacttttggttaatgtgctaccgctgacattagatgttcaagcatatgaaaactgcttgtgctgtaacttgtatgtctgtaacgtaaaaatctttccatatggtaaccggggaagcagaggtatatccttttctgctgttgtatctatttccagtttcctgctgaacttgcaaatgaatattcagcaatttccgttttgctc</dnaseqindica>|<br />
Link = [http://www.ncbi.nlm.nih.gov/nuccore/NM_001048755.1 RefSeq:Os01g0182600]|<br />
}}<br />
[[Category:Genes]]<br />
[[Category:Japonica mRNA]]<br />
[[Category:Oryza Sativa Japonica Group]]<br />
[[Category:Japonica Genes]]<br />
[[Category:Japonica Chromosome 1]]<br />
[[Category:Chromosome 1]]</div>Mika